Anthobium daliense, Shavrin & Smetana, 2017

Shavrin, Alexey V. & Smetana, Aleš, 2017, A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). I. Gracilipalpe, morchella and nigrum groups, Zootaxa 4365 (1), pp. 1-39 : 9-10

3.1.2.2. Anthobium daliense sp.n.

( Figs. 3 , 35–36 , 38 )

Type material examined: Holotype ♂ [plastic plate with aedeagus, abdominal tergite VIII, sternite VIII and apical segment in Canadian balsam was pinned under the plate with beetle]: ‘ CHINA: N-Yunnan Dali Bai | Nat.Aut.Pref. Diancang | Shan 3km W Dali 25°41.1'N | 100°06.8'E 2650-2750m | 29.8 0 3 A.Smetana [ C140 ]’ <rectangular label, printed>, ‘ HOLOTYPE | Anthobium | daliense sp.n. | Shavrin A. & Smetana A. 2017’ <red rectangular label, printed> ( NSMT).

Paratype ♀: ‘ CHINA: Yunnan, Dali | Cangshan ca 2500 m | 10.IV.2003 stream moss | G. de Rougemont leg.’ <rectangular label, printed>, ‘ PARATYPE | Anthobium | daliense sp.n. | Shavrin A. & Smetana A. 2017’ <red rectangular label, printed> (CRG).

Description. Measurements (n=2): HW: 0.90–0.92, HL: 0.55; AL (holotype): 1.89; OL: 0.27–0.30; PL: 0.72– 0.75; PW: 1.30–1.40; ESL: 1.90–1.97; EW: 1.72–1.84; AW: 1.18–1.57; MTbL (holotype): 1.15, MTrL (holotype): 0.55 (MTrL 1–4: 0.38; MTrL 5: 0.17); AedL (holotype): 0.52; TL: 4.00 (holotype)–4.60.

Body moderately wide, convex. Body and antennomeres 3–11 brown; lateral and basal portions of pronotum and apical segments of abdomen yellow-brown; mouthparts, antennomeres 1 and 2 and legs reddish brown. Body glossy, without microsculpture except of neck with isodiametric microsculpture. Head with irregular, sparse small punctation, with wide impunctate area between grooves; middle part of neck with very irregular, moderately deep and large punctation; pronotum with punctation as that on middle part of neck, more irregular and somewhat smaller on middle and markedly sparser on latero-apical portions; scutellum with several small punctures; punctation of elytra dense and large, smaller and coarser in prescutellar area, each elytron with six indistinct and tangled longitudinal rows of serial punctures, with moderately wide impunctated interspaces between third and fourth rows regular punctures more or less irregularly scattered; abdominal tergites with very sparse, indistinct punctures. Body glabrous; lateral sides of middle elevation of head near apical margin of grooves and infraorbital ridge near posterior third of eye with two pairs of long setae; anterior sides of middle elevation of pronotum with long setae; each elytron with irregular and very sparse long setae; abdominal tergites with regular short accumbent setation. Habitus as in Fig. 3 .

Head 1.6 times as wide as long; posterior portions of infraorbital ridges slightly convex, middle part of head moderately flattened; paratype with slightly impressed small area on middle part between grooves; grooves in front of ocelli deep and long, reaching apical third of length of eye; postocular ridges moderately smoothed; anterior portion with wide semicircular notch, slightly deeper behind antennal insertion. Eyes large, convex. Ocelli situated at about level of postocular ridges; distance between ocelli about one and half times as long as distance between ocellus and posterior margin of eye. Apical segment of maxillary palp with moderately narrow basal part slightly widened medially. Antenna with elongated antennomeres; measurements of antennomeres (holotype): 1: 0.21 × 0.08; 2: 0.16 × 0.06; 3: 0.17 × 0.06; 4–6: 0.16 × 0.06; 7–8: 0.16 × 0.07; 9–10: 0.16 × 0.08; 11: 0.23 × 0.08.

Pronotum 1.8 times as wide as long; anterior angles widely rounded; apical margin about as wide as posterior margin; anterior angles more or less rounded, slightly protruded anteriad; posterior angles moderately obtuse; latero-apical and lateral portions with regular very smoothed crenulation; disc of pronotum with very wide middle elevation and irregular, indistinct (holotype) to distinct, transverse semicircular impression on medio-basal third; lateral portions slightly explanate, each with deep pit at middle.

Elytra more or less subparallel-sided, distinctly longer than wide, reaching basal margin of abdominal tergite V (paratype) or VI, about two and half times as long as pronotum.

Metatarsus more than twice as long as metatibia.

Abdomen distinctly narrower than elytra, with a pair of small, oval tomentose spots in middle of tergite V; intersegmental membranes between tergites IV–VII narrow, covered by brickwall-like sculpture.

Male. Apical margin of abdominal tergite VIII and sternite VIII slightly sinuate. Aedeagus ( Fig. 35 ) with basal part gradually narrowed towards moderately widely rounded apex of median lobe; parameres wide, distinctly asymmetrical, with slightly longer left paramere, setiferous lobes markedly widened, with two short apical and preapical setae, median part of setiferous lobes with additional very short setae; internal sac narrow. Aedeagus laterally as in Fig. 36 .

Female. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded.

Comparative notes. Based on the shape of anterior angles of the pronotum and asymmetrical apical portions of parameres, A. daliense sp.n. is similar to the Himalayan A. gracilipalpe , from which it differs by the smaller and sparser punctation of head and pronotum, by the lack of longitudinal impression and shape of medio-basal impression on the middle elevation of the pronotum, as well as by the larger aedeagus with longer and stronger apical parts of parameres, and by different structure of the internal sac.

Distribution. Anthobium daliense sp.n. is known from two locations ( Fig. 38 ) in Diancang Shan in Yunnan, China.

Bionomics. The holotype was collected in narrow, deep creek ravine and was sifted from very wet accumulated leaves and various debris at the foot of a vertical wall (locality: C140).

Etymology. The specific epithet is the Latinized adjective derived from the name of the type locality near the Old Town Dali near Erhai Lake.

Figures

FIGURE 38. Distribution of Anthobium gracilipalpe group: A. gracilipalpe (black circles), A. daliense (black triangle), A. nivale (black square).

FIGURES1–3. Habitusof Anthobium gracilipalpe group:1—A.gracilipalpe (INDIA:Garhwal),2—A.nivale (holotype), 3— A. daliense (holotype). Scale bars:1.0 mm.

FIGURES 35–36. Aedeagusof Anthobium daliense (holotype):35—parameral view, 36—lateral view. Scale bar:0.1 mm.