Zyras (Zyras) facundus LAST, 1969

Assing, Volker, 2017, On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini), Beiträge Zur Entomologie = Contributions to Entomology 67 (2), pp. 213-246 : 228-229

Zyras (Zyras) facundus LAST, 1969

( Figs 2, 24 , 36 , 56–58)

Zyras (Zyras) facundus LAST, 1969: 279 .

Zyras (Zyras) semirufus CAMERON, 1939b: 18 ; preoccupied.

Type material examined: Lectotype ♂, present designation: “F. C. Drescher, G. Tangkoeban Prahoe, 4000–5000 Voet, Preanger, Java, 20.VII.1930 / Z. semirufa [sic] Cam. Type / Syntype / M. Cameron. Bequest. B.M. 1955-147 / Lectotypus ♂ Zyras semirufus Cameron , desig. V. Assing 2017 / Zyras facundus Last , det. V. Assing 2017 ” ( BMNH). Paralectotype ♀: same data as holotype ( BMNH).

Comment: The original description is based on an unspecified number of syntypes from “G. Tangkoeban Prahoe” ( CAMERON 1939b). LAST (1969) replaced Zyras semirufus CAMERON , a name preoccupied by Zyras semirufus (BERNHAUER, 1902) in the subgenus Glossacantha GEMMINGER & HAROLD, 1968 ( HLAVÁČ et al. 2011), with the nomen novum Z. facundus . Two syntypes were located in the Cameron collection. A male is designated as the lectotype.

Redescription: Body length 6.4–7.6 mm; length of forebody 2.9–3.1 mm. Coloration ( Figs 2, 24 , 36 ): forebody dark-brown to blackish-brown; abdomen pale-reddish with the posterior portions of tergites VII and VIII weakly infuscate; legs yellowish; antennae blackish-brown with antennomeres I–III slightly to distinctly paler and XI at least apically slightly paler; maxillary palpi brown to dark-brown with the apical palpomere yellowish.

Head ( Fig. 24 ) distinctly transverse, median portion extensively impunctate; punctures in lateral portions sparse and rather fine to moderately coarse. Eyes longer than postocular region in dorsal view. Antenna ( Fig. 2 ) 2.0 mm long; antennomeres IV approximately as long as broad, V weakly transverse, VI–X of gradually increasing width and increasingly transverse, X approximately 1.5 times as broad as long, and XI approximately as long as the combined length of IX and X.

Pronotum ( Fig. 24 ) 1.25–1.26 times as broad as long and 1.16–1.23 times as broad as head, broadest in anterior half, moderately convex in cross-section; punctation rather fine, sparse, and irregularly distributed, with extensive impunctate areas on either side of midline, on either side of the broadly impunctate midline with a series of punctures.

Elytra ( Fig. 24 ) 0.88–0.91 times as long as pronotum; punctation sparse and rather fine, regularly or slightly irregularly distributed. Hind wings present. Metatarsomere I slender, approximately as long as the combined length of II–IV.

Abdomen ( Fig. 36 ) narrower than elytra, with deep anterior impressions on tergites III–V; anterior impressions of tergites III–V each with a transverse row of weakly defined non-setiferous punctures, with a lateral setiferous puncture on either side and with four (tergites III–IV) or six (tergite V) setiferous punctures bearing long black setae near posterior margin; tergite VI with a transverse row of sparse non-setiferous punctures anteriorly, with a lateral setiferous puncture on either side and with six setiferous punctures bearing long dark setae near posterior margin; tergite VII with or without few scattered non-setiferous punctures anteriorly and with two transverse series of setiferous punctures posteriorly, posterior margin with palisade fringe; tergite VIII with setiferous punctures bearing long black setae in posterior twofifths, posterior margin convex, in the middle truncate to convex.

♂: posterior margin of sternite VIII strongly convex; median lobe of aedeagus 0.65 mm long and shaped as in Figs 56–57; paramere ( Fig. 58 ) longer than median lobe, 0.77 mm long, and with rather large and distinctly flattened apical lobe.

♀: posterior margin of sternite VIII weakly concave in the middle.

Comparative notes: Zyras facundus is characterized by rather massive antennae, a rather strongly transverse pronotum with distinctive punctation, rather sparse and fine punctation of the elytra, the coloration and punctation pattern of the pronotum, and by the morphology of the aedeagus. Regarding the modifications of the apical lobe of the paramere, this species is similar to Z. ambulans ASSING, 2017 from Thailand, from which it is distinguished by completely different coloration and much longer elytra alone.

Distribution: Zyras facundus is currently known only from the type locality in Java.

Figures

Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm.

Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm.

Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm.