Megalopsenella pouilloni, Jouault & Ngô-Muller & Pouillon & Nel, 2021

Jouault, Corentin, Ngô-Muller, Valérie, Pouillon, Jean-Marc & Nel, André, 2021, New Burmese amber fossils clarify the evolution of bethylid wasps (Hymenoptera: Chrysidoidea), Zoological Journal of the Linnean Society 191 (4), pp. 1044-1058 : 1049-1051

MEGALOPSENELLA POUILLONI SP. NOV.

( FIGS 5 , 6 )

urn:lsid:zoobank.org:act:578342A1-78A5-4F5E-9A08E5D3E1B46A83 .

Holotype: Female , accession number Po 55 (complete and well-preserved female specimen in a rounded piece of amber measuring 20 × 20 × 5 mm, without syninclusions), housed in the Musée d’Histoire Naturelle de Colmar, France.

Type locality and horizon: Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar; Lower Cenomanian, Upper Cretaceous.

Etymology: Named after Jean-Marc Pouillon, who donated the holotype.

Description: Female; body about 7 mm long, not depressed, almost smooth and not strongly foveolate; LFW 4.31 mm; castaneous; wings setose. LH 1.13 mm; WH 0.29 mm; WF 1.3 mm; HE 0.33 mm; OOL 0.55 mm; WOT 0.32 mm; DAO 0.14 mm; VOL 0.18 mm. Head not pubescent, smooth; mandible elongate, teeth not visible; six maxillary palpomeres, three (?) visible labial palpomeres; gena slightly projecting forward; reduced clypeus medially emarginate; antennal sockets partially closed dorsally by a tooth-shaped projection; antenna with 11 cylindrical flagellomeres; scape 0.47 × 0.22 mm, pedicel 0.1 2 × 0.1 2 mm, F1 0.1 4 × 0.1 2 mm (all measurements refer to length × width), F2 0.15 × 0.13 mm, F3 0.15 × 0.13 mm, F4 0.15 × 0.13 mm, F5 0.15 × 0.11 mm, F6 0.18 × 0.11 mm, F7 0.15 × 0.11 mm, F8 0.16 × 0.10 mm, F9 0.17 × 0.10 mm, F10 0.17 × 0.09 mm; F11 0.3 × 0.09 mm; occipital carina strongly marked; compound eye without microsetae; ocellar triangle not visible. Mesosoma with dorsum smooth, not sculptured;

KEY TO GENERA OF † PROTOPRISTOCERINAE 1. Antenna with 13 antennomeres with short scape; forewing venation usually developed but sometimes reduced...............................................................................................................................................................2

- Antenna with 12 antennomeres, scape elongated; forewing venation always reduced (Cenomanian: Myanmar) ................................................................................................................... Cretapristocera gen. nov.

2. Head elongated with small eyes .......................................................................................................................3

- Head almost globose with large eyes ...............................................................................................................4

3. Mandibles only with few teeth (five) along masticatory margin; legs with pro- and meta-femora clearly thickened; forewing cell 2R1 open (Cenomanian: Myanmar) ........................... Bethylitella Cockerell, 1917a

- Mandibles with small teeth along masticatory margin and with basal one elongated: legs, except pro- femora, only slightly thickened; forewing cell 2R1 almost closed (Eocene: Baltic amber) .............................. ................................................................................................................................ Protopristocera Brues, 1923

4. Notauli well developed, parapsidal furrow weak (Eocene: Oise amber) .... Gynopteron Falières & Nel, 2019

- Notauli weak, parapsidal furrow well-visible (Eocene: Baltic amber) ................. Bethylopteron Brues, 1933

Forewing covered with microtrichia and bordered with small setae; anterior border not angularly incurved anterior to pterostigma; seven closed cells present, and all veins C, Sc + R, Rs, M + Cu, Cu, A tubular; pterostigma rather broad; poststigmal abscissa of R1 long; hindwing 3.23 mm long with five hamuli. Metasoma c. 5.76 mm long, with short petiole; length of clearly visible tergites: T1 1.45 mm, T2 1 mm, T3 at least 0.8 mm; short part of sting visible.

Figures

Figure 5. Megalopsenella pouilloni, sp. nov. holotype Po55. A, habitus, in left lateral view. B, habitus, in right ventral view. Scale bars: 1 mm.

Figure 6. Megalopsenella pouilloni, sp. nov. holotype Po55. A, head detailed ventral view. B, fore and hindwings with nomenclature of cells (in bold) and veins. C, line drawing of habitus in right ventral view. Scale bars: 1 mm.