Edoneus atheatus Holthuis, 1978

Cai, Yixiong & Husana, Daniel Edison M., 2009, Cave Shrimps Of The Genus Edoneus Holthuis, 1978, From Luzon, The Philippines, With Descriptions Of Three New Species (Crustacea: Decapoda: Atyidae), Raffles Bulletin of Zoology 57 (1), pp. 51-63 : 52-53

Edoneus atheatus Holthuis, 1978

( Fig. 1 )

Edoneus atheatus Holthuis, 1978: 220 , Figs. 5 , 6 ; Balete & Holthuis,1992: 99.

Material examined. – Holotype: Male, cl 4.6 mm, RMNH D 31898, the Philippines, Luzon, Isabela Province Maddela district , Cave near Santiago [recte: the Philippines, Luzon, Quirino Province, Aglipay Municipality, Barrio Palasian, cave (no. 6) near Sitio Disiluad, (see Balete & Holthuis, 1992; Fransen et al., 1997)]; coll. F. Brandsma, 1974.

Paratypes: Two females, cl 4.7–4.9 mm, RMNH D 31899, data same as holotype .

Other materials: One male, cl 3.3 mm, 2 females, cl 3.5–3.6 mm, RMNH D 41183, the Philippines, north Luzon, Quirino Province, Aglipay Municipality, Barrio Palasian, Sitio Disiluad. Cave no. 6, coll. D. S. Balete, 30 Dec.1985 .

Diagnosis – Rostrum short, triangular in lateral view, unarmed, reaching nearly to end of eyes. Eyes small, unpigmented; pterygostomian margin broadly rounded; telson with a very tiny posteromedian projection; distal margin with intermediate pairs of setae much shorter than lateral pair of spines; carpus of first pereiopod moderately excavated anteriorly, twice as long as high; chela 1.7 times as long as broad; fingers as long as palm; carpus of second pereipod much slender than previous one, not excavated anteriorly, 6.2 times as long as high; chela 2.3 times as long as broad; fingers distinctly longer than palm; branchial formula incomplete, with 6 pairs of gills; no arthrobranch on pereiopods and maxillipeds; podobranch of second maxilliped lamina-shaped; uropod with diaeresis bearing about 15–24 movable spinules, lateral angle much shorter than movable spinules.

Habitat. – Collected from large flowing river inside a cave in northern Luzon.

Remarks. – Found only from the type locality ( Holthuis, 1978; Balete & Holthuis, 1992). Holthuis (1978) described the new genus and new species Edoneus atheatus , on the basis of three specimens. Balete & Holthuis (1992) reported some more specimens and provided more detailed information on the type locality, as well as detailed habitat information.

Distribution. – Luzon, the Philippines.

Figures

Fig. 1. Edoneus atheatus. A, telson; B, second maxilliped; C, fist pereiopod; D, second pereiopod; E, uropodal diaeresis. Scale bars: A, E = 0.1 mm; B–D = 0.3 mm. (male, cl 2.9 mm, RMNH).

Fig. 5. Edoneus sketi, new species: A, first pereiopod; B, second pereiopod; C, third pereiopod; D, dactylus of third pereiopod; E, fifth pereiopod; F, dactylus of fifth pereiopod; G, male first pleopod; H, male second pleopod; I, uropodal diaeresis. Scale bars: A–C, E = 0.5 mm; D, F, I = 0.1 mm; G, H = 0.2 mm; (male, cl 4.1 mm, paratype, ZRC).

Fig. 6. Edoneus marulas, new species: A, cephalothorax and cephalic appendages, lateral view; B, telson; C, distal portion of telson; D, preanal carina; E, scaphocerite; F, mandible; G, maxillula; H, maxilla; I, first maxilliped; J, second maxilliped; K, third maxilliped. Scale bars: A = 1 mm; B, E = 0.5 mm; C = 0.1 mm; D, F–K = 0.3 mm. (male, cl 3.9 mm, paratype, ZRC).