Archaeotinodes regiomontana Melnitsky & Ivanov

Melnitsky, Stanislav I. & Ivanov, Vladimir D., 2013, Three new caddisflies species of the fossil genus Archaeotinodes (Insecta: Trichoptera: Ecnomidae) from the Baltic Amber, Zootaxa 3635 (3), pp. 261-268 : 264-265

Archaeotinodes regiomontana Melnitsky & Ivanov , new species

Figs. 3–4

Description ( Fig. 3–4 ). Thorax brown, abdomen and head pale brown; legs, antennae and wings yellow. DC in forewings short. Structure of sternite V similar to structure of sternite V in Archaeotinodes petropolitana sp. nov. Spur formula 3.4.4. Measurements: body length 4.6 mm; forewing length 5.2 mm.

Male genitalia. Sternite IX massive, protruded backwards. Gonopods short, each composed of dorsal branch rounded at apex and extended spine-like process (ventral branch). Ventral lobes of lower appendages fused, this structure divided into two apically acute processes, and the ventral branch of each lower appendage curved in lateral view. Aedeagus well sclerotized. Cerci long, widening towards apex; bases of preanal appendages narrower than their apical parts. Segment X membranous, divided into two long blades entirely covered by preanal appendages. Lower appendages, sternite IX, and outer surface of preanal appendages covered with long, pale hairs.

Comparison. Archaeotinodes regiomontana sp. nov. is similar to Archaeotinodes pauper Ulmer, 1912 . It differs from the latter in the apical part of the fused ventral branches of the lower appendages. The cerci of the new species are broader than in A. pauper and contiguous dorsally, in contrast to the separated cerci of Ulmer's species.

Holotype male. PIN, № 364/56, amber, Eocene.

Etymology. From the Latin Regiomontana - Königsberg, the former name of Kaliningrad.

Note. The sternal structure of abdominal segment V associated with the pheromone glands in fossil caddisflies previously have been observed in representatives of five families: Philopotamidae , Hydropsychidae , Psychomyiidae , Polycentropodidae , and Glossosomtidae (Ulmer 1912; Mey 1988; Ivanov & Melnitsky 2005, 2006; Melnitsky 2009). In Archaeotinodes petropolitana sp. nov. and A. regiomontana sp. nov., the anterior margin of abdominal sternite V has a clearly marked, thick, cuticular plate apparently associated with the pheromone gland orifice situated near the antecostal suture. This structure is typical for the representatives of the genus Ecnomus in the recent fauna (Ivanov & Melnitsky 1999, 2002).

Figures

FIGURE 3. Archaeotinodes regiomontana sp. nov., holotype PIN, № 364 / 56.

FIGURE 4. Archaeotinodes regiomontana sp. nov., holotype PIN, № 364 / 56, male genitalia, lateral. Scale bar = 0.1 mm. Abbreviations: AE—aedeagus, DL—dorsal lobe of lower appendage, PA—preanal appendage, SX—segment X, VL—ventral lobe of lower appendage.