Ctenochauliodes

Liu, Xingyue, Hayashi, Fumio & Yang, Ding, 2011, Taxonomic notes and updated phylogeny of the fishfly genus Ctenochauliodes van der Weele (Megaloptera: Corydalidae), Zootaxa 2981, pp. 23-35 : 24-25

Key to males of Ctenochauliodes

1. Wings grayish brown throughout ( Yang & Liu 2010: plate IX: 61)............................ C. griseus Yang & Yang

- Wings hyaline or subhyaline, with dark markings............................................................ 2

2. Hindwings with distinct blackish brown markings from base to apex ( Yang & Liu 2010: plate X: 67)................... 3

- Hindwings immaculate or with only a few brownish markings on apex........................................... 4

3. Male tenth tergum with apex twisted inward ( Liu & Yang 2006b: Fig. 5 B); male tenth gonocoxite with apex hook-like and curved dorsad ( Liu & Yang 2006b: Fig. 5 A)............................................ C. abnormis Yang & Yang

- Male tenth tergum with apex not twisted inward (Liu & Yang 2006: Fig. 16 B); male tenth gonocoxite with apex not curved dorsad ( Liu & Yang 2006:b Fig. 16 A)..................................................... C. yangi Liu & Yang

4. Left lobe of male tenth gonocoxite in lateral view with apex obtuse or acutely pointed, but not curved dorsad............. 5

- Left lobe of male tenth gonocoxite in lateral view with apex acutely pointed and curved dorsad........................8

5. Forewing with numerous small brownish spots on proximal half of costal area ( Yang & Liu 2010: plate X: 64); left lobe of male tenth gonocoxite in lateral view strongly elevated dorsad ( Yang & Liu 2010: Fig. 121a).... C. punctulatus Yang & Yang

- Forewing without small brownish spots on proximal half of costal area; left lobe of male tenth gonocoxite in lateral view fee- bly elevated dorsad....................................................................................6

6. Forewing proximally with a broad subtriangular brown marking ( Liu & Yang 2010: plate X: 65); lobes of male tenth gonocoxite with apex rounded and not curved ventrad ( Liu & Yang 2006b: 14A).................... C. sagittiformis Liu & Yang

- Forewing proximally without broad subtriangular brown marking; lobes of male tenth gonocoxite with apex acutely pointed or curved ventrad........................................................................................ 7

7. Forewing sparsely with several small blackish brown spots ( Yang & Liu 2010: plate IX: 57); male tenth gonocoxite with left lobe acutely pointed on apex ( Liu & Yang 2006b: fig. 6A); a pair of digitiform lateral lobes presented beneath anus ( Liu & Yang 2006b: Fig. 6 A)............................................................. C. digitiformis Liu & Yang

- Forewing with dense brownish markings; male tenth gonocoxite with left lobe rounded on apex; lateral lobes beneath anus tuberculate........................................................................................... 8

8. Markings on forewing dispersedly dotted, without band-like pattern ( Fig. 1 ); male tenth gonocoxite with right lobe strongly produced into a subtriangular apex from middle portion ( Fig. 9 )........................... C. khasianus (van der Weele)

- Markings on forewing more or less fused as several narrow and band-like stripes on distal half ( Figs. 2–4 ); male tenth gonocoxite with right lobe gradually narrowed and produced into a subtriangular apex from proximal portion ( Liu & Yang 2006b: Fig. 8 C).................................................................... C. nigrovenosus (van der Weele)

9. Forewing densely patterned, with numerous small brownish spots ( Fig. 5 )........................................ 10

- Forewing sparsely patterned, with band-like stripes on distal half ( Yang & Liu 2010: plate IX: 58).................... 11

10. Male tenth gonocoxite with left lobe nearly as long as right lobe, and both lobes curved dorsad with hook-like tips in lateral view ( Liu & Yang 2006b: Fig. 15 A, C).................................................... C. similis Liu & Yang

- Male tenth gonocoxite with right lobe much longer than left lobe, and straightly directed posteriad with spinous tip in lateral view ( Liu & Yang 2006b: Fig. 12 A, C)..................................................... C. stigmosus sp. nov.

11. Male tenth gonocoxite with left lobe as long as right lobe ( Liu & Yang 2006b: Fig. 9 C)......... C. fujianensis Yang & Yang

- Male tenth gonocoxite with right lobe much longer than left lobe............................................... 12

12. Male tenth gonocoxite with right lobe at least two times as long as left lobe ( Liu & Yang 2006b: Fig. 11 C)............................................................................................. C. meridionalis Yang & Yang

- Male tenth gonocoxite with right lobe less than two times as long as left lobe ( Liu & Yang 2006b: Fig. 7 C)................................................................................................ C. elongatus Liu & Yang

Figures

FIGURES 4 – 5. Habitus images of Ctenochauliodes spp. 4. C. nigrovenosus (van der Weele), holotype male of C. forcipatus Kimmins; 5. C. stigmosus sp. nov., holotype male. Scale bars = 5.0 mm.

FIGURES 16 – 21. Female eighth gonocoxites of Ctenochauliodes spp., ventral view. 16. C. fujianensis Yang & Yang; 17. C. elongatus Yang & Yang; 18. C. griseus Yang & Yang; 19. C. meridionalis Yang & Yang; 20. C. similis Liu & Yang; 21. C. stigmosus sp. nov. Gp 8: eighth gonapophyses.

FIGURES 6 – 10. Ctenochauliodes khasianus (van der Weele). 6. Male genitalia, lateral view; 7. Male tenth tergum, dorsal view; 8. Male tenth tergum, ventral view; 9. Male tenth gonocoxite, ventral view; 10. Sclerite beneath anus, ventral view. Gx 10: tenth gonocoxite; S 9: ninth sternum; sa: sclerite beneath anus; T 9 – 10: ninth to tenth tergum. Scale bars = 1.0 mm.

FIGURES 1 – 3. Habitus images of Ctenochauliodes spp. 1. C. khasianus (van der Weele), holotype male; 2. C. nigrovenosus (van der Weele), lectotype female; 3. C. nigrovenosus (van der Weele), lectotype male of C. friedrichi Navás. Scale bars = 5.0 mm.

FIGURES 11 – 15. Female genitalia of Ctenochauliodes spp. 11. C. nigrovenosus (van der Weele), lateral view of genitalia; 12. same, ventral view of eighth gonocoxites; 13. C. abnormis Yang & Yang, ventral view of eighth gonocoxites; 14. C. punctulatus Yang & Yang, ventral view of eighth gonocoxites; 15. C. yangi Liu & Yang, ventral view of eighth gonocoxites.