Anthomyza mcalpinei, Roháćek & Barber, 2016

Roháćek, Jindřich & Barber, Kevin N., 2016, Nearctic Anthomyzidae: a monograph of Anthomyza and allied genera (Diptera), Acta Entomologica Musei Nationalis Pragae (suppl.) 56, pp. 1-412 : 161-177

Anthomyza mcalpinei sp. nov.

( Figs 3 , 275 , 277, 278 , 280–293 , 295 , 380 )

Type material. HOLOTYPE: ♂, “ONT: SSMarie, S.of Alg-oma U.College, 25.vi.2001 KNBarber, sweeps, mostly Carex aquatilis 46°39.9'N 84°17.2'W ” and “ Holotypus ♂ Anthomyza mcalpinei sp. n., J. Roháček & K. N. Barber det. 2014” (red). The specimen is in perfect condition, with well visible gonostyli (see Fig. 295 ) ( DEBU, intact). PARATYPES: CANADA: ALBERTA: Elk Island, light mixed forest and marsh, 13.vi.1993, 1 ♀, T. Clarke leg. ( DEBU); w. border Elk Island N. P., Range Rd. 210, 0.5 km N Hwy #16, 53°34.52'N 112°57.09'W, sweeps, Calamagrostis canadensis , 21.vii.2008, 16 ♂♂ 8 ♀♀, K. N. Barber leg. ( CNCI); Calgary, Fish Creek Prov. Pk., 50°54.407'N 114°01.119'W, sweep, brush near ranch house, 25.vi.2010, 1 ♀; same locality but 50°55.739'N 114°03.312'W, sweep near creek, 22.vii.2010, 1 ♂, both J. E. Swann leg. (both BDUC); Dunvegan, 55°55.39'N 118°35.74'W, sweep south facing grass slope at dusk, 19.vii.2003, 1 ♂, S. Boucher leg. ( LEMQ 0040435); Dunvegan, north shore of Peace River, sweep vegetation along shoreline, 12.vii.1997, 4 ♂♂ 4 ♀♀, T. A. Wheeler leg. ( LEMQ 0039872, -75, -79, -80, -81, -84, -89, -90); Dunvegan, n. shore of Peace River, sweep grasses at edge of agricultural neld, 13.vii.1997, 10 ♂♂ 11 ♀♀ ( LEMQ 0039869, -870, -883, -885–888, -891–895, -897–900, -902–904, -907, -995), 14.vii.1997, 5 ♂♂ 1 ♀ ( LEMQ 0039867, -871, -876–878, -901), T.A. Wheeler leg.; Halfway Slough, 59°56.049'N 111°44.344'W, sweep, 5.vii.2012, 2 ♂♂, S. & K. Williamson leg. ( BDUC); ~ 14.4 km E Obed, Range Rd. 213 @ RR crossing, 53°32.19'N 117°01.02'W, sweeps, mostly Carex utriculata , 25.vii.2011, 1 ♂; Rocky Mtn. House, 52°22.57'N 114°56.34'W, pooter, grasses on bridge embankment, edge of parking area, 24.vii.2008, 1 ♀; ~ 40.8 km SSW Rocky Mtn.House, Hwy #752, 52°09.79'N 115°25.89'W, sweeps/pooter, roadside Poeae , 24.vii.2008, 1 ♂, all K. N.Barber leg. (all CNCI); 10 km NW Whitecourt, Hwy 32 at Sakwatamau River, 54°10'N 115°42'W, sweep vegetation at river edge, 15.vii.1997, 1 ♀, T.A. Wheeler leg. ( LEMQ 0039855); R[egional] M[unicipality] of Wood Buffalo, ABMI site #67 NE, 1 km E Winnifred Lake, 59.22785187°N 110.360909°W, organic soil, Berlese, 18.vi.2014, 1 ♂, ABMI neld crew ( ABO 3) leg. ( PMAE). BRITISH COLUMBIA: Big Beaver Creek, Alaska Highway DEC-263, 13.viii.1978, 1 ♂, P.H.Arnaud Jr.leg. ( CASC, genit. prep.); Fernie,Annex Pk., 49°30'43"N 115°04'07"W, 2.viii.2010, 14 ♂♂ 8 ♀♀, S. M. Paiero leg. ( DEBU 00334407, -09–11, -17–19, -22, -24, -27, -28, -32, -38, -39, -42, -48, -54, -56–58, -66, -68); Fernie, Annex Pk., 49°30.72'N 115°04.13'W, sweeps, wet ditch, mostly Equisetum arvense & E. laevigatum , 17.vii.2011, 1 ♂ 1 ♀; same locality but sweeps, wet ditch, Carex utriculata , 17.vii.2011, 13 ♂♂ 15 ♀♀, 18.vii.2011, 3 ♂♂ 6 ♀♀, 19.vii.2011, 13 ♂♂ 2 ♀♀; Fernie, Annex Pk., 49°30.66'N 115°04.16'W, sweeps, pond margin, Carex utriculata , 18.vii.2011, 2 ♂♂, 19.vii.2011, 1 ♂, all K. N. Barber leg. (all CNCI); Kinbasket Lake, BC Hydro drawdown study, Malaise trap ( WIL 83-01), 18.vii.2008, 1 ♀, (00MRFTb01), 10.vii.2010, 1 ♀, (09 MCOT 01), 27–28.vi.2009, 1 ♀, (84 MCOT 15), 4–5.viii.2009, 1 ♀, (84 MTRT 01), 4–5.viii.2009, 1 ♂, (84MTR- Ta15), 4.vii.2010, 1 ♀, (88 MCOT 01), 10.vii.2010, 1 ♀, (91 MTRT 01), 10–11.vii.2009, 1 ♀, all Cooper Beauchesne & Assoc. Ltd. leg. (all RBCM); ~8.0 km SE Valemount, edge of Kinbasket Lake, 52°46.65'N 119°10.38'W, sweeps, mostly Carex utriculata , 23.vii.2011, 1 ♀, K. N. Barber leg. ( CNCI); Kootenay Region, Arrow Lake, 11U 449421E 5623320N, sweeping/searching, 5.vi.2010, 2 ♂♂, J. K. Sharkey ( LGL Ltd.) leg. ( RBCM); Mission City, 8.vi.1953, 1 ♂, E. Mason leg., 8.vi.1953, 1 ♂, 4.vii.1953, 1 ♀, 18.vii.1953, 1 ♂ 1 ♀, W. R. M. Mason leg. ( CNCI); Mt. Robson Prov. Pk., Hwy #16, small road towards Mt.Robson, 53°03'N 119°15'W, forest noor, swamp, (Universität Bielefeld, Ca1519), 6.viii.2002, 1 ♀, M. v. Tschirnhaus leg. ( ZSMC, in ethanol); Revelstoke, 2.vii.1973, 2 ♂♂ 2 ♀♀, H. J. Teskey leg. ( CNCI); Robson, 18.vi.1958, 1 ♀, 5.vii.1967, 1 ♂ 3 ♀♀, H. R. Foxlee leg. ( UBCZ); Vancouver, Point Grey, 23.vii.1973, 1 ♂, J. R. Vockeroth leg. ( CNCI). MANITOBA: ~ 14 km SW Falcon Lake, jct. Hwy #1 & Rd. 86E, 49°38.24'N 95°29.89'W, sweeps, graminoids, mostly Carex spp., 29.vii.2011, 1 ♂ 1 ♀, K. N. Barber leg.; Int. Peace Gardens, Turtle Mtn. For. Res., 17.vii.1958, 1 ♂, R. B. Madge leg., 7.viii.1958, 2 ♂♂, J. G. Chillcott leg.; 5 mi W Norgate, Riding Mtn. Nat. Pk., aspen-spruce swamp, 16.viii.1958, 1 ♂, J. G. Chillcott leg. (all CNCI); 4 km E Ste. Rita, Hwy 15 at Brokenhead R., sweeping sedge and grass along shore, 9.vii.1992, 1 ♂ 2 ♀♀, T. A. Wheeler leg. ( LEMQ 0039896, -905, -906); 5 mi SW Shilo, 13.viii.1958, 1 ♂; 5 mi SW Shilo, noodplain community near tamarack bog, 22.vii.1958, 1 ♀, both R. L. Hurley leg.; ~ 6 km SW Shilo, 49°45.19'N 99°41.28'W, edge of wet area, sweeps, incl. Equi. ssuviatile , Carex , Calamagrostis , Impatiens , 10.vii.2011, 1 ♂, K. N. Barber leg.; Treesbank, Assiniboine R., 18.vi.1958, 1 ♀, J. F. McAlpine leg. (all CNCI). NEW BRUNSWICK: Chamcook, 9.viii.1957, 1 ♂, G. E. Shewell leg.; Sackville, Tantramar Marsh, 14.vii.1983, 2 ♀♀, J. R. Vockeroth leg.; St. Andrews, 8.viii.1957, 4 ♂♂, G.E.Shewell leg.(all CNCI). NEWFOUNDLAND: Avalon Peninsula, Portugal Cove, Indian Meal Line, 15.vii.1982, 1 ♂ 1 ♀, [no collector] ( NFRC); Portugal Cove, pasture sweep, 1.vii.1987, 1 ♂ 2 ♀♀, T. A. Wheeler leg. ( DEBU); Terra Nova Nat. Pk., 7.vii.1961, 1 ♂, C. P. Alexander leg. ( USNM, genit. prep.). NOVA SCOTIA: C[ape] B[reton] H[ighlands] Nat. Pk., Lone Shieling, PG732859, over small stream in maple forest, 1.vii.1983, 1 ♀, J. R. Vockeroth leg.; same locality but Pleasant Bay, PG684871, wet hardwood forest, 10.vii.1984, 1 ♀, H. J. Teskey leg.; Kentville, 6.viii.1958, 1 ♂, J. R. Vockeroth leg.; Halifax Co., Lawrencetown, 19–20.vii.1967, 1 ♀, D. M.Wood leg.; Lockeport, 18.vii.1958, 1 ♂ 2 ♀♀, 25.vii.1958, 1 ♀, J. R. Vockeroth leg. (all CNCI); Petite Riviere, 44°13.2'N 64°26.2'W, sweep vegetation above ocean beach, 23.vii.2002, 1 ♂, J. Forrest & T.A.Wheeler leg. ( LEMQ 0040360); Sydney, [-]. vii.1967, 1 ♂ 2 ♀♀, N. L. H. Krauss leg. ( USNM). ONTARIO: 4 mi W Arthur, 25.vii.1985, weedy ditch, 1 ♂, K. N. Barber leg.; Belwood, 14.vii.1972, 1 ♀, D. H. Pengelly leg. (both DEBU); Blackburn, Mer Bleue, 27.vi.1955, 1 ♀, W. R. M. Mason leg. ( CNCI); Bruce Co., Crane River, 21.vii.1996, 1 ♂; Bruce Co., Miller Lake, 21.vii.1996, 1 ♂, both S. A. Marshall leg.; Bruce Co., Hwy #6 @ Willow River, 45°10.43'N 81°31.21'W, sweeps, mostly Carex aquatilis , 21.vi.2008, 2 ♂♂, K. N. Barber leg. (all DEBU); Bruce Penin[sula], 1 km NNW Ague Lake, spring creek, 45°03'48"N 81°24'34"W, sweeping Calamagrostis canadensis in a fen, 1.vii.2010, 2 ♂♂ 9 ♀♀, J. Roháček leg. ( SMOC); Bruce Peninsula N. P., bank of Crane River nr. Hwy 6, 45°08.9'N 81°28.1'W, sweeps/pooter, Calamagrostis & Carex , 31.vii.1997, 11 ♂♂ 12 ♀♀, K. N. Barber leg., sweeps, grasses / sedges, 31.vii.1997, 2 ♂♂ 1 ♀, S. A. Marshall leg.; Bruce Peninsula N. P., Dorcas Bay Rd. at Willow Creek, 45°09.4'N 81°34.4'W, sweeps, mostly creekside graminoids, 3.vii.1999, 3 ♂♂ 5 ♀♀; Bruce Peninsula N. P., Emmett Lake, 45°13.4'N 81°29.3'W, sweeps, graminoids, edge of marsh, 2.vii.1999, 1 ♂ 1 ♀; Bruce Peninsula N. P., Singing Sands, 45°11.6'N 81°34.4'W, sweeps, grasses along dry ditch, 31.vii.1997, 2 ♂♂ 3 ♀♀, 4.vii.1998, 3 ♂♂ 3 ♀♀; same locality but 45°11.6'N 81°35.2'W, sweeps, Carex , 5.vii.1998, 1 ♀, all K. N. Barber leg. (all DEBU); ~ 5 km SE Cochrane, 49°01.16'N 80°57.93'W,sweeps, railside Equisetum spp., graminoids, herbs, 18.vii.2009, 1 ♀; ~ 13.5 km S Cochrane, 48°56.65'N 81°00.18'W, hydro right-of-way, sweeps, mostly Carex utriculata , 9.vii.2012, 1 ♀, 12.vii.2013, 1 ♀; ~ 35 km WSW Dubreuilville, 2 km SE jct. Hwys #17 & #519, 48°17.16'N 84°53.34'W, sweeps, roadside Calamagrostis / Scirpus , 23.vii.2003, 1 ♂ 1 ♀; ~ 3.8 km ENE Dugwal, 48°35.33'N 80°57.90'W, sweeps, wet ditch, Carex utriculata , Equisetum ssuviatile , Scirpus , 13.vii.2013, 2 ♂♂ 3 ♀♀; ~ 5 km SSE Echo Bay, Jct. Bar River Rd./Government Rd., 46°26.52'N 84°02.80'W, sweeps, Calamagrostis , 27.viii.2003, 2 ♂♂ 1 ♀; ~ 59 km NNW Elliot Lake, s. of Rocky Is. Lake, 46°50.16'N 83°03.05'W, sweeps, mostly Carex aquatilis in fen, 3.vii.2010, 5 ♂♂ 6 ♀♀, all K. N. Barber leg. (all CNCI); ~ 59 km NNW Elliot Lake, S of Rocky Island Lake, 46°50.16'N 83°03.05'W, 455 m, sweeping, mostly Carex rostrata ? [more likely C. utriculata ] in fen, 3.vii.2010, 1 ♀; ~ 61 km NNW Elliot Lake,Three Lakes, 46°49.94'N 83°06.31'W, 425 m, sweeping grasses [mostly Glyceria striata ] near lakeshore, 3.vii.2010, 1 ♂ 4 ♀♀ (1 ♀ genit. prep.), all J. Roháček leg. (all SMOC); ~ 63 km NNW Elliot Lake, s. of Rocky Is. Lake, 46°49.80'N 83°09.08'W, sweeps, streamside graminoids, 4.vii.2010, 2 ♂♂ 2 ♀♀, K. N. Barber leg. ( CNCI); same locality but 425 m, sweeping streamside graminoids, 4.vii.2010, 19 ♂♂ 12 ♀♀, J. Roháček leg. ( SMOC); ~ 66 km NNW Elliot Lake, Rocky Island Lake, 46°50.82'N 83°08.76'W, sweeps, Scirpus [ microcarpus ] on dried shoreline, 4.vii.2010, 2 ♂♂ 4 ♀♀, K. N. Barber leg. ( CNCI); same locality but 405 m, sweeping Scirpus sp. [ S. microcarpus ] on dried shoreline, 4. vii.2010, 1 ♂ 2 ♀♀, J. Roháček leg. ( SMOC); Fathom Five N. Pk., Cove Is., Bass Bay shore, fen, p.m. sweeps, 25.vi.1995, 1 ♂; same locality but shore of Bass Bay, sweep, 26.vii.1995, 1 ♂, both S.A. Marshall leg. (both DEBU); Fergus, 43°41.5'N 80°23.2'W, Grand R. noodplain, sweeps/pooter, Bromus inermis , 17.vii.1997, 1 ♀, K. N. Barber leg. ( CNCI); Finland, S of Caliper Lake, sweep sedge at beaver dam, 10.vii.1992, 3 ♂♂ 1 ♀, T. A. Wheeler leg. ( LEMQ 0039865, -66, -68, -73); Greenwater P. Pk., 49°11.05'N 81°16.04'W, sweeps, mostly lakeside Calamagrostis , 18.vii.2009, 2 ♂♂ 2 ♀♀ ( DEBU 01501993–96); Greenwater P. Pk., 49°10.93'N 81°16.37'W, sweeps, Phalaris arundinacea in creek noodplain, 21.vii.2009, 8 ♂♂ 3 ♀♀ ( DEBU 01502108–18); Greenwater P. Pk., 49°10.91'N 81°16.28'W, sweeps, Carex , Calamagrostis , Phalaris in creek noodplain, 21.vii.2009, 17 ♂♂ 18 ♀♀, K. N. Barber leg. ( DEBU 01502140–45, -47–75); Guelph, 3.vii.1980, 2 ♂♂ 1 ♀, S. A. Marshall leg., 3.viii.1980, 1 ♂, K. N. Barber leg. ( DEBU); 6 km W Hagar, 46.47°N 80.48°W, sweep river edge, at rest area, 29.vi.2007, 1 ♂ 1 ♀, J. Mlynarek leg. ( LEMQ 0040531, -532); ~ 5 km NE Heyden, Hwy 552, 46°41.37'N 84°16.85'W, sweeps, roadside ditch/ stream, graminoids, 31.vii.2006, 1 ♂ 3 ♀♀; ~ 6 km N Heyden, Hwy #17 base of “Mile Hill”, 46°41.74'N 84°20.60'W, sweeps, graminoids under alder, 7.viii.2005, 2 ♂♂; same locality but 46°41.75'N 84°20.71'W, sweeps, graminoids in wet meadow, 7.viii.2005, 2 ♂♂; Icewater Creek WS [watershed], 12.7 km NNE Searchmont, mi.10.5 Whitman Dam Rd., alder thicket, 21.vi.1986, 2 ♂♂, 7.vii.1986, 1 ♀; Icewater Creek WS [watershed], 46°53.72'N 84°03.39'W, sweeps, riparian ferns, graminoids, 5.viii.2006, 1 ♂ 2 ♀♀, all K. N. Barber leg. (all CNCI); Lake Superior Prov. Park, Crescent Lake Trail, 47°16.73'N 84°33.12'W, sweeping Clintonia , ferns, Aralia , Maianthemum under Betula / Acer , 9.vii.2010, 1 ♀, J. Roháček leg. ( SMOC); ~ 11.9 km N Kejick, 49°43.89'N 95°04.14'W, sweeps, wet ditch, Calamagrostis canadensis , 30.vii.2008, 1 ♂; Manitoulin Is., Carter Bay, 45°36.3'N 82°08.5'W, sweeps, Pearly Everlasting [ Anaphalis margaritacea ], 30.vi.1999, 1 ♂; Manitoulin Is., Michael’s Bay Pk., 45°36.0'N 82°06.2'W, sweeps, grasses near shore, 28.vii.1997, 1 ♀; Manitoulin Is., 0.7 km N Michael’s Bay Pk., 45°36.5'N 82°06.2'W, sweeps, roadside grasses, 28.vii.1997, 4 ♂♂ 2 ♀♀, sweeps, Carex / Calamagrostis in fen nat, 5.vii.1998, 5 ♂♂ 6 ♀♀, sweeps/pooter, graminoids in fen nat, 28.vii.1997, 1 ♂ 2 ♀♀, 30.viii.2004, 3 ♂♂ 1 ♀; Manitoulin Is., ~2.2 km N Cold Springs, Perch Ck. @ Hwy 540, 45°53.2'N 82°06.3'W, sweeps, various grasses / sedges in noodplain, 5. vii.1998, 5 ♂♂ 2 ♀♀ (1 ♀ genit. prep.), sweeps/pooter, Calamagrostis canadensis , 1.viii.1997, 7 ♂♂ 4 ♀♀; same locality but 45°53.1'N 82°06.2'W, sweeps/pooter, Calamagrostis canadensis , 4.vii.1999, 13 ♂♂ 4 ♀♀, all K. N. Barber leg. (all CNCI); Manitoulin I., Misery Bay Nature Reserve, 45°47.64'N 82°44.93'W, sweeping, mostly Carex from wetland boardwalk, 2.vii.2010, 1 ♀, J. Roháček leg. ( SMOC); same locality but sweeps, mostly Carex stricta from wetlands boardwalk, 2.vii.2010, 2 ♀♀, K. N. Barber leg. ( DEBU 01502486, -87); Hwy #17, ~ 8.5 km NW Marathon, 48°47.69'N 86°26.07'W, sweeps, roadside graminoids, 2 ♀♀, 31.vii.2008, K. N. Barber leg.; Maynooth, swept from beaver meadow, 5.ix.1953, 1 ♂ 1 ♀, J. F. McAlpine leg.; Midland, 30.vii.1956, 2 ♀♀, J. G. Chillcott leg. (all CNCI); Moosonee, 51°16.07'N 80°38.66'W, sweeps, shoreline graminoids, 9.vii.2014, 1 ♀; Moosonee, 51°16.17'N 80°39.10'W, sweeps, mostly Carex utriculata , Scirpus , in wet hydro cut, 10.vii.2014, 1 ♀; Moosonee, 51°16.36'N 80°39.11'W, sweeps, railside ditch, mostly Equisetum ssuviatile , Carex spp., 10.vii.2014, 2 ♂♂ 4 ♀♀, 11.vii.2014, 1 ♂ 1 ♀; Moosonee, 51°16.55'N 80°39.01'W, sweeps, mostly Carex spp., wet forest trail, 11.vii.2014, 1 ♀; Moosonee, 51°16.63'N 80°38.87'W, sweeps, Calamagrostis , Carex , drier edge of sedge meadow, 9.vii.2014, 2 ♂♂ 2 ♀♀; Moosonee, 51°16.68'N 80°38.65'W, sweeps, mostly C [arex] utriculata , C. aquatilis , wet sedge meadow, 10.vii.2014, 1 ♂ 3 ♀♀; Moosonee, 51°16.99'N 80°38.37'W, sweeps, mostly Equisetum ssuviatile , Carex spp., wet sedge meadow, 10.vii.2014, 1 ♂; ~ 8 km SSW Nipigon, Hwy #628, 48°57.09'N 88°19.71'W, sweeps, damp roadside, mixed graminoids, 16.vii.2008, 1 ♀, all K. N. Barber leg. (all CNCI); One-sided [= Caliper] Lake, 27.vi.1960, 1 ♂, Kelton & Whitney leg.; Ottawa, 14.ix.1955, 1 ♀, 12.vii.1956, 1 ♂, J. R. Vockeroth leg. (all CNCI); Otter Rapids, 50°10.80'N 81°38.59'W, sweeps, Carex sp., 19.vii.2009, 1 ♀, sweeps, roadside Equisetum spp., 19.vii.2009, 1 ♂; Otter Rapids, 50°10.87'N 81°38.56'W, sweeps, Calamagrostis canadensis in hydro cut, 20.vii.2009, 1 ♂ 1 ♀; ~ 7 km SSE Otter Rapids, 50°07.52'N 81°35.92'W, sweeps, Scirpus sp., 20.vii.2009, 2 ♂♂ 1 ♀, all K. N. Barber leg. (all CNCI); Pancake Bay Prov. Park, 46°58.11'N 84°42.72'W, sweeping from boardwalk, mostly emergent sedges / Equisetum , 9.vii.2010, 7 ♂♂ 1 ♀, J. Roháček leg. ( SMOC); Pancake Bay P.Pk., 46°58.11'N 84°42.72'W, sweeps from boardwalk, mostly emergent sedges / Equisetum , 17.vii.2004, 1 ♂ ( DEBU 01500322), 24.vii.2004, 2 ♂♂ ( DEBU 01500573, -74), 7.viii.2004, 2 ♂♂ 1 ♀ ( DEBU 01501085–87), 27.vi.2005, 2 ♀♀ ( DEBU 01501608, -09); Pancake Bay P. Pk., 46°58.12'N 84°42.75'W, sweeps, mostly graminoids/ Typha , near wetland boardwalk, 2.viii.2004, 4 ♂♂ 1 ♀ ( DEBU 01500966–70), all K. N. Barber leg.; Hwy 101 at Prairie Bee River (west side bridge), 47°51.81'N 83°54.33'W, sweeps, mostly Carex utriculata , 14.vii.2013, 3 ♂♂ 1 ♀; ~ 4.5 km E Rosseau, on Aspdin Rd., 45°15.88'N 79°34.88'W, sweeps, mostly Carex in sedge meadow, 7.vii.2005, 1 ♂ 1 ♀, all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, Algoma U[niversity] College, sweeps, 20.vii.1987, 2 ♂♂ 1 ♀; S[ault] S[ainte] Marie, S. of Algoma U[niversity] College, 46°29.9'N 84°17.2'W,sweeps, Phalaris arundinacea , 12.vii.1997, 1 ♂ 2 ♀♀, 22.vi.1998, 2 ♂♂, sweeps, trailside Carex / Scirpus , 14.vii.1998, 1 ♂ 1 ♀, sweeps, Carex lacustris , 12.vii.1997, 3 ♂♂ 2 ♀♀, sweeps, Typha , 11.vi.1997, 1 ♀, sweeps/pooter, grasses under Populus / Betula , 11.vii.1997, 1 ♀, sweeps, grassy edge of woods, 11.vii.1997, 1 ♂, sweeps, graminoids mostly Carex aquatilis , 11.vi.1997, 5 ♂♂ 9 ♀♀, sweeps/pooter, Calamagrostis canadensis , 12.vii.1997, 14 ♂♂ 29 ♀♀, 3.viii.1997, 2 ♂♂ 2 ♀♀ (all CNCI), 26.viii.1997, 12 ♂♂ 11 ♀♀ ( USNM), 12.vii.2002, 2 ♂♂ 1 ♀, sweeps, Calamagrostis canadensis , 28.viii.1997, 7 ♂♂ 1 ♀, 29.viii.1997, 56 ♂♂ 5 ♀♀, 22.vi.1998, 3 ♂♂ 9 ♀♀ ( CNCI), 26.vi.1998, 31 ♂♂ 41 ♀♀ ( BYUC, CLEV, CMNH, CSCA, CSUC, EMEC, KNWR, MCZN, MEMU 3 ♂♂ 4 ♀♀ each; BDUC 4 ♂♂ 5 ♀♀), 29.vii.1999, 2 ♂♂, 31.vii.2002, 3 ♀♀, sweeps, mostly Calamagrostis canadensis , 28.vii.2001, 6 ♂♂ 1 ♀, 29.vii.2001, 1 ♂ 2 ♀♀, 6.viii.2001, 7 ♂♂, 4.viii.2002, 4 ♂♂ 1 ♀, sweeps, Calamagrostis canadensis & Carex aquatilis , 23.vii.1997, 16 ♂♂ 3 ♀♀, 28.viii.1997, 2 ♂♂ 2 ♀♀ ( CNCI), sweeps, Carex / Calamagrostis , 3.viii.2002, 5 ♂♂ 5 ♀♀ ( CASC), 10.viii.2002, 1 ♂ 1 ♀, 5.ix.2002, 1 ♀, pooter, Carex / Calamagrostis , 10.viii.2002, 1 ♀ ( CNCI), all K. N. Barber leg.; same locality but sweeps, Carex / Calamagrostis , used in lab rearing, [with various dates of death], 6.viii.2002, 3 ♂♂ 9 ♀♀; same locality but lab-reared, Phalaris arundinaceae , from 3 ♂♂ 9 ♀♀, 46°29.9'N 84°17.2'W, sweeps, Carex / Calamagrostis , 6.viii.2002, 20°C, L:D 16:8, 50–70% RH, moist 3 cm sections P. arundinaceae in 5 cm plates, oviposition: 15–21.viii.2002, puparium: 21.ix.2002, adult: 4.x.2002, 1 ♀, all K. N. Barber leg. (all CNCI); same locality but sweeps/pooter, Carex aquatilis , 3.viii.1997, 1 ♂ 7 ♀♀, sweeps, Carex aquatilis , 12.vii.1997, 15 ♂♂ 27 ♀♀, 3.viii.1997, 2 ♂♂ 4 ♀♀ ( CNCI), 22.vi.1998, 3 ♂♂ 3 ♀♀ ( NMPC), 26.vi.1998, 7 ♂♂ 13 ♀♀, 29.vii.1998, 1 ♂, 28.vi.2002, 1 ♂ 4 ♀♀, 29.vi.2002, 5 ♂♂, 1.vii.2002, 2 ♂♂ 3 ♀♀, 9.vii.2002, 2 ♀♀ ( CNCI), 12.vii.2002, 6 ♂♂ 8 ♀♀ ( LACM), 31.vii.2002, 6 ♂♂ 5 ♀♀ ( AMNH), sweeps, mostly Carex aquatilis , 13.vi.2001, 1 ♂, 14.vi.2001, 1 ♂ 1 ♀, 17.vi.2001, 1 ♀, 21.vi.2001, 7 ♂♂ 7 ♀♀ (incl. pair in copula), 22.vi.2001, 1 ♂ ( CNCI), 25.vi.2001, 26 ♂♂ 17 ♀♀ ( CNCI 24 ♂♂ 15 ♀♀, SMOC 2 ♂♂ 2 ♀♀), 28.vi.2001, 1 ♂ 3 ♀♀, 11.vii.2001, 17 ♂♂ 5 ♀♀ (incl. 3 pairs in copula), 15.vii.2001, 9 ♂♂ 7 ♀♀ (1 ♂ wing illustration) ( CNCI), 17.vii.2001, 35 ♂♂ 27 ♀♀ ( MTEC, OSAC, PMAE, RBCM, SEMC, UBCZ, UCRC, UGCA 4 ♂♂ 3 ♀♀ each; WFBM 3 ♂♂ 3 ♀♀), 18.vii.2001, 2 ♂♂ 6 ♀♀, 21.vii.2001, 3 ♂♂ 1 ♀, 28.vii.2001, 6 ♂♂ 7 ♀♀, 29.vii.2001, 1 ♂ 2 ♀♀, 2.viii.2002, 4 ♂♂ 9 ♀♀, 4.viii.2002 ( CNCI), 5 ♂♂ 3 ♀♀ ( INHS), sweeps, mixed graminoids, 18.vi.1998, 5 ♂♂ 2 ♀♀ ( CNCI), all K. N. Barber leg.; same locality but 46°29.88'N 84°17.19'W, sweeps, Phalaris arundinacea , 18.vi.2005, 1 ♀, sweeps, Calamagrostis canadensis , 18.vii.2004, 4 ♂♂, 26.viii.2003, 1 ♂ 2 ♀♀, sweeps, Carex aquatilis , 1.vii.2003, 1 ♂ 3 ♀♀, 26.viii.2003, 7 ♂♂ 6 ♀♀ (incl. pair in copula), 18.vi.2005, 2 ♀♀, 21.vii.2005, 6 ♂♂ 3 ♀♀ ( CNCI), 27.vii.2005, 6 ♂♂ 4 ♀♀ ( LEMQ), sweeps, mostly Carex aquatilis , 18.vii.2004, 7 ♂♂ 15 ♀♀, sweeps, trampled graminoids, mostly Carex aquatilis , 5.ix.2004, 3 ♀♀, 1.viii.2005, 8 ♂♂ 12 ♀♀, pooter, mostly Carex aquatilis , 21.viii.2004, 2 ♂♂ 1 ♀ ( CNCI); same locality but 46°29.82'N 84°17.17'W, sweeps, mostly Carex aquatilis near river, 21.vii.2005, 5 ♂♂ 2 ♀♀ ( CNCI), all K. N. Barber leg.; S[ault] S[ainte] Marie, S. of Algoma University, 46°29.88'N 84°17.19'W, sweeps, mostly Carex aquatilis , 29.vi.2008, 1 ♂ 3 ♀♀, sweeps, Carex aquatilis , Calamagrostis canadensis , 5.vii.2008, 3 ♂♂ 3 ♀♀, sweeps, Carex aquatilis , Typha latifolia , 6.viii.2008, 5 ♂♂ 3 ♀♀; same locality but 46°29.81'N 84°17.15'W, sweeps, Carex aquatilis , Calamagrostis canadensis (by river), 5.vii.2008, 2 ♂♂ 2 ♀♀, all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, Baseline Rd., 46°31.40'N 84°24.40'W, Malaise #1, Aster [ Doellingeria ], Rubus , Equisetum , Carex , Solidago , in aspen clearing, 18–29.vii.2005, 1 ♀, sweeps, Thalictrum , Rubus , Equisetum , Carex , ferns under aspen, 25.vi.2005, 1 ♀; same locality but 46°31.40'N 84°24.45'W, sweeps, edge of forest, Solidago , Rubus , Equisetum , grasses, 25.vi.2005, 2 ♂♂, 29.vi.2005, 1 ♂ 1 ♀; same locality but w. of creek, 46°31.52'N 84°24.63'W, sweeps, graminoids on path, 22.vii.2005, 8 ♂♂ 2 ♀♀, sweeps, Carex under ash / aspen, 22.vii.2005, 2 ♂♂ 1 ♀; same locality but 46°31.61'N 84°24.68'W, sweeps, Carex edge of alder thicket, 22.vii.2005, 5 ♂♂ 3 ♀♀, sweeps, graminoids/ composites, 20.viii.2008, 4 ♂♂ 1 ♀, all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, Bellevue Pk., 46°30.1'N 84°18.1'W, sweeps, mostly Calamagrostis , 7–8.vii.2000, 2 ♀♀; S[ault] S[ainte] Marie, Birchwood Pk., mixed forest, 15.vi.1986, 1 ♂, 28.vi.1986, 13 ♂♂ 5 ♀♀, 13.viii.1986, 1 ♀, all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, Bristol Pl. Pk., 46°30.8'N 84°16.6'W, sweeps, Phalaris arundinacea , 18.vii.1997, 1 ♀, 28.vi.1998, 7 ♂♂ 17 ♀♀, sweeps/pooter, Phalaris arundinacea , 1.viii.1997, 1 ♀, 8.viii.1997, 1 ♂ 2 ♀♀; same locality but 46°30.77'N 84°16.66'W, sweeps, Phalaris arundinacea under Populus , 7.viii.2008, 1 ♂, all K. N. Barber leg. (all CNCI); Sault Ste. Marie, Finn Hill, 46°31.48'N 84°17.36'W, sweeping graminoid vegetation, 7.vii.2010, 7 ♂♂ 4 ♀♀, J. Roháček leg. ( SMOC, 1 ♀ genit. prep., 1 ♂ 1 ♀ photographed); S[ault] S[ainte] Marie, Finn Hill, 46°31.48'N 84°17.36'W, sweeps, mostly Scirpus microcarpus , 18.vi.2005, 1 ♂, 13.viii.2005, 2 ♂♂ 2 ♀♀, 14. viii.2005, 14 ♂♂ 8 ♀♀, 16.viii.2005, 2 ♂♂ 1 ♀, 25.viii.2005, 2 ♂♂, 12.ix.2005, 1 ♀, 16.ix.2005, 1 ♂, 8.vii.2006, 4 ♂♂ 3 ♀♀; same locality but 46°31.63'N 84°17.29'W, sweeps, mostly Scirpus microcarpus , 8.vii.2006, 1 ♂ 1 ♀; same locality but 46°31.6'N 84°17.3'W, sweeps, graminoids/ composites, 19.vii.2005, 3 ♀♀; same locality but 46°31.60'N 84°17.30'W, pooter, Carex stipata stipata , 27.vii.2004, 1 ♂ 1 ♀; same locality but 46°31.63'N 84°17.33'W, sweeps, Carex stipata stipata , 19.vii.2004, 1 ♂ 3 ♀♀, 20.vii.2004, 3 ♂♂ 2 ♀♀, 1.viii.2005, 1 ♀, pooter, Carex stipata stipata , 25.vii.2004, 2 ♀♀, sweeps, Calamagrostis canadensis , 26.vi.2007, 4 ♂♂ 1 ♀, sweeps, mostly Calamagrostis canadensis , 6.vii.2008, 2 ♂♂ 1 ♀, sweeps, mostly Carex stipata stipata , 26.vi.2007, 3 ♂♂ 2 ♀♀, sweeps, mostly Carex / Calamagrostis , edge of Populus tremuloides , 25.vi.2009, 2 ♂♂ 2 ♀♀, all K. N. Barber leg. (all CNCI), sweeping boggy meadows, mostly Carex stipata stipata , 7.vii.2010, 12 ♂♂ 18 ♀♀ (1 ♂ 3 ♀♀ genit. prep.), 12.vii.2010, 2 ♂♂, J. Roháček leg. ( SMOC); same locality but 46°31.66'N 84°17.34'W, sweeps, mostly Calamagrostis canadensis , 8.viii.2008, 1 ♂ 1 ♀, 10.viii.2008, 1 ♀ ( CNCI); same locality but 46°31.67'N 84°17.32'W, sweeps, Calamagrostis canadensis , 20.vii.2004, 3 ♂♂ 3 ♀♀ ( CNCI 1 ♂ 1 ♀, SMOC 2 ♂♂ 2 ♀♀), 30.vii.2004, 1 ♂ 1 ♀ ( CNCI); same locality but 46°31.7'N 84°17.5'W, sweeps, mostly sedges in trail, Calamagrostis canadensis , 4.vii.2002, 5 ♂♂ 1 ♀, 7.vii.2002, 1 ♂ 3 ♀♀ (incl. pair in copula) ( CNCI), all K. N. Barber leg.; Sault Ste. Marie, Fish Hatchery Road, near Coldwater Creek, 46°34.29'N 84°17.21'W, sweeping graminoids, Impatiens , 9.vii.2010, 7 ♂♂ 2 ♀♀, J. Roháček leg. ( SMOC, 1 ♂ 1 ♀ genit. prep.); S[ault] S[ainte] Marie, Ft. Creek Cons[ervation] Area, 46°32.5'N 84°20.8'W, sweeps/pooter, Phalaris arundinacea , 9.viii.1997, 1 ♂, sweeps, Scirpus sp., 9.viii.1997, 1 ♀; S[ault] S[ainte] Marie, Hwy #17 city limits, 46°36.58'N 84°17.83'W, sweeps, mostly Carex / Calamagrostis in wet area, 16.viii.2004, 1 ♂ 5 ♀♀, 23.viii.2004, 2 ♂♂ 3 ♀♀, sweeps, mostly Carex / Calamagrostis in wet area, 19.vii.2008, 1 ♂ 2 ♀♀, 13.viii.2008, 1 ♂ 2 ♀♀, sweeps, Calamagrostis canadensis in wet area, 16.viii.2004, 6 ♂♂ 1 ♀; same locality but 46°36.62'N 84°17.91'W, sweeps, mostly riparian sedges, 16.viii.2004, 2 ♂♂ 5 ♀♀, all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, Kinsmen Pk., 46°35.7'N 84°16.7'W, sweeps, mostly Carex , 11.viii.2002, 1 ♀; S[ault] S[ainte] Marie, Landslide Rd., Coldwater Ck. noodplain, 46°33.8'N 84°16.6'W, sweeps, mostly Carex sp., 7.viii.1997, 1 ♂ 3 ♀♀, sweeps/pooter, Calamagrostis canadensis , 7.viii.1997, 2 ♂♂ 1 ♀; same locality but 46°33.94'N 84°16.66'W, sweeps, graminoids in wet ditch, 26.viii.2008, 2 ♂♂ 1 ♀, all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, Sault Coll[ege] Outdoor Lab, 46°32.1'N 84°18.2'W, sweeps, Carex sp., 25.vii.1997, 4 ♂♂ 2 ♀♀, sweeps, Phalaris arundinacea , 25.vii.1997, 1 ♂, sweeps, trailside grasses, Acer / Betula , 25.vii.1997, 1 ♂, sweeps, short trail grasses, Acer / Betula , 25.vii.1997, 2 ♂♂ 4 ♀♀, sweeps, graminoids in open, 25.vii.1997, 1 ♀, sweeps, Carex in clearing, 7.ix.1997, 1 ♂ 1 ♀; same locality but 46°32.08'N 84°18.21'W, sweeps, Carex lacustris in opening, 17.vii.2005, 1 ♂, 2.viii.2005, 1 ♀, sweeps, Scirpus sp. in opening, 17.vii.2005, 1 ♂; S[ault] S[ainte] Marie, hydro cut nr. Sault Coll[ege] Outdoor Lab, 46°32.1'N 84°18.0'W, sweeps, mostly sedges, 10.vii.2002, 1 ♂ 4 ♀♀, sweeps, Scirpus cyperinus , 24.viii.2005, 2 ♂♂, 12.viii.2008, 1 ♂ 3 ♀♀, all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, 2 nd Line E, 46°32.3'N 84°16.6'W, sweeps, Calamagrostis canadensis , 26.vi.1999, 2 ♂♂ 2 ♀♀, sweeps, graminoids in open meadow, 26.vi.1999, 1 ♂; S[ault] S[ainte] Marie, Thayers Acres, 46°35.54'N 84°15.53'W, sweeps, emergent Carex sp., 1.vii.2007, 1 ♀, all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, Voyageur Trail, 46°35.48'N 84°15.23'W, sweeps, Calamagrostis canadensis , 9.vii.2006, 1 ♂, [K. N. Barber] leg.; S[ault] S[ainte] Marie, Whitensh Is./St. Mary’s Is., 46°30.68'N 84°21.20'W, sweeps, riparian graminoids, 8.viii.2004, 5 ♂♂ 7 ♀♀, K. N. Barber leg. (all CNCI); ~ 10 km W S[ault] S[ainte] Marie, Airport Rd., 46°29.9'N 84°28.9'W, natural gas r[igh]t-of-way, sweeps, graminoids, composites, Equisetum , Rubus , ferns, 25.viii.2008, 2 ♂♂ 2 ♀♀; same locality but 46°29.72'N 84°28.96'W, natural gas r[igh]t-of-way, sweeps, graminoids, Equisetum , herbs, 5.viii.2009, 2 ♂♂, sweeps,mostly ferns, graminoids, 5.viii.2009, 3 ♂♂ 5 ♀♀, sweeps, mostly Scirpus / Calamagrostis , 4.viii.2004, 1 ♂, all K. N. Barber leg. (all CNCI); same locality but sweeping graminoids, composites, Equisetum , Rubus , ferns, 12.vii.2010, 6 ♂♂ 7 ♀♀, J. Roháček leg. ( SMOC); NW S[ault] S[ainte] Marie, Red Rock, 46°36.21'N 84°32.85'W, sweeps, graminoids, ferns, Impatiens , 4.viii.2006, 2 ♂♂ 1 ♀; Searchmont, N Hwy #552, 46°50.3'N 84°04.4'W, sweeps, roadside sedges / grasses, 10.vii.1998, 2 ♀♀; ~ 5 km SE Searchmont, km 6.2 Ranger Lk. Rd., 46°45.52'N 83°59.51'W, sweeps, sedges at beaver dam outnow, 23.vi.2007, 1 ♀, all K. N. Barber leg.; 12.4 km NNE Searchmont, mi. 10 Whitman Dam Rd., herb/grass meadow by Goulais R., 23.vi.1986, 1 ♂ 1 ♀, 25.vi.1986, 1 ♂ 2 ♀♀, 3.vii.1986, 1 ♀, 9.vii.1986, 1 ♂, D. J. M. Harvey leg.; ~ 18 km NNE Searchmont, Goulais River WS [watershed], ~mi. 15 Whitman Dam Rd., 46°55.3'N 83°56.3'W, sweeps, grass under jackpine, 30.vii.1999, 1 ♀, K. N. Barber leg. (all CNCI); S of Sioux Narrows, sweeping sedge at beaver dam, 10.vii.1992, 2 ♂♂ 3 ♀♀, T. A. Wheeler leg. ( LEMQ 0039856, -60, -61, -64, -82); ~ 21 km NNE Smooth Rock Falls, 49°20.91'N 81°32.01'W, sweeps, ditchside Equisetum spp., grasses, herbs, 19.vii.2009, 1 ♂ 2 ♀♀, 8.vii.2012, 1 ♀; 2 km E Sowerby, Hwy 17 @ Harris Ck. Rd., 46°17.6'N 83°21.3'W, sweeps, noodplain grasses, 1.viii.1997, 1 ♂; 0.4 km S Stonecliffe, Pine Valley Rd. @ RR crossing, 46°11.9'N 77°52.8'W, sweeps, graminoids, 9.vii.2001, 5 ♀♀, all K. N. Barber leg.; Swastika, 7.vii.1987, 1 ♀, J. R. Vockeroth leg. (all CNCI); ~ 74 km NNE Thessalon, 46°53.94'N 83°16.23'W, shore of Mississagi River, sweeping graminoids and Equisetum spp. on muddy shore, 5.vii.2010, 2 ♂♂ 5 ♀♀, J. Roháček leg. ( SMOC); same locality but sweeps, graminoids, herbs, Equisetum spp., 5.vii.2010, 2 ♂♂ 5 ♀♀, sweeps, graminoids, herbs, Equisetum spp., 17.vii.2010, 6 ♂♂ 5 ♀♀, sweeps, graminoids, herbs, Equisetum spp., 12.ix.2010, 1 ♂, K. N. Barber leg. ( CNCI); ~ 92 km NNE Thessalon, nr. Mountain Ash Lake, 47°02.98'N 83°10.88'W, sweeping Carex aquatilis on edge of wetland, 4.vii.2010, 6 ♂♂ 7 ♀♀, J. Roháček leg. ( SMOC); ~ 29 km SW Timmins, 48°19.12'N 81°44.79'W, sweeps, mostly Carex spp./ Calamagrostis , 18.vii.2009, 2 ♂♂; ~ 34 km N Timmins, 48°45.88'N 81°21.71'W, sweeps, Carex spp., 18.vii.2009, 5 ♂♂ 1 ♀; ~ 25 km W Upsala, 49°08.83'N 90°48.20'W, sweeps, wet area, mostly Calamagrostis canadensis , 30.vii.2008, 3 ♂♂ 3 ♀♀, all K. N. Barber leg. (all CNCI); Waterloo Reg., Blair, RARE, Blair Trail, 43°22'38"N 80°20'34"W, 20.vi.2006, 3 ♂♂ 4 ♀♀, M. D. Bergeron leg. ( DEBU 00271089, -090, -093–95, -106, -100); Waubamick [sic Waubamik], [-]. vii.1915, 2 ♂♂, H. S. Parish leg. ( USNM). PRINCE EDWARD ISLAND: Eglington Bay, sweep vegetation, 20.vii.1996, 1 ♂, N. deVille leg. ( LEMQ 0039852). QUEBEC: Beech Grove, 7.vi.1955, 1 ♀, J. F. McAlpine leg.; Beechgrove, 45°39'N 76°08'W, 29.vi.1962, 2 ♂♂ 1 ♀, 27.vi.1984, 1 ♀, 5.vii.1984, 1 ♀, 24.vi.1988, 3 ♂♂ 1 ♀, all J. R. Vockeroth leg.; Papineau Co., Buckingham Twp., 1.viii.1962, 1 ♀, L. K. Smith leg.; Cascapedia, 2.viii.1954, 1 ♀, J. E. H. Martin leg. (all CNCI); Chemin-du-Lac Bog, 47°45'38.3"N 69°31'43.2"W, sweeping, natural site, T1, 30.vi.2006, 1 ♂, A. G. Taillefer leg. ( LEMQ 0040320); Eardley, 23. vii.1968, 1 ♀, P. Ward leg. ( CNCI); Gaspé, Forillon N. P., Ruisseau Castor, 48°54'N 64°21'W, sweep beaver dam, 6.viii.2000, 1 ♂, H.Varady-Szabo leg. ( LEMQ 0039874); Gaspé, Forillon N. P., Cap des Rosiers, 48°50'N 64°12'W, sweep grass, 6.viii.2000, 1 ♂ 1 ♀, H. Varady-Szabo leg. ( LEMQ 0039858, -63); Gaspesie, near Percé, 48°37'N 64°11'W, sweep, 1.viii.2001, 1 ♂, V. Dion & S. Boucher leg. ( LEMQ 0039850); Gatineau Co., Masham Township, 27.vii.1995, 2 ♂♂, E. Ikeda leg. ( LEMQ 0039853, -54); Godbout, dunes near ferry dock, swept, eclector, (Universität Bielefeld, X 988), 24.viii.1994, 2 ♂♂, M. v. Tschirnhaus leg. ( ZSMC, in ethanol); Harrington L., Gatineau Pk., 3.vii.1963, 2 ♂♂ 3 ♀♀, J. R. Vockeroth leg.; Hull, nr. Fairy Lake, 27.vii.1959, 1 ♂, L. K. Smith leg. (all CNCI); Île Bonaventure, 48°30'N 64°10'W, 3 km from Côte de Percé, sweep grass, 28.vii.2000, 1 ♀, A. Thibault leg. ( LEMQ 0039859); Îles de la Madeleine, Île de la Grand[e] Entrée, Chemin du Bassin Ouest, 47°32.96'N 61°32.64'W, sweep neld along road, 9.vii.2004, 2 ♂♂ 1 ♀, S. Boucher leg. ( LEMQ 0040500, -502, -503); Îles Penchées, Les Escoumins, 48°22'N 69°22'W, sweep Juncus , Lathyrus on sand beach, 1.viii.2001, 1 ♂, M. Giroux leg. ( LEMQ 0039851); Lac Phillipe, 45°37'N 76°W, 7.vii.1968, 2 ♂♂, J. R. Vockeroth leg.( CNCI); Lac Roddic, 16 km S Maniwaki, 22.vi.1991, 1 ♀, M. Barták leg. ( MBPC, genit. prep.); Laurentide Pk., 7.viii.1956, 2 ♂♂, A. H. Sturtevant leg.; Mantane [sic Matane], 12.viiii.1967, 1 ♂ (gent. prep.),A. Stone leg. (all USNM); Newport, 14.viii.1983, 3 ♂♂ 1 ♀, B. M. Bissett leg.; Kam[ouraska] Co., Parke Reserve, sweeping Kalmia angustifolia , 11.vii.1957, 1 ♀, G. E. Shewell leg. (all CNCI); Percé, Bridgeville, 48°36'N 64°19'W, sweep vegetation in peat bog, 8.viii.2000, 1 ♀, H. Varady-Szabo leg. ( LEMQ 0039857); Percé, Pointe-St.-Pierre, 48°37'N 64°10'W, sweep vegetation, 8.viii.2000, 2 ♂♂, H. VaradySzabo leg. ( LEMQ 0039849, -62); Québec [City], 30.vii.1952, 1 ♂, G. Steyskal leg.; “Quebec CAN”, 5.viii.1930, 1 ♂, A. L. Melander leg. (both USNM). SASKATCHEWAN: Assiniboia, 27.vi.1955, 1 ♀, J. R. Vockeroth leg.; Beaver Creek Cons. Area, ~ 13 km S Saskatoon, 51°58.6'N 106°43.0'W, sweeps, mostly grasses under Betula / Populus , 12.vii.1999, 1 ♀, K. N. Barber leg.; Christopher Lake, 11.vii.1959, 1 ♂ 2 ♀♀, A. & J. Brooks leg. (all CNCI). UNITED STATES OF AMERICA: COLORADO: 3.5 mi S Florissant, Sanborn Ranch, swept from aspen grove nr. girls camp, 17.vii.2000, 1 ♀, B. A. Foote leg. ( CMNH). IDAHO: Bonner Co., 6.5 mi W Granite Lake, Cirsium arvense , 30.vi.1976, 1 ♀, S. T. Rose leg. ( WFBM); Priest Lake, 1.viii.1916, 1 ♀, A. L. Melander leg. ( USNM). MICHIGAN: Houghton Co., 28.vi.1955, 1 ♀, R. R. Dreisbach leg., 20.vi.1960, 1 ♀, R. & K. Dreisbach leg.; Keweenaw Co., 26.vi.1955, 1 ♀, R. R. Dreisbach leg., 22.viii.1959, 1 ♀, R. & K. Dreisbach leg., 27.vii.1953, 1 ♂, G. Steyskal leg.; Keweenaw Co., Isle Royale, 5., 8.vii.1938, 1 ♀, 15., 17.vii.1938, 1 ♂, G. Steyskal leg.; Menominee Co., 29.viii.1952, 1 ♀, R. R. Dreisbach leg.; Newaygo Co., 27.vi.1953, 1 ♂, R. R. Dreisbach leg.; St. Ignace, 31. viii.1954, 1 ♂, R. L. Fischer leg. (all USNM). MINNESOTA: Eaglesnest, 13.vii.1957, 1 ♀, W. V. Balduf leg. ( INHS 40,176); Aitkin Co., 14 mi W Willow River, 46.334°N 93.096°W, 31.vi.1994, 1 ♂, 27.viii.1994, 2 ♀♀, 26.vii.1995, 1 ♀, 22.vi.1997, 1 ♀, 5.vii.1997, 1 ♂, D. E. Hansen leg. ( CNCI). MONTANA: 15 mi S Big Fork, 4.vii.1967, 2 ♂♂; 4 mi E Big Fork, 11.viii.1967, 1 ♀; 15.0 mi NE Polson, 2.vii.1966, 1 ♂; 20 mi S Swan Lake, 14.vii.1967, 1 ♀, all B. A. Foote leg.; 20 mi N Whitensh, 31.vi.1966, 2 ♂♂ 1 ♀, T.Krystowski leg. (all USNM). NEW HAMPSHIRE: Kinsman Notch, 7.vii.1931, 1 ♂, J. M.Aldrich leg. ( USNM). NEW YORK: St. Lawrence Co., Colton, Raquette R., 24.vii.1963, 1 ♂, W. W. Wirth leg. ( USNM); St. Lawrence Co., ~ 25 km SE S.Colton, 44°19.07'N 74°43.42'W, sweeps, mostly Carex , edge of wetland, 27.vii.2006, 1 ♀, K. N. Barber leg.; St. Lawrence Co., Huggards, 4.viii.1957, 2 ♀♀, J. R. Vockeroth leg. (all CNCI); Clinton Co., Saranac, 17.vii.1952, 1 ♀, W. A. MacDonald leg. ( USNM). OREGON: Deschutes Co., Bend, 8.viii.1951, 1 ♂, M. R.Wheeler leg. ( AMNH, genit. prep.). WASHINGTON: Olympic Nat. Park, Kalaloch, 8.vii.1968, 1 ♀, W. W. Wirth leg. ( USNM); Pierce Co., Tacoma, 12.viii.1982, 1 ♀, T. L. Whitworth leg. ( LACM). WISCONSIN: Washburn Co., T39N R12W B32, 25.vi.1953, 1 ♂ 2 ♀♀, 3.vii.1953, 1 ♀, 4.vii.1953, 1 ♂ 1 ♀, 24.vii.1953, 1 ♂, 30.viii.1953, 1 ♀, T39N R12W B33, 20.vi.1953, 1 ♂, T39N R12W B33, 29.vi.1953, 1 ♂, T39N R13W B30, 4.vii.1953, 1 ♂, R. H. Jones leg. ( USNM).

Other material examined (not included in type series). CANADA: ONTARIO: Bruce Peninsula N. P., bank of Crane River nr. Hwy 6, 45°08.9'N 81°28.1'W, sweeps, grasses / sedges, 31.vii.1997, 1 ♂, S. A. Marshall leg. ( DEBU, head & thorax crushed); Greenwater P. Pk., 49°10.91'N 81°16.28'W, sweeps, Carex , Calamagrostis , Phalaris , in creek noodplain, 21.vii.2009, 1 ♂, K. N. Barber leg. ( DEBU 01502146, deformed T 4 & T 5); Ottawa, 13.vii.1963, 1 ♀, J. R.Vockeroth leg. ( CNCI, headless); Sault Ste. Marie, Finn Hill, 46°31.63'N 84°17.33'W, sweeping boggy meadows, mostly Carex stipata stipata , 7.vii.2010, 4 ♂♂, J. Roháček leg. ( SMOC, 3 ♂♂ used for molecular work, other ♂ headless). QUEBEC: Québec [City], 30.vii.1952, 1 ♂, G. Steyskal leg. ( USNM, abdomen missing). UNITED STATES OF AMERICA: MASSACHUSETTS: Woods Hole, [-]. vii.1918, 1 ♀, A. H. Sturtevant leg. ( USNM, wings missing).

Description. Male. Total body length 2.46–3.02 mm; general colour closely resembling that of A. dissors Collin, 1944 , thus largely yellow but with highly variable brown to greyish brown darkenings on mesonotum and preabdominal terga; pleural part of thorax completely yellow and epandrium always blackish brown ( Figs 3 , 277 ). Head about as long as high, anteriorly slightly angular in pronle with face slightly receding, normally entirely yellow except for brownish ocellar triangle, very rarely (in darkest specimens) with lateral areas of occiput somewhat brownish darkened. Occiput slightly concave, laterally usually slightly darker yellow (rarely brownish, see above) than medially; in lighter yellow medial area with two divergent elongate spots of silvery white microtomentum meeting above foramen. Frons relatively narrow, yellow and largely dull, only ocellar triangle brown; frontal triangle with sparse but distinct silvery golden glittering microtomentum. Orbits lighter yellow, with sparse silvery whitish microtomentum being attenuated posteriorly (usually behind middle ors). Frontal triangle relatively narrow, reaching to anterior third of frons. Frontal lunule small but distinct, pale yellow. Face moderately narrow, medially slightly concave, yellow to whitish yellow and separated from parafacialia by distinct golden-orange marginal stripe also reaching onto ventral margin of gena; parafacialia and gena whitish yellow, with silvery white microtomentum; postgena and mouthparts pale yellow. Cephalic chaetotaxy (all setae black): pvt relatively long and strongly crossed; oc (usually) or vti (rarely) longest of cephalic setae but oc can rarely be shortened and weak; vte usually shorter but sometimes almost as long as vti; posterior ors almost as long as vte; 3 ors, 2 posterior long (middle ors slightly shorter than posterior ors) and 1 shorter anterior ors setula (this sometimes unusually strong and longer than half length of middle ors); 2–3 pairs of medial microsetulae in anterior third of frons; 1 small setula behind vte; postocular setulae (only 7–8) short, situated mainly dorsally; postgena with several setulae and 2 short posteroventral setae; 1 long vi (as long as middle ors), subvibrissa reduced, slightly longer than peristomals; only 5–6 relatively long but nne peristomal setulae. Palpus yellow, with 1 distinct dark ventral preapical seta and a series of 9–10 setulae subapically and ventrally. Eye broadly ovoid, wider anteroventrally, with longest diameter oblique and about 1.3 times as long as shortest. Shortest genal height 0.14–0.15 times as long as shortest eye diameter. Antenna strongly geniculate, entirely yellow; 1st nagellomere with short white pilosity. Arista with basal segments ochreous and distal setiform part blackish brown, about 1.9–2.0 times as long as antenna, short-ciliate (cilia shorter than those on 1st nagellomere).

Thorax slightly narrower than head. Scutum with colouration ranging from largely yellow with faint ochreous brown darkenings anteriorly between dc and prs-sa lines and/or medially (between dc lines) in front of scutellum, through to distinctly brown except for yellow medial area in anterior half and humeral-notopleural area to almost completely dark greyish brown with only humeral-notopleural area (this always) yellow. Scutellum always more or less dark, ochreous brown to dark brown. Dorsum of thorax grey microtomentose and relatively dull. Pleural part of thorax more shining than scutum, entirely yellow (no darker dorsal stripe in contrast to A. dissors ). Postscutellum and postnotum pale to dark brown, the latter sometimes yellow in dorsal half. Thoracic chaetotaxy: 1 hu (longer than posterior npl) plus 1–3 hu setulae; 2 npl (anterior distinctly longer than hu, posterior shorter); 1 relatively weak prs (about as long as or shorter than hu); sa and pa usually slightly longer than prs; 2 long postsutural dc (the smaller anterior seta about as long as anterior npl, posterior longest of thoracic setae) and 5–6 dc microsetae in front of them (the hindmost distinctly enlarged); 4 (rarely only 3) rows of ac microsetae on suture but only 2 rows between dc with hindmost ac pair usually situated somewhat beyond level of posterior dc; 2 sc, laterobasal weak (as long as or shorter than sa), apical sc almost as long as posterior dc; 1 small, pale and hair-like ppl; 2 long stpl (anterior usually slightly shorter) and 5–6 upcurved setulae in dorsal half of sternopleuron (sometimes with 1 setula in front of anterior stpl in addition); ventral part of sternopleuron with a cluster of 6–7 longer setae. Scutellum rounded triangular, very slightly convex dorsally. Legs pale yellow, only distal half to three-fourths of last tarsal segment of all tarsi dark brown. f 1 with ctenidial spine not very strong, usually only slightly longer than maximum width of t 1 and with a number of long but nne setae in posterodorsal and posteroventral rows. f 2 simply setose; f 3 with a row of posteroventral setae, 5–7 of them in distal half shortened and distinctly thickened; t 2 with short ventroapical seta; other tibiae and tarsi simply setulose, only fore and hind basitarsus with 2–3 slightly enlarged setulae ventrobasally; mid basitarsus with 1 somewhat longer and thicker ventrobasal setula. Wing ( Fig. 275 ) elongate and relatively narrow, with pale yellowish ochreous veins and membrane. C with distinct sparse spinulae among hair-like setosity between apices of R 1 and R 2+3. R 2+3 long, bent parallel to C with apex slightly upcurved to it; R 4+5 very slightly bent to straight, parallel with almost straight M. Discal cell (dm) long and narrow; r-m situated very slightly to distinctly in front of the middle of cell dm. Apical portion of CuA 1 slightly to distinctly longer than dm-cu and almost reaching wing margin; A 1 short, ending far from it. Alula distinct but narrow. Wing measurements: length 2.30–3.10 mm, width 0.71–0.95 mm, Cs 3: Cs 4 = 1.07–1.24, rm\dm-cu: dm-cu = 2.24–3.00. Haltere yellowish white, with stem more yellow.

Abdomen dorsally with variable extent of brown colouring on yellow ground, ventrally always pale yellow. Colour pattern of preabdominal terga (T1–T5) ranging from largely yellow to predominantly brown; lightest specimens with only T1–T3 anteromedially brownish (or T1 completely yellow), medium dark specimens with anteromedial brown spots enlarged (often with T2–T3 largely to completely brown) and also present on T4–T5, darkest specimens with T1–T3 completely and T4–T5 largely brown with only sides and posterior margins yellow. T1–T5 sparsely greyish microtomentose and subshining, relatively shortly and sparsely setose. T1 and T2 distinctly separate, only laterally fused. T1–T2 slightly shorter and more transverse than T3–T5, the latter subequal in size, all reaching onto lateroventral sides of abdomen. Preabdominal sterna pale yellow, relatively narrow and becoming wider posteriorly; S1 short and transverse, S2 about as long as wide, S3 slightly, S4 more and S5 (widest sternum) most transverse. S2–S5 densely nnely setose, only S1 bare and with darker posterior marginal stripe. T6 submembranous, short, transverse, bare, almost unpigmented. S6 and S7 yellow, both with brown anterior marginal ledge; S6 with 2–3 (usually 2), S7 with 1–3 (usually 2) setae; S8 longer than epandrium, but paler brown, with only anterolateral marginal area more or less yellow, setose in posterior half.

Genitalia. Epandrium ( Figs 280, 281 ) blackish brown (only in lightest specimens with short posteroventral area light brown to yellow), contrasting with yellow S6–S7 and lighter brown S8, moderately long and relatively broad (slightly wider than in A. dissors ), rather densely setose, with 2 or more pairs of longer and thicker setae dorsolaterally; anal nssure relatively small (slightly shorter than in A. dissors ), rounded triangular. Cercus short and pale, with nne setae, apical longest. Cerci more distant from each other than those of A. dissors . Medandrium ( Fig. 280 ) moderate, slightly narrower than in A. dissors , dorsally somewhat narrowed, with dorsolateral corners simple (not acute), ventrally with shallow broad emargination, bare. Gonostylus ( Figs 280, 281, 286 ) elongate and relatively slender, shorter than epandrial height, slightly bent medially and anteriorly, distally gradually tapering but apically less pointed than in A. dissors and posteroapically provided with 4 small teeth (2 apical and 2 subapical – Fig. 286 ); outer side mostly micropubescent, inner side with longer setae and only slightly concave (both as in A. dissors ). Hypandrium ( Fig. 282 ) closely resembling that of A. dissors , relatively slender, with anterior internal lobes small and low. Transandrium ( Fig. 283 ) straight and relatively thick, without distinct caudal process except for a pair of short medial sclerotizations transilient to spinose parts of basal membrane. Pregonite ( Fig. 282 ) generally formed as in A. pallida group and very similar to that of A. dissors , fused to hypandrium, with slender and acute anterior tooth, posteriorly with nne projection transilient to basal membrane, ventrally with 4–5 (one longer) setae (only 3 setae in A. dissors ). Postgonite ( Fig. 282 ) markedly different from that of A. dissors , shorter and widest proximally (in lateral view) and with broadly rounded apex, pale-pigmented, with 1 setula near middle of anterior margin and several sensillae on outer side. Dorsal internal sclerite (normally situated at base of postgonite) narrow and curved (visible in Fig. 285 attached to aedeagal part of folding apparatus). Basal membrane ( Figs 282, 283 ) with small spinulae covering two posterolateral elongate areas reaching also onto ventral side (as in A. dissors ). Aedeagal part of folding apparatus differing from that of A. dissors in having several dark blunt spines dorsolaterally. Connecting sclerite strong, dark, proximally slender and attached to phallophore, distally widened and provided with a cluster of dark spines ( Fig. 285 ) but, in contrast to A. dissors , with only nne spinulae in membrane posterior to these spines (without additional cluster of dark spines). Phallapodeme as in other members of the A. pallida group, relatively slender, with basal part deeply forked, and fulcrum situated very close to laterally projecting apex ( Fig. 285 ). Aedeagus ( Fig. 285 ) with very short phallophore and large long distiphallus that is bind from near its base. Saccus long as in A. dissors but usually less widened distally, membranous, with elongate slender basal sclerites and armed with fewer (usually 6) robust dark-pigmented spines. Filum also resembling that of A. dissors , long and arched but somewhat wider, with smaller membranous preapical lobes, and apex curved, slender, pointed and with a number of small spines ( Fig. 284 ). Ejacapodeme small, pale, with pointed (not digitiform) terminal end ( Fig. 285 ).

Female. Similar to male unless mentioned otherwise. Total body length 2.65–3.90 mm. Head with ocellar triangle pale brown and/or with brownish colour reduced to small spots at ocelli. Occiput always entirely yellow laterally. Orbit sometimes with 1 additional microsetula in front of short anterior ors setula. Outer side of 1st nagellomere always with large darker orange ochreous to pale brown anterodorsal spot below insertion of arista, also inner side sometimes faintly darkened. Thorax ( Fig. 278 ) usually almost completely yellow, at most with scutum and scutellum darker (orange) than humeral-notopleural area, only postscutellum and/ or postnotum ochreous to brownish darkened (postscutellum sometimes brown). f 3 without row of shortened and thickened posteroventral setae. Discal cell (dm) broader on average. Cross-vein r-m situated often in the middle of cell dm. Wing measurements: length 2.71–3.57 mm, width 0.91–1.19 mm, Cs 3: Cs 4 = 1.03–1.20, rm\dm-cu: dm-cu = 2.07–2.64. Abdomen with T1–T6 ( Fig. 278 ) usually entirely yellow to pale yellow (T5 and particularly T6 often somewhat darker yellow to ochreous), rarely with small medial brownish spots on T1–T6 (or on some of them), exceptionally with more (2–3) small spots on T6. T2–T5 shorter and more transverse than in male, T1–T2 narrower, T3–T5 wider and subequal in size, all wider than T6. Preabdominal sterna pale yellow, slightly narrower than in male, S2 as long as wide, S3 and S4 slightly (both suboblong) and S5 (trapezoidal) distinctly transverse, S5 largest and widest but narrower than postabdominal S6.

Postabdomen ( Figs 287, 289 , 380 ) relatively long, narrow from 7th segment, telescopic, having sclerites yellow to pale yellow with brown markings. T6 simple, large (but narrower than T5), tapered posteriorly and with broadly rounded posterior corners, usually dark yellow, rarely with 1 or 3 brownish spots, with relatively short and dense setae in posterior two-thirds, marginal setae longest. S6 transversely subtrapezoidal with anterior corners rounded or anterior margin almost semicircular, broader than S5, pale yellow and nnely densely setose. 7th segment with separate T7 and S7 ( Figs 289 , 380 ), the former long, subcylindrical, slightly narrower apically, dorsomedially strongly shortened due to deep incision ( Fig. 287 ), ventrally longer, with anterior margin narrowly divided and separated from anteromedial portion of S7, without anteroventral pouch-like lobes ( Fig. 289 ). T7 pale brown to completely dark brown dorsally, and with either one or two dark brown spots ( Fig. 287 ); brown (pale or dark) colour reaching onto sides of T7 and sometimes also anteroventrally where it can be markedly darkened; ventral part of 7th segment with large central area yellow, within which S7 is somewhat darker and still discernible ( Fig. 289 ), nnely setose and surrounded by distinctive micropubescence on membrane, 7th spiracle arising in anterolateral corners of membrane ( Figs 289 , 380 ). Dorsal and lateral parts of T7 with rather short setae mainly at posterior margin ( Figs 287 , 380 ). 8th segment nnely micropubescent laterally. T8 ( Fig. 287 ) about as long as wide, bent to form parallel sides, anteromedially slightly to distinctly emarginate; its colour ranging from pale ochreous yellow to brown, with sparse but long micropubescence and nne exclinate setae laterodorsally; S8 ( Fig. 289 ) about as long as T8, medially divided into 2 convex, nnely hirsute and micropubescent sclerites. Genital chamber (uterus) posteriorly with distinctive dark-pigmented internal sclerotization ( Figs 291, 292 ) formed by 1 compact crooked ventral sclerite (similar to that of A. dissors ), a pair of relatively long, nat and paler dorsal sclerites (these are absent in A. dissors ) and 1 subcircular, curved (in pronle) annular sclerite situated in front of the former. Membranous part of genital chamber large but without additional sclerotizations ( Fig. 290 ). Ventral receptacle ( Fig. 290 ) very slender (also basally), tube-like, hyaline, terminally with multiply twisted vermicular apex. Accessory gland small, vesicular, hyaline, on distally slightly dilated and sparsely ringed duct. Spermathecae (1+1) broadly suboval to ovoid ( Figs 288, 293 ), each with distinct, narrow but deep invagination and a number of short blunt spines in basal fourth surrounding duct insertion; duct very long and ending simply (without distinct cervix) in centre of spermathecal body. T10 small ( Fig. 287 ), suboval to rounded trapezoidal (distinctly wider than in A. dissors ), ochreous to yellow, with 2–3 pairs of setae (1 long) and reduced micropubescence. S10 pale brown to yellow, somewhat larger and wider than T10, suboval in ventral view ( Fig. 289 ), nnely setulose and micropubescent. Cercus relatively short ( Figs 291 , 380 ), not slender, with numerous nne and relatively short setae, apical and dorsopreapical longest.

Discussion. Anthomyza mcalpinei sp. nov. belongs to the A. pallida group, as denned above. Its closest relatives are the Palaearctic A. dissors and the W. Nearctic A. pullinotum sp. nov.; their close relationship can be justined not only from external appearance (thorax often with dorsal darkening, pleura always yellow) but particularly from highly similar structures in the male and female genitalia, including such nne details as: nnely spinulose basal membrane with acute spinulae laterally and nat short ones medially; subterminal membranous lobe-like dilation of the nlum of distiphallus; connecting sclerite distally with a cluster of dark spines; female T7+S7 with original T7 expanded as narrow lobes that almost meet ventromedially (sometimes even meeting in A. dissors ); and female genital chamber with compact unpaired ventral sclerite (interpreted to be a fusion product of a pair of ventral sclerites as known in other relatives of the A. pallida group).At least some of the above characters can be considered synapomorphic and diagnose the A. dissors subgroup. Interestingly, each of the three species belonging to this subgroup has a differently armed aedeagal part of the folding apparatus. Anthomyza pullinotum exhibits the plesiomorphic state of a group of nne dark tubercles, which is also seen in the rest of the A. pallida group outside of this subgroup. Derived states are seen in the other two species of the subgroup, with the tubercles in A. dissors reduced and hyaline, and those of A. mcalpinei reduced to a few enlarged wart-like spines. Anthomyza mcalpinei undoubtedly is the nearest ally of the Palaearctic species A. dissors , demonstrated not only by their external resemblance (including the colour variability of the body, mainly of the mesonotum and abdomen), but chieny by almost identical structures in the male and female genitalia, especially a similarly formed ventral sclerite and a reduced number of dorsal sclerites in the female genital chamber.

Anthomyza mcalpinei differs from both above relatives in having the occiput laterally normally only darker yellow (without brown areas behind eyes); the pleural part of the thorax entirely yellow and without a dark dorsal stripe (but pleura in A. pullinotum also is usually unicolourous yellow); male S6 and S7 yellow, contrasting with blackish brown epandrium; gonostylus apically not pointed but with distinct posteroapical teeth ( Fig. 286 ); aedeagal part of folding apparatus with a few dark blunt spines; saccus with fewer (usually 6) robust dark-pigmented spines; female thorax never brown dorsally; female 7th segment with a distinct S7 that is not fused with T7 ( Fig. 289 ); and female genital chamber with only a single pair of nat sclerites above the single compact and crooked ventral one.

Etymology. This common species is named in honour of J. F. McAlpine, a preeminent Canadian dipterist whose body of work on acalyptrate Diptera systematics serves as an inspiration to the junior author, whose graduate committee he served on.

Biology. There are fewer western records for A. mcalpinei compared to A. pengellyi , and include collections from pure stands of Calamagrostis canadensis (Alberta: w. border of Elk Island N. P.) and Carex utriculata (British Columbia: Fernie). In the east, the primary plants from which this species has been taken are the grasses Phalaris arundinacea and C. canadensis and sedges such as Carex aquatilis , C. utriculata and Scirpus microcarpus J. & C. Presl. There is a single record from “mostly” Carex stricta Lam. (Ontario: Manitoulin Is. – Misery Bay Nature Reserve) and repeated collections from Carex stipata Muhl. ex Willd. var. stipata (Ontario: Sault Ste. Marie – Finn Hill). Beaver dams and meadows are mentioned on labels, with some including additional reference to sedges (Ontario: S of Sioux Narrows, ~ 5 km SE Searchmont, Finland – S of Caliper Lake). A “pasture sweep” yielded A. mcalpinei (Newfoundland: Portugal Cove) while another less specinc collection indicates “ grasses at edge of agricultural neld” (Alberta: Dunvegan), a site shared with A. pengellyi and A. vockerothi . Various other plants such as Equisetum , Typha , Solidago , Rubus and ferns can be included but it is the graminoids in these plant mixtures or growing nearby that are the most likely hosts. A mixed riparian community (Ontario: Greenwater P. Pk., Fig. 279 ) was dominated by Carex spp., C. canadensis , and P. arundinacea (from higher to lower prevalence) while a drier adjacent upstream area, dominated by P. arundinacea , also yielded specimens. There is a singular site in Sault Ste. Marie, Ontario (S of Algoma University) where several hundred specimens were collected over a number of years. This is a small, narrow site that supports a mixture of C. canadensis and C. aquatilis , each transitioning to dominance at either end of a forest edge with a dense patch of P. arundinacea in the middle. This narrow stretch is bordered on the opposite side by a raised municipal snow dump and a wetter area dominated by Typha latifolia L. While most of the collection effort was expended in sampling from the C. aquatilis (primarily for syntopic A. gibbiger and A. orthogibbus ), A. mcalpinei was also very abundant on C. canadensis at this site, but also present at lower numbers on P. arundinacea . More than 150 eggs of A. mcalpinei were obtained from a collection of nine females with three males (Ontario: Sault Ste. Marie – S of Algoma University) and used in a rearing attempt over the period of 8 August to 20 September 2002. The three dominant species of plants from this site ( C. aquatilis , C. canadensis , and P. arundinacea ) were initially offered to the adults and the subsequent larvae, but on 14 September all (~34) surviving larvae were transferred to P. arundinaceae . The single puparium (21 September) which yielded an adult female (4 October) originated from eggs laid on P. arundinacea ; the pupariation period at 20°C was 13 days. The remaining nve larvae were transferred to 10°C on 3 November but were dead by 29 November. Despite the lack of success reported here, one or more of these graminoids are likely to serve as a natural host for A. mcalpinei . Adults of A. mcalpinei have been collected from 3 June (Minnesota: Willow River) to 16 September (Ontario: Sault Ste. Marie).

Distribution. This commonly collected species has a transcontinental distribution in Canada and the northern United States of America that is similar to, but latitudinally narrower than that of A. pengellyi . It has not yet been recorded in New Mexico in the southwest or in the northern jurisdictions of Alaska, Yukon, Northwest Territories and Labrador, although it has been recorded from Newfoundland and Prince Edward Island in the northeast. The list of records includes Canada: Alberta, British Columbia, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Ontario, Prince Edward Island, Quebec, Saskatchewan; United States of America: Colorado, Idaho, Massachusetts, Michigan, Minnesota, Montana, New Hampshire, New York, Oregon, Washington, Wisconsin (see Table 2, Fig. 599 ).

Figures

Figs 2–3. Nearctic species of the genus Anthomyza. 2 – A. shewelli sp. nov., a pair (male on left) on leaf of Carex sp. (Canada:Ontario:Dubreuilville, Magpie River), body length ca.2.4 mm (male) and 3.1 mm (female); 3 –A.mcalpinei sp. nov., living male, laterally (Canada:Ontario:Sault Ste. Marie, Finn Hill), body length ca.2.8 mm. Photo by J. Roháček.

Figs 274–276. Wings of the Nearctic species of the Anthomyza pallida group. 274 – A. pengellyi sp. nov., male, wing length 2.65 mm (Canada: Ontario); 275 – A. mcalpinei sp. nov., male, wing length 2.7 mm (Canada: Ontario); 276 – A. pullinotum sp. nov., male, wing length 2.3 mm (Canada: Yukon Territory). Photo by K. N. Barber.

Figs 277–279.Living Anthomyza mcalpinei sp.nov.and its habitat.277 – A.mcalpinei sp. nov., male,laterodorsally,body length ca. 2.8 mm; 278 – female, laterodorsally, body length ca. 3.5 mm (both from Canada: Ontario: Sault Ste. Marie – Finn Hill); 279 – wetland at a lake innow with Carex, Calamagrostis, Phalaris and other graminoids in Greenwater Prov.Park (Canada:Ontario), habitat of A. mcalpinei. Photo by J. Roháček (Figs 277, 278) and K. N. Barber (Fig.279).

Figs 280–286. Anthomyza mcalpinei sp. nov., paratype male (Canada: Ontario). 280 – external genitalia, caudally; 281 – the same, laterally; 282 – hypandrial complex, laterally; 283 – transandrium, caudally; 284 – apex of nlum, subventrally (widest extension); 285 – aedeagal complex, laterally; 286 – gonostylus, ventrolaterocaudally (widest extension). Scales = 0.05 mm (Fig. 284) and 0.1 mm (others).

Figs 287–293. Anthomyza mcalpinei sp. nov., paratype female (Canada: Ontario). 287 – postabdomen, dorsally; 288 – spermatheca; 289 – postabdomen, ventrally; 290 – distal part of female genital chamber, laterally; 291 – female internal sclerites and apex of postabdomen, laterally; 292 – internal sclerites, ventrally; 293 – spermatheca. Scales = 0.2 mm (Figs 287, 289), 0.05 mm (Figs 288, 293) and 0.1 mm (others).

Figs 294–298. Primary types of the Anthomyza pallida group. 294 – A. pengellyi sp. nov, holotype male, laterally, body length 2.44 mm; 295 – A.mcalpinei sp. nov., holotype male, laterally, body length 2.94 mm; 296 – A. pullinotum sp. nov., holotype male, laterally, body length 2.74 mm; 297 – the same, head and thorax, laterally; 298 – the same, abdomen laterally. Photo by J. Roháček.

Figs 376–380. Female T7+S7 and/or apex of postabdomen of Anthomyza species, laterally. 376 – A. concolor (Thomson, 1869) (USA: California); 377 – A. occidentalis sp. nov., paratype (USA: Oregon); 378 – A. pullinotum sp. nov., paratype (Canada: Alberta); 379 – A. vockerothi sp. nov., paratype (Canada: Ontario). 380 – A. mcalpinei sp. nov., paratype (Canada: Ontario). Scales = 0.2 mm.

Fig. 599. Map of North America showing distributions of two members of Cluster 3 and one member of Cluster 4. Cluster 3: Anthomyza equiseti sp. nov. and A. mcalpinei sp. nov.; Cluster 4: Arganthomyza carbo Roháček & Barber, 2013. The square symbols for A. carbo are enlarged to improve visibility at common localities; there are several records (spots) common to A. equiseti and A. mcalpinei where the latter is not visible, particularly three records running south from James Bay (Ontario: Moosonee). See “Distributional patterns” for explanation of Clusters.