Anthomyza vulgaris, Roháćek & Barber, 2016

Roháćek, Jindřich & Barber, Kevin N., 2016, Nearctic Anthomyzidae: a monograph of Anthomyza and allied genera (Diptera), Acta Entomologica Musei Nationalis Pragae (suppl.) 56, pp. 1-412 : 328-346

Anthomyza vulgaris sp. nov.

( Figs 520, 521 , 525 , 545 , 547, 550–573 )

Type material. HOLOTYPE: ♂, “CAN: ON: ~25kmWNW Ignace, 05.vii.2012, KN Barber , sweeps, fen, mostly Carex utriculata with grasses 49°29.52’N 92°00.83’W ” and “ Holotypus ♂ Anthomyza vulgaris sp. n., J. Roháček & K. N. Barber det. 2014” (red). The specimen is in good condition, with exposed genitalia and highly visible gonostyli (see Fig. 520 ) ( CNCI, intact). PARATYPES: CANADA: ALBERTA: Grand Prairie, ex. Ulmus sp., 21.viii.2008, 1 ♂, J. Donnelly leg. ( CNCI, 1 ♂ genit. prep.). BRITISH COLUMBIA: Mt. Robson Prov. Pk., Hwy #16, small road towards Mt.Robson, 53°03'N 119°15'W, forest noor, swamp, (Universität Bielefeld, Ca1519), 6.viii.2002, 1 ♀, M.v.Tschirnhaus leg. ( ZSMC, in ethanol); Revelstoke, 2.vii.1973, 1 ♀, H. J. Teskey leg.; Robson, 1–20.vii.1949, 1 ♀, 7.vii.1950, 3 ♀♀ (1 ♀ genit. prep.), H. R. Foxlee leg. (all CNCI); Salmon Arm, Shuswap Lk., shore, 13.vii.1949, 2 ♂♂, H. B. Leech leg. ( CASC, 1 ♂ genit. prep.); Sicamous, sweep near rail tracks, 50°50.427'N 118°59.097'W, 26.vi.2011, 1 ♀, J. E. Swann & D. R. Edwards leg. ( BDUC). LABRADOR: 0.25– 1 mi SE, 1 mi up Kenemich R., vegetation on boggy trail, 13–15.viii.1958, 1 ♂ 1 ♀, B[ritish] S[chools] E[xploring] S[ociety] Expedition leg. ( BMNH, both genit. prep.). MANITOBA: Aweme, 19.vii.1917, 1 ♀, 10.viii.1917, 1 ♀, 28.viii.1917, 1 ♀, N. Criddle leg. ( CNCI); Aweme, Criddle farm, 49°42.5'N 99°36.1'W, sweep grasses near house, 15.viii.1999, 1 ♂, T.A. Wheeler leg. ( LEMQ 0039427), 8.vii.2000, 11 ♂♂ 6 ♀♀, V. Crecco leg. ( LEMQ 0039411–422, -424, -484, -485, -496, -512), sweep prairie near house, 13.vi.1999, 1 ♀ ( LEMQ 0039515), 8.vii.2000, 3 ♂♂ ( LEMQ 0039189, -96, -97), 19.vi.1999, 4 ♂♂ 2 ♀♀ ( LEMQ 0039342–45, -351, -475), T. A. Wheeler leg.; Bald Head Hills, 13 mi N Glenboro, 12.vii.1958, 1 ♀; 9 mi N Forrest, 29.vi.1958, 1 ♂, both R. L. Hurley leg. (both CNCI); 5 km N Gardenton, Tallgrass Prairie Reserve, 49°10.71'N 96°40.76'W, sweep roadside, 13.vii.2000, 3 ♂♂ 3 ♀♀, Crecco & Wheeler leg. ( LEMQ 0039163–65, -69–71), sweep prairie at roadside, 6.vii.2000, 5 ♂♂ 2 ♀♀, T.A. Wheeler leg. ( LEMQ 0039198–203, -208), sweep in tallgrass prairie, 12.vi.1999, 1 ♀ ( LEMQ 0039530), 17.vi.1999, 2 ♀♀ ( LEMQ 0039434, -543), 20.vi.1999, 1 ♂ 1 ♀, J. Perusse leg. ( LEMQ 0039438, -517), 6.vii.2000, 1 ♂ 1 ♀, T.A.Wheeler leg. ( LEMQ 0039204, -513); Ninette, 15.vii.1958, 1 ♀, N. B. Chillcott leg., ex. Betula glandulosa , 15.vii.1958, 1 ♂ 3 ♀♀ (1 ♂ genit. prep.), J. G. Chillcott leg., 1 ♀, R. L. Hurley leg.; 3 mi S Shilo, ex. leafy spurge, 30.vi.1958, 2 ♂♂ 5 ♀♀, R. L. Hurley leg.; 5 mi SW Shilo, 22.vii.1958, 2 ♂♂ 4 ♀♀, C. D. F. Miller leg., 13.viii.1958, 1 ♀, swept from open prairie, 11.vii.1958, 7 ♂♂ 6 ♀♀ (1 ♀ genit. prep.), Andropogon - Stipa prairie community, 22.vii.1958, 4 ♂♂ 4 ♀♀, at margin of grain neld, 22.vii.1958, 3 ♂♂ 2 ♀♀, tamarack, 2.viii.1958, 1 ♀, J. G. Chillcott leg., noodplain community nr. tamarack bog, 11.vii.1958, 1 ♀, J. G. Chillcott leg., 22.vii.1958, 1 ♂, R. L. Hurley leg. (all CNCI); 1 km E Stuartburn, 49°08'N 96°44.7'W, sweep vegetation along roadside, 12.vi.1999, 1 ♀, V. Crecco leg. ( LEMQ 0039516); Treesbank, 20. vii.1915, 1 ♂ 1 ♀, 27.viii.1915, 1 ♀ ( USNM), 6.viii.1915, 1 ♂ 4 ♀♀ ( CNCI, 1 ♂ 1 ♀ with Aldrich det. as Anthomyza gracilis ), N. Criddle leg.; Whitemouth, 10.vii.1958, 1 ♂, J. G. Chillcott leg. ( CNCI); Winnipeg, St. Charles Rine Range, 49°54.2'N 97°20.3'W, sweep in tallgrass prairie, 4.vii.1997, 1 ♀, D. Pollock leg. ( LEMQ 0039532), 24. vi.1998, 2 ♂♂ 4 ♀♀ ( LEMQ 0039433, -439, -442, -444, -452, -549), 1.vii.1998, 4 ♂♂ 4 ♀♀ ( LEMQ 0039537–40, -45–48), 3.vii.1998, 1 ♀ ( LEMQ 0039533), 8.vii.1998, 8 ♂♂ 9 ♀♀ ( LEMQ 0039425, -426, -428, -432, -436, -461, -466, -467, -527, -528, -531, -550, -554–558), 10.vii.1998, 1 ♂ 4 ♀♀ ( LEMQ 0039446, -463, -468, -469, -551), 15.vii.1998, 1 ♂ 2 ♀♀ ( LEMQ 0039541, -52, -53), 17.vii.1998, 1 ♂ 2 ♀♀ ( LEMQ 0039529, -35, -44), 22.vii.1998, 4 ♂♂ 7 ♀♀ ( LEMQ 0039462, -453, -456, -457, -464, -465, -534, -524–526, -536), P. Bouchard leg.; Winnipeg, St. Charles Rine Range, Go Back Prairie, 49°54.6'N 97°20.3'W, sweep in tallgrass prairie, 18.vi.1999, 1 ♀ ( LEMQ 0039431), 13.viii.1999, 1 ♂ ( LEMQ 0039450), V. Crecco leg.; Winnipeg, St.Charles Rine Range, Arrowhead block, 49°54.6'N 97°20.5'W, sweep in tallgrass prairie, 30.vi.1999, 1 ♀ ( LEMQ 0039499), 7.vii.1999, 4 ♂♂ 3 ♀♀ ( LEMQ 0039440, -443, -445, -447–449, -454), 14.vii.1999, 4 ♂♂ 1 ♀ ( LEMQ 0039451, -55, -58–60), S. McMillan leg., 13.viii.1999, 3 ♀♀ ( LEMQ 0039435, -41, -42), 18.viii.1999, 1 ♂ ( LEMQ 0039437), T.A. Wheeler leg., 11.viii.1999, 3 ♀♀ ( LEMQ 0039423, -429, -430), V. Crecco leg., 14.vii.2000, 7 ♂♂ 15 ♀♀ ( LEMQ 0039494, -495, -497, -498, -500–502, -504–511, -514, -518–523), Crecco & Wheeler leg. NEW BRUNSWICK: Birch Cove, nr. Chamcook, 14. viii.1957, 1 ♀, G. E. Shewell leg. ( CNCI); Cape Pelée, 46°13.79'N 64°14.5'W, sweep vegetation at chalet, 31. vii.2004, 1 ♂ 1 ♀, S. Boucher leg. ( LEMQ 0040524, -26); Chamcook, 9.viii.1957, 2 ♀♀; Chamcook, Glebe Road, 14.vii.1965, 1 ♀, all G. E. Shewell leg. (all CNCI); Grand Falls, 8.viii.1956, 1 ♂ 1 ♀, A. H. Sturtevant leg. ( USNM); Ingonish, North Bay, 7–8.vii.1984, 1 ♂ 1 ♀, H. J. Teskey leg.; Kouchibouguac N. Pk., Code 6030X, 7.vii.1977, 8 ♂♂ 6 ♀♀ (1 ♀ genit. prep.), J. F. McAlpine leg.; Halifax Co., Lawrencetown, 19–20.vii.1967, 2 ♀♀, D. M. Wood leg.; Maces Bay, Hwy 790, 9.vii.1971, 5 ♂♂ 8 ♀♀ (1 ♂ genit. prep.), B. V. Peterson leg. (all CNCI); near McGraw Brook, Road 108, 46.8°N 66.2°W, sweeping, 30.vii.2001, 1 ♂ 1 ♀, M. Giroux leg. ( LEMQ 0039188, -95); Middle Sackville, 45°55.4'N 64°21.4'W, sweep vegetation along old rail line, 19.vii.2002, 6 ♂♂ 3 ♀♀, J. Forrest & T. Wheeler leg. ( LEMQ 0039173, -39186, -39187, -39192, -39194, -40343, -40346–40348); Pt.Lepreau, 10.viii.1956, 1 ♂, A. H. Sturtevant leg. ( USNM); Sackville, near Mt. Allison Univ., 45°53.9'N 64°22.5'W, sweep old garden in vacant lot, 19.vii.2002 3 ♂♂ 5 ♀♀, J. Forrest & T. Wheeler leg. ( LEMQ 0040330–32, -34, -35, -37, -38, -40); Sackville, Tantramar Marsh, 14.vii.1983, 1 ♂ 6 ♀♀, J. R. Vockeroth leg. ( CNCI). NOVA SCOTIA: Crescent Beach, 44°13.8'N 64°23.2'W, sweep dune vegetation, 24.vii.2002, 2 ♀♀, J. Forrest & T. Wheeler leg. ( LEMQ 0039166, -67); Hampton, 44°54.7'N 65°20.5'W, sweep dense vegetation along path above ocean beach, 26.vii.2002, 6 ♂♂ 4 ♀♀, J. Forrest & T. Wheeler leg. ( LEMQ 0040350–57, -59, -64); Lockeport, 18.vii.1958, 4 ♂♂ 7 ♀♀, 24.vii.1958, 1 ♀, 25.vii.1958, 2 ♂♂ 2 ♀♀, J. R. Vockeroth leg. ( CNCI); Moser River, sweep vegetation along salt shore, 18.viii.1996, 4 ♂♂ 3 ♀♀, T. A. Wheeler leg. ( LEMQ 0039352–55, -58–60); Paquette Lk., 5 km S South Harbour, 5.vii.1983, 1 ♂ 1 ♀, A. Borkent leg. ( CNCI); Peggy’s Cove, sweep grasses near shore W of lighthouse, 19.viii.1996, 1 ♀, T. A. Wheeler leg. ( LEMQ 0039219); S. Harbour, PG961944, 11.vii.1984, 1 ♀, H. J. Teskey leg., PG929935, marshy lake shore, brackish water, Carex , Juncus , Iris , 27.vi.1983, 1 ♀, J. R. Vockeroth leg.; S. Harbour Beach, PG949958, sand beach, Ammophila & Lathyrus , 3.vii.1983, 2 ♂♂ 1 ♀, J. R.Vockeroth leg.; Springneld, 6.viii.1958, 1 ♀, J. R. Vockeroth leg. (all CNCI). ONTARIO: Algoma Mills, Lauzon Village Rd., 0.5 km N Hwy.17, 46°11.71'N 82°48.37'W, sweeps, Carex , Calamagrostis , edge of fen, 21.vii.2007, 5 ♂♂ 5 ♀♀, K. N. Barber leg. ( CNCI);Algonquin Pk., 23.vi.1962, 1 ♀, R. J. Pilfrey leg., 27.vii.1981, 1 ♀, D. J. Mountjoy leg. ( DEBU); Algonquin Park, 26–30. vii.1955, 2 ♂♂ 9 ♀♀, Sabrosky leg. ( USNM); Algonquin, mixed wood, 1.vi.1991, 1 ♀, M. Barták leg. ( MBPC, genit. prep.); Algonquin P. Pk., nr. Brent, Cedar Lake, 46°01'39"N 78°29'58"W, 29.vi.2008, 2 ♂♂ 2 ♀♀; Nipissing Co., Algonquin P. Pk., Tea Lk. Dam, Oxtongue River, 19.vii.1991, 1 ♀, all B. J. Sinclair leg. (all CNCI); 4 mi W Arthur, weedy ditch, 25.vii.1985, 14 ♂♂ 25 ♀♀, K. N. Barber leg. ( DEBU); Atikokan, 14 mi E on Hwy. 11, 3–4. vii. 1978, 1 ♀, H. J. Teskey leg. ( CNCI); 10 km E Atikokan, on Hwy.11, grass/ sedge / Typha along roadside, 11. vii.1992, 1 ♀, T.A.Wheeler leg. ( LEMQ 0039334);Atwood Twp., clearing, raspberry, 24.vii.1992, 1 ♂, S. Hughson leg. ( DEBU); Baptiste Lake, 45°10'N 78°00'W, sweep near lake shore, 25.vii.2000, 1 ♂ ( LEMQ 0039364), sweep at edge of forest, 23.vii.2000, 3 ♀♀ ( LEMQ 0039216, -217, -363), sweep old railway line, clearings in forest, 15. vii.2001, 1 ♀ ( LEMQ 0039370), J. Forrest leg.; Batchawana, 10.vii.1955, 1 ♀, J. Gustafson leg. ( LEMQ 0039214); Belwood, 8.vii.1972, 1 ♀, 16.vii.1972, 1 ♂ 2 ♀♀, D. H. Pengelly leg. ( DEBU); Bergland, 4.viii.1960, 1 ♀, S. M. Clark leg. ( CNCI); Bissett, [-]. vi.1954, 1 ♀, H. D. Stalker leg. ( AMNH); Blackburn, 12.vii.1954, 4 ♂♂ 8 ♀♀; Blackburn, Mer Bleue, 29.vi.1954, 1 ♀, 27.vi.1955, 1 ♂ 4 ♀♀, all W. R. M. Mason leg.; 1.5 km E Bruce Mines, 46°18.1'N 83°46.4'W, sweeps, short roadside grasses, 1.viii.1997, 1 ♀, K. N. Barber leg. (all CNCI); Bruce Co., Dorcas Bay, 45°11'N 81°35'W, 22.vi.2008, 1 ♀, S. A. Marshall leg. ( DEBU 00299114); Bruce Co., Bruce Pen. N. P., George Lake, 15.viii.2007, 1 ♀, S. M. Paiero leg. ( DEBU 00286986); Bruce Co., Dunks Bay, 45°14.8'N 81°38.5'W, 30.vii.1997, sweeps, roadside veg., 1 ♂ 1 ♀; Bruce Co., Hwy#6 @ Willow River, 45°10.43'N 81°31.21'W, sweeps, mostly Carex aquatilis , 21.vi.2008, 1 ♀; Bruce Peninsula N. P., Bartley Lake, 45°12.9'N 81°29.3'W, sweeps, shoreline graminoids, 2.vii.1999, 1 ♀; Bruce Peninsula N. P., Cameron Lk. Rd., 45°12.9'N 81°33.0'W, sweeps, roadside grasses, 3.vii.1997, 1 ♀; Bruce Peninsula N. P., Crane River, 45°07.0'N 81°32.1'W, sweeps, grass/ sedge fen meadow, 3.vii.1999, 2 ♀♀; Bruce Peninsula N. P., Crane River below Lake Scugog, 45°07.0'N 81°32.1'W, sweeps, riverside vegetation, 3.vii.1999, 3 ♀♀, all K. N. Barber leg.; Bruce [Pen.] N. P., Cyprus Lake campground, trail nr. Horse Lake, 24–25.vi.1996, 1 ♀, S. A. Marshall leg.; Bruce Peninsula N. P., Dorcas Bay Rd. at Willow Creek, 45°09.4'N 81°34.4'W, sweeps, mostly creekside graminoids, 3.vii.1999, 2 ♂♂ 1 ♀, K. N. Barber leg. (all DEBU); Bruce Co., Bruce Pen. N. P., Lower Andrew Lk., portage trail, sweep, 8.vi.2000, 1 ♂ 1 ♀, C. S. Onodera leg. ( DEBU 00080038, -56); same locality but 45°13'N 81°27'W, 24.viii.2003, 2 ♂♂, S. A. Marshall leg. ( DEBU 00298601, -27); Bruce Peninsula N. P., Singing Sands, 45°11.6'N 81°34.7'W, sweeps, Agropyron sp., 29.vii.1997, 1 ♂ 11 ♀♀, sweeps, Calamagrostis canadensis , 29.vii.1997, 2 ♂♂ 5 ♀♀, sweeps, Agropyron / Calamagrostis , 30.vii.1997, 4 ♂♂ 13 ♀♀, 5.vii.1998, 10 ♂♂ 39 ♀♀; same locality but 45°11.51'N 81°34.66'W, sweeps, grasses on sand, 22.vi.2008, 3 ♂♂ 6 ♀♀, all K. N. Barber leg. (all DEBU); Bruce Peninsula N. P., Dorcas Bay, Singing Sands, 45°11'N 81°35'W, sweep shore, 2.vii.2001, 1 ♂, M. Pollet leg. ( LEMQ 0039185); 1 km S Chalk River, St.Andrew’s UC, Forest View Cemetery, 46°00.4'N 77°25.9'W, sweeps, mostly grasses, 6.vii.2001, 2 ♂♂ 11 ♀♀; ~ 13.9 km W Chapleau, 47°49.20'N 83°35.42'W, hydro right-of-way, sweeps, mostly Carex utriculata , grasses, 23.vi.2013, 1 ♂, 13.vii.2013, 6 ♂♂ 1 ♀, 7.vii.2014, 5 ♂♂ 1 ♀ (all CNCI); ~ 5 km SE Cochrane, 49°01.16'N 80°57.93'W, sweeps, railside Equisetum spp., graminoids, herbs, 18.vii.2009, 27 ♂♂ 48 ♀♀ ( AMNH, CASC, LACM, USNM 5 ♂♂ 7 ♀♀ each; 7 ♂♂ 20 ♀♀ CNCI), all K. N. Barber leg.; Deux Riv[ieres], 8.vii.1954, 1 ♀, A.H. Sturtevant leg.( USNM); ~ 32 km NW Dowling, Hwy #144 @ Moncrieff Ck., 46°46.53'N 81°36.50'W, sweeps, graminoids on muddy noodplain, 29.vii.2007, 1 ♀; Dryden, 49°47.1'N 92°48.4'W, sweeps, grasses in low area, 10.vii.1999, 3 ♂♂ 9 ♀♀ (all CNCI, 1 ♂ genit. prep.); Dryden, 49°47.27'N 92°48.62'W, sweeps, mixed graminoids/herbs, 17.vii.2008, 11 ♂♂ 14 ♀♀ ( BDUC, BYUC 4 ♂♂ 6 ♀♀ each; CNCI 3 ♂♂ 2 ♀♀); ~ 35 km WSW Dubreuilville, 2 km SE Jct. Hwys.#17 & #519, 48°17.16'N 84°53.34'W, sweeps, Calamagrostis / Scirpus , 23.vii.2003, 4 ♀♀, sweeps, roadside vegetation incl. wet ditch, 31. vii.2008, 2 ♂♂ 14 ♀♀, sweeps, Equisetum sp., 31.vii.2008, 2 ♀♀ ( CNCI), all K. N. Barber leg.; Dundas, 25.vi.1980, 4 ♂♂ 2 ♀♀, S. Beierl leg. ( DEBU); 5 km W Dyer Bay, Crane Lake Rd., 45°10.9'N 81°23.8'W, sweep alvar savanna, 18.vii.2000, 4 ♂♂ 16 ♀♀, V. Crecco & T. Wheeler leg. ( LEMQ 0039470–74, -76–83, -86–92); Elliot Lake, 46°22.23'N 82°36.49'W, sweeps, mixed graminoids incl. Carex utriculata , 29.vi.2013, 7 ♂♂ 3 ♀♀, K. N. Barber leg. ( CNCI); ~ 55 km NNW Elliot Lake, S of Rocky Island Lake, 46°49.32'N 82°59.54'W, 455 m, sweeping vegetation with predominant Scirpus sp., 3.vii.2010, 1 ♀, J. Roháček leg. ( SMOC, genit. prep.); ~ 66 km NNW Elliot Lake, Rocky Island Lake, 46°50.82'N 83°08.76'W, sweeps, Scirpus [ microcarpus ] on dried shoreline, 4.vii.2010, 1 ♂, K. N. Barber leg. ( CNCI), 405 m, sweeping Scirpus sp. on dried shoreline, 4.vii.2010, 5 ♂♂ 8 ♀♀, J. Roháček leg. ( SMOC, 1 ♂ 1 ♀ genit. prep.); Elora, 27.vi.1977, 1 ♀, K. N. Barber leg., 1 ♀, E.A. Innes leg. ( DEBU); ~ 11 km ESE English River, Hwy#17, 49°09.42'N 90°50.11'W, sweeps, Equisetum ssuviatile , 17.vii.2008, 1 ♀, K. N. Barber leg. ( CNCI); Erin, 25.vi.1979, 2 ♂♂, 19.vii.1979, 1 ♂ 1 ♀, J. Ernst leg. ( DEBU); 8 km NE Espanola, Hwy 17 at Spanish R., 46.3°N 81.67°W, sweep veg. at rest area, 28.vi.2007, 1 ♀, J. Mlynarek leg. ( LEMQ 0040529), 2 ♀♀, T. A. Wheeler leg. ( LEMQ 0040284, -533); Finland, 28.vi.1960, 1 ♂ 5 ♀♀, Kelton & Whitney leg., 28.vi.1960, 1 ♀, 17.vii.1960, 2 ♀♀, S. M.Clark leg.(all CNCI); Finland, S of Caliper Lake, sweep sedge at beaver dam, 10.vii.1992, 9 ♂♂ 10 ♀♀, T. A. Wheeler leg. ( LEMQ 0039289, -312–314, -316–326, -328–331); Fitzroy Harbour, 11.vii.1938, 2 ♂♂, O. Peck leg. ( CNCI, 1 ♂ genit. prep.); Flinton, St. John the Evangelist RC Church, 44°41.3'N 77°12.8'W, sweeps, mowed grasses on stable sand, 9.vii.2001, 1 ♀; ~7.0 km E Foleyet, 48°14.34'N 82°20.75'W, hydro right-ofway, sweeps, mostly Carex utriculata , 13.vii.2013, 1 ♂; Fraserdale, 49°50.71'N 81°36.99'W, sweeps, graminoids, herbs in disturbed area, 19.vii.2009, 1 ♂ 9 ♀♀, all K. N. Barber leg. (all CNCI); Goderich, 22.vi.1977, 1 ♂ ( DEBU); Greenwater P. Pk., 49°11.05'N 81°16.04'W, sweeps, mostly emergent Equisetum ssuviatile , 18.vii.2009, 1 ♀ ( DEBU 01501988); Greenwater P. Pk., 49°11.34'N 81°17.04'W, sweeps, grasses /herbs in hydro cut, 21.vii.2009, 8 ♂♂ 23 ♀♀ ( DEBU 01502254–84), all K. N. Barber leg.; Goulais River, Pine Shores Rd, G. R. Cemetery, 46°41.91'N 84°25.71'W, sweeps, mowed grasses on stable sand, 30.vi.2007, 1 ♂ 3 ♀♀; Goulais River, end of Island Rd., 46°43.57'N 84°24.45'W, sweeps, mud nats, Equisetum ssuviatile , Dulichium arundinaceum , 9.vii.2007, 1 ♂, all K. N. Barber leg. (all CNCI); Goulais River, Sand Bay, 46°44.81'N 84°32.68'W, sweeping Juncus and Carex at margin of fen pools, 10.vii.2010, 1 ♂, J. Roháček leg. ( SMOC, genit. prep.); Wellington Co., Guelph, UG campus, sweep, 17.viii.1992, 1 ♀, H. Nadel leg. ( RBCM); Wellington Co., Guelph, U of G Arboretum (north), neld sweep, 12. vii.1993, 2 ♂♂, (south), neld sweep, 17.vii.1993, 2 ♂♂ 2 ♀♀, D. C. Caloren leg.; U of Guelph Arboretum, sweep, young goldenrod, 18.vii.1995, 1 ♂, J. M. Dow leg. (all DEBU); Guelph, University Arboretum, sweeping over boggy meadow, 19.viii.1994, 1 ♀, J. Roháček leg. ( SMOC, genit. prep.); Guelph, 23.vi.1976, 1 ♀, J. M. Cumming leg., 18.viii.1976, 1 ♀, D. Levin leg., 21.vi.1976, 1 ♀, 3.vii.1980, 1 ♀, S. A. Marshall leg., 20.vi.1977, 1 ♀, 21. vi.1977, 1 ♂, 5.vii.1977, 1 ♂ 1 ♀, W. A. Atwater leg., 23.vi.1977, 1 ♀, 5.vii.1977, 1 ♂ 2 ♀♀, 16.vi.1978, 1 ♀, 22.vi.1978, 1 ♀, 28.vi.1978, 1 ♂ 3 ♀♀, 11.vii.1978, 1 ♂, 1.viii.1980, 1 ♀, 2.viii.1980, 9 ♂♂ 11 ♀♀, K. N. Barber leg., 12.vii.1978, 1 ♀, N. Pierce leg., 8.vi.1978, 1 ♂, 13.vii.1978, 1 ♂ 2 ♀♀, B. Warner leg., 6.vii.1979, 1 ♀, K. L. Bailey leg., 26.vii.1979, 2 ♀♀, 27.vii.1979, 1 ♀, D. Lewis leg., 11.vii.1979, 1 ♀, 30.vii.1980, 1 ♀, S. Beierl leg., 18.vii.1980, 3 ♀♀, K. H. Harvey leg., 17.vii.1980, 1 ♂ 1 ♀, L. L. Krailo leg., 13.vii.1983, 2 ♂♂ 5 ♀♀, R. Damaio Smith leg., 31.vii.1986, 1 ♀, Z. Y. Huang leg., pan traps, 18–26.vii.1980, 1 ♀, 24.vi.–14.vii.1981, 1 ♂, Malaise trap, 1–7.vii.1985, 1 ♀, K. N. Barber leg.; Havlock, 4.vii.1977, 1 ♀, J. W. McCreadie leg. (all DEBU); Hawk Jct., 48°05.39'N 84°33.62'W, 17.vi.2004, sweeps, mowed grasses on stable sand, Community Centre, 2 ♂♂ 7 ♀♀, Pine Grove Cemetery, 1 ♀, K. N. Barber leg. ( CNCI); Hepworth, 4.vii.1979, 1 ♀, D. L. Krailo leg. ( DEBU), 26.vii.1997, 1 ♀, S. A. Marshall leg. ( DEBU 00075280); ~ 1.9 km E MB border, Hwy#17, 49°44.42'N 95°07.61'W, sweeps, mixed graminoids, 30.vii.2011, 4 ♂♂ 3 ♀♀; Jct.Hwys 17 & 108, 46°12.8'N 82°34.3'W, sweeps, roadside graminoids near wet area, 1.viii.1997, 1 ♂ 3 ♀♀, all K. N. Barber leg. (all CNCI); Hwy 17N & Trout Lake Road, 46°37.563'N 84°17.019'W, sweep, roadside, 23.vii.2011, 2 ♀♀, J. E. Swann & D. R. Edwards leg. ( BDUC); Hills, 11.vii.1978, 1 ♀, M. Lichtenberg leg.; Hilton Township, Malaise at edge of hardwood forest & neld, 4.ix.1992, 1 ♂ (genit. prep.), J. E. Swann leg. (both DEBU); Hilton Twp., 46°14.874'N 83°53.739'W, abandoned neld, Malaise, 21–26.vii.2011, 1 ♂, J. E. Swann & D. R. Edwards ( BDUC); Icewater Creek WS [Watershed], 13.5 km NNE Searchmont, mi.11.5 Whitman Dam Rd., sandy access road, 17.vi.1986, 1 ♂, 19.vi.1986, 1 ♀, [D. J.] M. Harvey leg., 21.viii.1985, 1 ♀, 20.vi.1986, 4 ♀♀, 1.vii.1986, 1 ♂ 3 ♀♀; ~ 25 km WNW Ignace, 49°29.49'N 92°00.71'W, sweeps, wet area, mostly Calamagrostis canadensis , 30.vii.2008, 7 ♂♂ 17 ♀♀, sweeps, fen, mostly Carex utriculata with grasses, 30.vii.2011, 1 ♂ 1 ♀; same locality but 49°29.48'N 92°00.75'W, sweeps, fen, mostly Carex utriculata with grasses, 30.vii.2011, 1 ♂ 2 ♀♀; same locality but 49°29.52'N 92°00.83'W, sweeps, fen, mostly Carex utriculata with grasses, 4.vii.2012, 9 ♂♂ 14 ♀♀, 5.vii.2012, 14 ♂♂ 7 ♀♀, 6.vii.2012, 2 ♂♂ (1 ♂ genit. prep.), all K. N. Barber leg. (all CNCI); Innisnl, 44°19.346'N 79°35.009'W, sweep around pond embankment, 28.vii.2011, 1 ♂, J. E. Swann & D. R. Edwards leg. ( BDUC); Bruce Co., Inverhuron P. Pk., sweep rocky stream bank and vegetation, 5.vii.2001, 1 ♀, M. Pollet leg. ( LEMQ 0040467); Iroquois Falls, overgrown wet shrubby Sphagnum bog, 18.vi.1987, 2 ♂♂, J. R. Vockeroth leg. ( CNCI); Kirkland Lk., 25.vi.1980, 1 ♂, J. Cashaback leg. ( DEBU); Lake Duncan, 1.5 mi N, 30.vi.1991, 1 ♀, Pont leg. ( BMNH); Muskoka Distr., Lake Kahshe, 5 km S Gravenhurst, 20–26.vi.2003, 2 ♂♂ 4 ♀♀, O. Lonsdale leg. ( DEBU 00226770, -773, -778, -808, -822, -830); Kenora Distr., Lake of the Woods, 6 km NNW Sioux Narrows, N side Long Bay, Bur Oak prairie remnant, sweep, 22.vi.1998, 1 ♂, Oldham & Bakowsky leg. ( DEBU 00154994); Killarney P. Pk., near Hwy 637, 46°02.66'N 81°19.83'W, sweep net in sedge meadow with grasses & Rubus , 11.vii.2004, 1 ♂ 2 ♀♀, J. Forrest leg. ( LEMQ 0040317–19); Lake Superior P. Pk., Agawa Bay Campgd., 47°20.3'N 84°37.2'W, sweeps, grasses on sand along trail, 4.vii.2000, 1 ♀ ( DEBU); Lake Superior P. Pk., Hwy 17 near jct. Agawa Rock, 47°22.31'N 84°41.23'W, sweeps, mostly Carex utriculata , 12.vii.2014, 1 ♀ ( DEBU 01503947), both K. N. Barber leg.; Algoma District, Lake Superior Prov. Pk., Rabbit Blanket Lake, 43°41'24"N 80°23'12"W [coordinates do not match locality], 400 m, 20.vi.2001, 1 ♂, S. A. Marshall leg. ( DEBU 00170278); 6 km S Langton Concession Rd. 8, S Hwy 45, 42°42.01'N 80°31.93'W, sweep clearing near Carolinean forest, 30.vi.2002, 1 ♀, S. Boucher leg. ( LEMQ 0040476); Lefroy, 16.vi.1982, 2 ♂♂ 2 ♀♀, D. Morris leg.; Luther Marsh Bog, sweeps, 27. vi.1985, 1 ♀, K. N. Barber leg. (all DEBU); ~ 6.3 km E Macleod, 49°41.37'N 86°51.41'W, sweeps, graminoids incl. Carex utriculata & C. aquatilis substricta , 6.vii.2012, 1 ♂; Manitoulin Is., Carter Bay, 45°36.3'N 82°08.5'W, sweeps, roadside vegetation, 30.vi.1999, 1 ♂; Manitoulin Is., ~2.2 km N Cold Springs, Perch Ck. at Hwy 540, 45°53.2'N 82°06.3'W, sweeps/pooter, Calamagrostis canadensis , 1.viii.1997, 2 ♀♀, sweeps, various grasses / sedges in noodplain, 3.vii.1998, 4 ♂♂ 37 ♀♀, 5.vii.1998, 3 ♂♂ 47 ♀♀ (1 ♂ genit. prep.); same locality but 45°53.1'N 82°06.2'W, sweeps, various grasses / sedges in noodplain, 4.vii.1999, 5 ♂♂ 28 ♀♀ (1 ♂ 1 ♀ genit. prep.), all K. N. Barber leg. (all CNCI); Manitoulin Is., 10 km SW Gore Bay, 45°51.6'N 82°31.5'W, sweep dry meadow at roadside, 17.vii.2000, 1 ♀, V. Crecco & T. Wheeler leg. ( LEMQ 0039191); Manitoulin I., ditch at Honord [sic Honora], 3. vii.1998, 1 ♂, S. A. Marshall leg. ( DEBU 00075281); Manitoulin I., Kip Fleming Tract, ~ 8 km SW Gore Bay, 45°52'13"N 82°32'31"W, oak savannah/alvar, 2–3.vii.2010, 7 ♂♂ 12 ♀♀, S. M. Paiero leg.( DEBU 00329614, -622, -623, -629, -652, -674, -675, -682, -684, -685, -719, -726, -733, -734, -741, -744, -746, -749, -765); Manitoulin I., Michael’s Bay, 8.viii.2003, 1 ♀ ( DEBU 01131678), nr. boat launch, 9.viii.2003, 1 ♂ 8 ♀♀ ( DEBU 00253642–649, -653), S. A. Marshall leg.; Manitoulin Is., Michael’s Bay Pk., 45°36.0'N 82°06.2'W, sweeps, grasses near shore, 28.vii.1997, 1 ♂ 3 ♀♀; same locality but 45°36.0'N 82°06.3'W, sweeps, lakeside grasses, 4.vii.1999, 2 ♂♂ 10 ♀♀; Manitoulin Is., 0.7 km N Michael’s Bay Pk., 45°36.5'N 82°06.2'W, sweeps, Carex / Calamagrostis in fen nat, 5. vii.1998, 1 ♀, all K. N. Barber leg. (all CNCI); Manitoulin Is., Misery Bay [Prov.] Nature Reserve, 45°47.42'N 82°44.07'W, sweeps, trailside graminoids, 27.vii.2004, 4 ♂♂ 3 ♀♀, K. N. Barber leg. ( DEBU 01501485–91); Manitoulin I., Misery Bay Prov. Nat. Res., 45°47'28"N 82°44'58"W, alvar, 4–5.vii.2010, 3 ♂♂ 3 ♀♀, S. M. Paiero leg.( DEBU 00329436, -449, -451, -458, -466, -467); Manitoulin Is., Misery Bay [Prov.] Nature Reserve, 45°47.64'N 82°44.93'W, sweeps, mostly Carex stricta from wetlands boardwalk, 2.vii.2010, 2 ♀♀, K. N. Barber leg. ( DEBU 01502491, -92), sweeping, mostly Carex [ stricta ] from wetland boardwalk, 2.vii.2010, 2 ♂♂ 5 ♀♀, J. Roháček leg. ( SMOC, 1 ♂ 1 ♀ genit. prep.); Manitoulin Is., nr. Misery Bay [Prov.] Nature Reserve, Little Lake Huron Road, 45°47.93'N 82°45.52'W, sweeps, mostly Carex , Juncus on wet alvar, 2.vii.2010, 1 ♂ 1 ♀, K. N. Barber leg. ( CNCI), sweeping, mostly Carex , Juncus on wet alvar, 2.vii.2010, 3 ♂♂ 3 ♀♀, J. Roháček leg. ( SMOC, 1 ♂ genit. prep.); Manitoulin Is., Providence Bay at Mindemoya R., 26.vi.1992, 1 ♂, T.A. Wheeler leg. ( LEMQ 0039307); Manitoulin I., Sand Bay, 45°48'06"N 82°47'36"W, dunes, sweep grassy area by stream, 25.vi.2003, 1 ♂, M.Buck leg.( DEBU 01131826); Hwy#17, ~8.5 km NW Marathon, 48°47.69'N 86°26.07'W, sweeps, roadside graminoids, 31.vii.2008, 3 ♂♂ 9 ♀♀, K. N. Barber leg.; Marmora, 26.vi.1952, 2 ♀♀, J. R. Vockeroth leg., 16.vi.1952, 1 ♂, J. R. McGillis leg., 27.vi.1952, 1 ♀, 4.vii.1952, 1 ♂, C. Boyle leg. (all CNCI); Marten River, 8.vii.1954, 1 ♂ 1 ♀, A. H. Sturtevant leg. ( USNM); Mattawa, 16.vi.1987, wet alder thicket, rich undergrowth, 1 ♀, J. R. Vockeroth leg.; Maynooth, 2.viii.1952, 1 ♀, J. F. McAlpine leg.; Metcalf, 15.vii.1982, 1 ♀, 12.vii.1983, 1 ♀, 11.vii.1985, 1 ♀, B. E. Cooper leg.; 2 mi N Metcalf, 28.vi.1982, 1 ♂ 3 ♀♀, 4.vii.1984, 1 ♂ 1 ♀, B. E. Cooper leg. (all CNCI); Midland, 8.vii.1970, 1 ♀, J. T. Huber leg.; Peterboro[ugh] Co., Miller Creek C[onservation] A[rea], 1.vii.2000, 1 ♀, W. J. Crins leg. (both DEBU); Dufferin Co., Mono Cliffs Prov. Pk., Spillway Trail, 44°02'53"N 80°04'35"W, sweeping, 21.vi.2003, 26 ♂♂ 20 ♀♀ ( DEBU 00225619, -20, -22, -24, -26, -30–34, -36–38, -40–43, -45, -46, -48–53, -55–58, -62, -63, -65–70, -83–88, -90–92); Thunder Bay Distr., Neys Prov. Pk., beach nr. Little Pic River, sweeping, 16.vii.2002, 1 ♂ 1 ♀ ( DEBU 00253173, -74), all M. Buck leg.; ~ 8 km SSW Nipigon, Hwy#628, 48°57.09'N 88°19.71'W, sweeps, damp roadside, mixed graminoids, 16.vii.2008, 18 ♂♂ 60 ♀♀ ( CLEV, CMNH, CSCA 4 ♂♂ 6 ♀♀ each; 6 ♂♂ 42 ♀♀ CNCI), K. N. Barber leg.; North Bay, 8.vii.1954, 2 ♂♂ 1 ♀; N[orth] Bay, 18.vi.1964, 2 ♀♀, all A. H. Sturtevant leg. (all USNM); North Branch, 18.vii.1960, 2 ♀♀, S. M. Clark leg.; Lanark Co., N. Burgess Twp., 23.vi. 198[-], 1 ♂ 2 ♀♀, D. M. Wood leg.; Norway Point, Lake of Bays, 2.viii.1919, 1 ♀, J. McDunnough leg.; One-Sided [= Caliper] Lake, 19.vii.1960, 1 ♂, S. M. Clark leg., 26.vi.1960, 2 ♀♀, 29.vi.1960, 1 ♀, 16–17.vi.1960, 1 ♀ (genit. prep.), Kelton & Whitney leg. (all CNCI); Orangeville, 25.vi.1976, 1 ♀, 29.vi.1976, 1 ♂, E. A. Innes leg. ( DEBU); Ottawa, 11.vii.1938, 1 ♀, A. Brooks leg., 22.vii.1954, 2 ♀♀, W. R. M. Mason leg., 26.vii.1955, 2 ♀♀, J. G. Chillcott leg., 4.vii.1964, 1 ♂, J. R. Vockeroth leg. ( CNCI), 17.vii.1980, 1 ♂, Shelley Neilson leg. ( DEBU); Ottawa, Mer Bleu[e] Bog, 14.vi.1980, 11 ♂♂ 12 ♀♀, 15.vi.1980, 2 ♂♂ 1 ♀, 24.vi.1980, 23 ♂♂ 10 ♀♀, K. N. Barber leg. ( DEBU); Ottawa, Mer Bleue Bog, 45°23.7'N 75°30.8'W, sweeps, grasses on sand, 6.viii.2000, 1 ♀, K. N. Barber leg.; 5 mi E Ottawa, Mer Bleu[e], taken in Sphagnum bog, Malaise trap, 25.vi.1964, 1 ♀, D. D. Munroe leg. (both CNCI); Otter Rapids, 50°11.04'N 81°38.34'W, sweeps, grasses, herbs in disturbed area, 19.vii.2009, 4 ♂♂ 12 ♀♀; Otter Rapids, 50°10.91'N 81°38.54'W, sweeps, mostly Calamagrostis canadensis in hydro cut, 19.vii.2009, 6 ♂♂ 2 ♀♀, sweeps, mostly C. canadensis , E. ssuviatile in hydro cut, 20.vii.2009, 9 ♂♂ 10 ♀♀; Otter Rapids, 50°10.87'N 81°38.56'W, sweeps, mostly Calamagrostis canadensis in hydro cut, 20.vii.2009, 1 ♀; Otter Rapids, 50°10.85'N 81°38.65'W, sweeps, grasses in hydro cut, 20.vii.2009, 1 ♂ 4 ♀♀; Otter Rapids, 50°10.96'N 81°37.88'W, sweeps, grasses, herbs on roadside slope, 20.vii.2009, 15 ♂♂ 15 ♀♀, sweeps, mostly Schedonorus arundinaceus , on roadside slope, 20.vii.2009, 7 ♂♂ 8 ♀♀, all K. N. Barber leg. (all CNCI); Pancake Bay P. Pk., 46°57.74'N 84°42.63'W, sweeps, beach grasses, 7.viii.2004, 20 ♂♂ 61 ♀♀ ( DEBU 01501125–205, 2 ♂♂ genit. prep.), sweeps, Ammophila breviligulata on beach sand, 9.vii.2010, 8 ♂♂ 11 ♀♀ ( DEBU 01502501–19), K. N. Barber leg., sweeping, Ammophila breviligulata on beach sand, 9.vii.2010, 29 ♂♂ 31 ♀♀, J. Roháček leg. ( SMOC, 2 ♂♂ 2 ♀♀ genit. prep.); Pancake Bay P. Pk., 46°58.13'N 84°42.70'W, sweeps, open grassy area near highway, 7.viii.2004, 4 ♂♂ 6 ♀♀ ( DEBU 01501108–17); Pancake Bay P. Pk., 46°58.10'N 84°42.71'W, sweeps, mostly Carex nr. wetland boardwalk, 24. vii.2004, 1 ♀ ( DEBU 01500696), all K. N. Barber leg.; Pancake Bay P. Pk., 46°58.11'N 84°42.72'W, sweeping from boardwalk, mostly emergent sedges / Equisetum , 9.vii.2010, 1 ♂ 2 ♀♀, J. Roháček leg. ( SMOC, all genit. prep.); 10 km NW Penetanguishene,Awenda P. Pk., Second Lake, sweep sedge at shoreline, 13.vii.1992, 2 ♀♀, T.A. Wheeler leg. ( LEMQ 0039302, -09); Petawawa, 4.vii.1938, 2 ♀♀, H. S. Parish leg. ( USNM); Thunder Bay Distr., mouth of Pic River, N side, 48°36'N 86°18'W, 20.vii.2001, 1 ♂, M. Buck leg. ( DEBU 00169253); Pinery P. Pk., Grand Bend, 6.vii.1983, 1 ♂ 1 ♀, K. N. Barber leg. ( DEBU); ~ 1 km NW Ponsonby, 43°38.2'N 80°22.9'W, sweeps/pooter, roadside, mostly Poa pratensis & Poa compressa , 17.vii.1997, 13 ♂♂ 38 ♀♀ ( CSUC, EMEC 4 ♂♂ 6 ♀♀ each; CNCI 5 ♂♂ 26 ♀♀), K. N. Barber leg.; Lambton Co, Port Franks, Karner Blue Sanctuary, 15.viii.1996, 1 ♀, Malaise, 13–15.vi.1996, 1 ♂, 25–30.vii.1996, 1 ♂, J. Skevington leg. ( DEBU); Hwy 101 at Prairie Bee River (west side bridge), 47°51.81'N 83°54.33'W, sweeps, mostly Carex utriculata , 14.vii.2013, 1 ♀, K. N. Barber leg.; Prescott, 19.vi.1963, 1 ♀, J. R. Vockeroth leg. (both CNCI); Priceville, 15.vi.1959, 1 ♂, 5.vii.1960, 1 ♀; Primrose, 7.vii.1960, 1 ♂ 1 ♀, all D. H. Pengelly leg. (all DEBU); Pukaskwa N. P., SW of Admin. Bldg., 48°36'06"N 86°17'19"W, Vaccinium /lichen clearing, Malaise, 19–22.vii.2001, 1 ♂, M. Buck leg. ( DEBU 00182608); Puslinch, 16.vii.1979, 1 ♀, B. Merchant leg. ( DEBU); Rainy River, 22.vi.1960, 1 ♀, Kelton & Whitney leg. ( CNCI); 10 km S of Richmond, sweep in fen, 15.vii.1999, 1 ♀, S. E. Brooks leg. ( LEMQ 0040468); Wellington Co., Rockwood, sweeps, 17.vii.2004, 2 ♂♂ 3 ♀♀, M. Buck leg. ( DEBU 00335657, -58, -63, -65, -67); ~ 4.5 km E Rosseau, on Aspdin Rd., 45°15.88'N 79°34.88'W, sweeps, mostly Carex in sedge meadow, 7.vii.2005, 1 ♀; ~ 2 km E Rossport, Hwy#17, picnic area, 48°50.3'N 87°29.4'W, sweeps of graminoids, 9.vii.1999, 4 ♂♂ 6 ♀♀; ~ 2.4 km E Rutherglen, 46°16.1'N 79°00.0'W, sweeps, grasses on disturbed sand hill, 6.vii.2001, 1 ♂ 17 ♀♀; St. Joseph Island, Beech Beach, 46°07.1'N 83°53.5'W, sweeps, grasses on stable sand, 9.vii.2000, 4 ♀♀, all K. N. Barber leg. (all CNCI); St. Lawrence Is. N. Pk. [now Thousand Islands N. Pk.], Grenadier I[nformation] Centre, [each with] Code: 1-244K-132, -245, -252, -357, 26. vi.1975, 3 ♂♂ 1 ♀, H. J. Teskey leg., [each with] Code: 1-241H-9 to -13, 26.vi.1975, 1 ♂ 4 ♀♀, E. Sigler leg. ( CNCI); Hald[imand]-Norfolk Reg., Manester Tract, 6 km NNW St. Williams, 30.vi.2000, 2 ♂♂ 1 ♀, M. ParchamiAraghi leg. ( DEBU 00137124, -25, -43), 8.vii.2000, 1 ♀, M. Buck leg. ( DEBU 00137198), sandy neld, sweep, 8.vi.2001, 8 ♂♂ 12 ♀♀, M. Buck leg. ( DEBU 00158292–296, -299, -301, -303, -305, -308, -309, -312–314, -320, -321, -327, -334–336), 15.vi.2001, 3 ♂♂ 3 ♀♀, S. M. Paiero leg. ( DEBU 00158674, -684, -692, -694, -703, -708), sandy neld, yellow pans, 15.vi.2001, 1 ♀, Buck, Paiero & McKendry leg. ( DEBU 00158401); SSM [Sault Ste. Marie], 14.viii.1981, 1 ♂, G.Aiudi leg. ( DEBU); S[ault] S[te.] Marie, S. of Algoma U[niversity] College, 46°29.9'N 84°17.2'W, sweeps, Carex aquatilis , 28.vi.2002, 1 ♀, 29.vi.2002, 1 ♀, 1.vii.2002, 1 ♀, 31.vii.2002, 1 ♂, sweeps, mostly Carex aquatilis , 25.vi.2001, 1 ♀, 18.vii.2001, 1 ♀, 2.viii.2002, 1 ♀, 6.viii.2002, 1 ♀, sweeps, Calamagrostis canadensis , 26.vi.1998, 1 ♀, 2.viii.2002, 1 ♀, sweeps/pooter, Calamagrostis canadensis , 12.vii.2002, 1 ♀; same locality but 46°29.88'N 84°17.19'W, sweeps, mostly Carex aquatilis , 18.vii.2004, 1 ♀; same locality but 46°29.93'N 84°17.21'W, sweeps, Elymus repens , disturbed snow dump site, 6.viii.2008, 1 ♂ 2 ♀♀; S[ault] S[te.] Marie, Allen’s Side Road, 46°31.66'N 84°24.13'W, meadow sweeps above creek, 14.viii.2004, 2 ♂♂ 1 ♀, all K. N. Barber leg. (all CNCI); S[ault] S[te.] Marie, Baseline Rd., 46°31.41'N 84°24.57'W, sweeps, sparse veg. on disturbed site, 26.vi.2005, 15 ♂♂ 11 ♀♀ ( KNWR, MEMU 4 ♂♂ 4 ♀♀ each; CNCI 7 ♂♂ 3 ♀♀, 4 ♂♂ genit.prep.); same locality but 46°31.40'N 84°24.45'W, sweeps, edge of forest, Solidago , Rubus , Equisetum , grasses, 25.vi.2005, 3 ♂♂ 2 ♀♀, 10.vii.2005, 2 ♀♀; S[ault] S[te.] Marie, Bristol Pl. Pk., 46°30.8'N 84°16.6'W, sweeps/pooter, Phalaris arundinacea , 27.vii.1997, 2 ♀♀, 8.viii.1997, 1 ♂ (all CNCI), all K. N. Barber leg.; S[ault] S[te.] Marie, Finn Hill, 46°31.6'N 84°17.3'W, sweeps, graminoids/ composites, 19.vii.2005, 7 ♂♂ 13 ♀♀, sweeps, meadow graminoids, 1.vii.2002, 2 ♂♂ 1 ♀, sweeps, meadow graminoids/herbs, 20.vii.2004, 2 ♂♂ 36 ♀♀; same locality but 46°31.66'N 84°17.34'W, sweeps, mostly Calamagrostis canadensis , 6.vii.2008, 3 ♂♂ 3 ♀♀, 8.vii.2008, 5 ♂♂ 3 ♀♀, 10.vii.2008, 1 ♂ 2 ♀♀, sweeps, graminoids, herbs, composites, 6.vii.2008, 7 ♀♀, 10.vii.2008, 3 ♂♂ 12 ♀♀; same locality but 46°31.65'N 84°17.34'W, sweeps, graminoids, herbs, composites, 19.viii.2008, 8 ♂♂ 21 ♀♀ (all CNCI); same locality but 46°31.67'N 84°17.32'W, sweeps, Calamagrostis canadensis , 20.vii.2004, 6 ♀♀ ( CNCI), 30.vii.2004, 11 ♂♂ 26 ♀♀ ( MCZC, MTEC 4 ♂♂ 6 ♀♀ each; CNCI 3 ♂♂ 14 ♀♀); same locality but 46°31.63'N 84°17.33'W, sweeps, graminoids, herbs, composites, edge of Populus tremuloides , 6.vii.2008, 10 ♂♂ 16 ♀♀ ( CNCI), 8.viii.2008, 16 ♂♂ 18 ♀♀ ( NMPC, OSAC 4 ♂♂ 6 ♀♀ each; CNCI 8 ♂♂ 6 ♀♀), 10.vii.2008, 8 ♂♂ 14 ♀♀ ( CNCI), 25.vi.2009, 20 ♂♂ 25 ♀♀ ( PMAE, RBCM, SEMC 4 ♂♂ 6 ♀♀ each; CNCI 8 ♂♂ 7 ♀♀, incl. pair in copula), sweeps, Juncus effusus , 19.vii.2004, 1 ♀, sweeps, Scirpus cyperinus , 19.vii.2004, 2 ♂♂ 4 ♀♀, sweeps, Carex stipata stipata , 19.vii.2004, 1 ♂ 3 ♀♀, 20.vii.2004, 2 ♂♂ 6 ♀♀, 9.vii.2005, 1 ♀, 1.viii.2005, 3 ♂♂ 2 ♀♀ (1 ♂ genit. prep.), 25.vi.2009, 1 ♂, sweeps, mostly Carex stipata stipata , 26.vi.2007, 9 ♂♂ 13 ♀♀, 13.vii.2007, 1 ♂ 2 ♀♀ ( CNCI), K. N. Barber leg.; Sault Ste. Marie, Finn Hill, 46°31.63'N 84°17.33'W, sweeping boggy meadows, mostly Carex stipata stipata , 7.vii.2010, 19 ♂♂ 21 ♀♀ (2 ♂♂ 2 ♀♀ genit. prep., 2 ♀♀ photographed alive), 12.vii.2010, 4 ♂♂ 2 ♀♀ (1 ♂ 1 ♀ genit. prep.); same locality but 46°31.48'N 84°17.36'W, sweeping graminoid vegetation, 7.vii.2010, 1 ♂, sweeping, mostly Scirpus microcarpus , 7.vii.2010, 3 ♂♂ (1 ♂ genit. prep.), all J. Roháček leg. (all SMOC); S[ault] S[te.] Marie, nr. Greenwood P[ublic] S[chool], 46°34.1'N 84°21.2'W, sweeps, trail and railside vergetation, 17.vi.1999, 3 ♂♂ 4 ♀♀; S[ault] S[te.] Marie, Kinsmen Pk., 46°35.5'N 84°16.6'W, sweeps, trailside graminoids in mixed forest, 7.viii.1997, 1 ♀; S[ault] S[te.] Marie, Landslide Rd., Coldwater Ck. noodplain, 46°33.8'N 84°16.6'W, sweeps, mostly Carex sp., 7.viii.1997, 2 ♂♂ 3 ♀♀, sweeps/pooter, Calamagrostis canadensis , 7.viii.1997, 1 ♂ 2 ♀♀; S[ault] S[te.] Marie, Sault Coll[ege] Outdoor Lab, 46°32.1'N 84°18.2'W, sweeps, trailside grasses, Acer / Betula , 25.vii.1997, 1 ♀, sweeps, mixed meadow veg., 25.vii.1997, 3 ♀♀; S[ault] S[te.] Marie, 2 nd Line E, 46°32.3'N 84°16.6'W, sweeps, graminoids in open meadow, 26.vi.1999, 1 ♂, sweeps, trailside grasses, 26.vi.1999, 3 ♀♀; S[ault] S[te.] Marie, Hwy #17 city limits, 46°36.58'N 84°17.83'W, sweeps, Calamagrostis canadensis , in wet area, 16.viii.2004, 2 ♂♂ 5 ♀♀, sweeps, Calamagrostis canadensis , in wet area, 13.viii.2008, 3 ♀♀, sweeps, mostly Carex / Calamagrostis in wet area, 16.viii.2004, 3 ♀♀, 23.viii.2004, 4 ♂♂ 6 ♀♀ (1 ♂ genit. prep.), sweeps, mostly Carex / Calamagrostis in wet area, 19.vii.2008, 3 ♂♂ 3 ♀♀, sweeps, Carex / Calamagrostis , in wet area, 13.viii.2008, 1 ♀; same locality but 46°36.62'N 84°17.85'W, sweeps, Carex gynandra in alder thicket, 4.vii.2016, 2 ♀♀, all K. N. Barber leg. (all CNCI); ~ 10 km W S[ault] S[te.] Marie,Airport Rd., airport entrance, 46°29.7'N 84°29.2'W, sweeps, mowed grass, 1.viii.1999, 3 ♀♀, 19.vi.2000, 1 ♂ 3 ♀♀, 30.vi.2002, 1 ♂ 11 ♀♀ ( CNCI), 8.vi.2001, 31 ♂♂ 32 ♀♀ ( UBCZ, UCRC, UGCA, WFBM 4 ♂♂ 6 ♀♀ each; SMOC 4 ♂♂ 4 ♀♀, CNCI 11 ♂♂ 4 ♀♀, 1 ♂ genit. prep. & wing illustration, 1 ♂ 2 ♀♀ genit. prep.); ~ 10 km W S[ault] S[te.] Marie, Airport Rd., 46°29.0'N 84°29.2'W, sweeps, grasses in dry, sandy ditch, 25.vii.1999, 2 ♀♀; same locality but 46°29.9'N 84°28.9'W, natural gas r[igh]t-of-way, sweeps, low veg. sandy area, 1.vii.1999, 3 ♀♀, all K. N. Barber (all CNCI); same locality but 46°29.72'N 84°28.96'W, natural gas r[igh]t-of-way, sweeps, mostly Scirpus / Calamagrostis , 4.viii.2004, 4 ♂♂ 2 ♀♀, sweeps, mostly ferns, Equisetum arvense , 23.vii.2009, 1 ♀, sweeps, graminoids, Equisetum , herbs, 5.viii.2009, 1 ♂ 3 ♀♀, K. N. Barber leg. ( CNCI), sweeps, graminoids, composites, Equisetum , Rubus , ferns, 14.vii.2010, 6 ♂♂ 8 ♀♀, K. N. Barber leg. ( CNCI), 12.vii.2010, 14 ♂♂ 11 ♀♀, J. Roháček leg. ( SMOC, 1 ♂ 4 ♀♀ genit. prep.); MacIntyre Road, north of Sault Ste. Marie, 46°37.403'N 84°18.320'W, sweep, roadside, 23.vii.2011, 4 ♀♀, J. E. Swann & D. R. Edwards ( BDUC); Searchmont, N Hwy #552, 46°50.3'N 84°04.4'W, sweeps, roadside sedges / grasses, 10.vii.1998, 1 ♀; same locality but 46°51.3'N 84°02.4'W, sweeps, graminoids around gravel pit, 24.vii.1998, 1 ♀, both K. N. Barber leg.; 12.4 km NNE Searchmont, mi.10 Whitman Dam Rd., herb/grass meadow by Goulais R., 23.vi.1986, 2 ♀♀, 3. vii.1986, 1 ♀, D.J. M. Harvey leg., 23.vi.1986, 11 ♂♂ 13 ♀♀, K.N. Barber leg.(all CNCI); 18 km NNE Searchmont, mi.15 Whitman Dam Rd., grassy access road, 19.vi.1986, 1 ♀, 24.vi.1986, 1 ♀, 10.vii.1986, 2 ♀♀, 29.vi.1987, 2 ♀♀; ~ 18.8 km NNE Searchmont, Goulais River WS [Watershed], ~mi.15 Whitman Dam Rd., 46°55.7'N 83°56.2'W, sweeps, jackpine plantation, 29.vii.1999, 1 ♀, all K. N. Barber leg.; Sioux Narrows, 25.vi.1960, 2 ♀♀ (1 ♀ genit. prep.), Kelton & Whitney leg. (all CNCI); 20 km SW Smith’s Falls, sweeping meadow, 7.viii.1992, 2 ♂♂ 1 ♀, T. A. Wheeler leg. ( LEMQ 0039327, -32, -33); ~ 21 km NNE Smooth Rock Falls, 49°20.91'N 81°32.01'W, sweeps, Equisetum ssuviatile in wet ditch, 19.vii.2009, 2 ♀♀, sweeps, ditchside Equisetum spp. [including E. ssuviatile ], grasses, herbs, 19.vii.2009, 8 ♂♂ 10 ♀♀, 8.vii.2012, 11 ♂♂ 11 ♀♀, K. N. Barber leg.; S[outh] March, 31.vii.1968, 1 ♂, J. R. Vockeroth leg. (all CNCI); Thunder Bay Distr., Stanley, roadside nr. oaks, sweep, 23.vi.2001, 6 ♂♂ 2 ♀♀, S. A. Marshall leg. ( DEBU 00169950, -53–55, -60, -61, -44, -59); Thunder Bay District, Stanley Cemetery Prairie, 4.4 km SE Kakabeka Falls, grassy neld ( F [estuca] hallii), sweep, 27.vi.1998, 1 ♂ 5 ♀♀, Oldham & Bakowsky leg. ( DEBU 00155001, -05, -06, -09, -11, -12); 0.4 km S Stonecliffe, Pine Valley Rd. @ RR crossing, 46°11.9'N 77°52.8'W, sweeps, graminoids, 9.vii.2001, 9 ♂♂ 14 ♀♀, K. N. Barber leg.; Sydney Field Sta., nr. Foxboro, 8.vii.1970, 4 ♂♂ 12 ♀♀, J. F. McAlpine leg. (all CNCI);Algoma District, Thessalon, 12.vi.1965, 1 ♂ 1 ♀, K. P. Butler leg. ( LEMQ); Thessalon, 15.vi.1963, 1 ♀, R. G. Brumpton leg. ( DEBU); ~ 74 km NNE Thessalon, 46°53.94'N 83°16.23'W, shore of Mississagi R., sweeps, graminoids, herbs, Equisetum spp., 5.vii.2010, 1 ♂, 17.vii.2010, 2 ♂♂ 9 ♀♀, K. N. Barber leg. ( CNCI), sweeping graminoids and Equisetum spp.on muddy shore, 5.vii.2010, 1 ♀, J. Roháček leg. ( SMOC, genit.prep.); ca. 100 km W Thunder Bay at Portage Bay Rd., swept along road, 11.vii.1992, 1 ♂ 4 ♀♀, T.A.Wheeler leg. ( LEMQ 0039310, -365, -373, -374, -382); ~ 29 km SW Timmins, 48°19.12'N 81°44.79'W, sweeps, Carex spp./ Calamagrostis , 18.vii.2009, 3 ♂♂ 7 ♀♀; ~ 25 km W Upsala, 49°08.83'N 90°48.20'W, sweeps, wet area, mostly Calamagrostis canadensis , 30.vii.2008, 4 ♂♂ 6 ♀♀, all K. N. Barber leg. (all CNCI); Waubamic [sic Waubamik], 14.vi.1915, 2 ♂♂; Waubamick [sic Waubamik], [-]. vi.1915, 2 ♀♀, [-]. vii.1915, 2 ♂♂ 4 ♀♀, H. S. Parish leg. (all USNM); ~ 5 km S Wawa, Kinniwabi Pines Motel, 48°46.93'N 86°37.07'W, sweeps, mowed grasses on stable sand, 17.vi.2003, 1 ♀, K. N. Barber leg. ( CNCI); 100 km S Wawa, sweeping grass/ Typha along road, 30.vi.1992, 1 ♂ 8 ♀♀, T.A.Wheeler leg. ( LEMQ 0039296–301, -306, -371, -372); White River, 48°35.5'N 85°16.6'W, sweeps, mixed grasses edge of parking lot, 9.vii.1999, 1 ♂ 2 ♀♀; White River, “Black’s Cabin”, 48°36.71'N 85°15.61'W, sweeps, mowed grass/herbs on stable sand, 14.vi.2003, 2 ♂♂, all K. N. Barber leg. (all CNCI); 60 km S White River, sweeping grass/ Typha along road, 30.vi.1992, 1 ♀, T. A. Wheeler leg. ( LEMQ 0039311); Wiarton, 1.vii.1962, 1 ♀, G. Thorpe leg. ( CNCI); Wolf Is., 25.vii.1980, 1 ♂ 2 ♀♀, S. Beierl leg. ( DEBU); Wolf Lake env. nr. Dorset, peat-bog, sweeping over peat-bog meadow, 5.viii.1994, 1 ♀, J. Roháček leg. ( SMOC). PRINCE EDWARD ISLAND: Eglington Bay, sweep vegetation, 20.vii.1996, 3 ♂♂ 2 ♀♀, N. deVille leg. ( LEMQ 0039211–13, -218, -356); Greenwich N. P., 46°26.6'N 62°41.7'W, sweep open areas on trail and parking area, 3.viii.2004, 2 ♂♂ 3 ♀♀, S. Boucher leg. ( LEMQ 0040462–65, -91, 1 ♂ genit.prep.), 3.viii.2004, 1 ♀, V. Dion leg.( LEMQ 0040522); Red Point, 10.viii.1963, 1 ♀, R. L. Randell leg. ( LEMQ 0040478). QUEBEC: Aylwin Twp., between Aylwin Stn. & Marks Crossing, ~ 46°00'00"N 76°05'00"W, mixed forest with jack pine, sandy soil, 21.vi.2012, 1 ♂ 2 ♀♀, O. Lonsdale leg.; Beechgrove, 27.vi.1984, 2 ♂♂ 2 ♀♀, B. M. Bissett leg., 23.vi.1951, 4 ♂♂ 8 ♀♀, J. F. McAlpine leg., 7.vi.1955, 1 ♂, R. Lambert leg., 7.vi.1955, 1 ♂ 1 ♀, D. G. F. Cobb leg.; Beechgrove, 45°39'N 76°08'W, 29.vi.1962, 35 ♂♂ 33 ♀♀ (1 ♀ genit. prep.), 27.vi.1984, 1 ♂, 24.vi.1988, 3 ♂♂ 2 ♀♀, J. R. Vockeroth leg.; Chandler, 2.vii.1971, 1 ♀, B. V. Peterson leg. (all CNCI); Charlevoix, Ste-Aimé-des-Lacs, 47°41'N, 70°18.5'W, sweep vegetation, 5.viii.2001, 2 ♂♂ 1 ♀, S. Boucher leg.( LEMQ 0039161, -62, -68); Ferland-et-Boileau belvédère du Lac Ha!Ha!, 47°59.85'N 70°46.7'W, 518 m, sweep, 16.viii.2004, 1 ♀, M. Giroux leg. ( LEMQ 0040280); Gaspesie, l’Anse á Griffon, near Parc Forillon, 48°55'N 64°19'W, sweep, 31.vii.2001, 2 ♂♂ 2 ♀♀ ( LEMQ 0039176, -77, -80, -81); Gaspesie, Parc Forillon, Plage de Penouille, 48°51'20.9"N 64°24'45.3"W, sweep along trail, 16.viii.2006, 2 ♂♂ 3 ♀♀ ( LEMQ 0040302, -03, -05, -06, -08), all S. Boucher leg.; Gaspé, Forillon N. P., Cap des Rosiers, 48°50'N 64°12'W, sweep grass, 6.viii.2000, 2 ♀♀, H. Varady-Szabo leg. ( LEMQ 0039260, -61); Gaspé, Plage Haldimand, 48°47'N 64°22'W, sweep vegetation, 4.viii.2000, 6 ♂♂ 13 ♀♀, D. Raby leg. ( LEMQ 0039220–223, -226–233, -241, -242, -246, -379, -403, -409, -410), 8 ♂♂ 18 ♀♀, H. Varady-Szabo leg. ( LEMQ 0039357, -367–369, -375–378, -380, -381, -384–389, -392–399, -401, -405); Gaspé, Haldiman[d], St.-Jean R., 48°47'N 64°22'W, sweep salt marsh vegetation, 31.vii.2000, 1 ♀, H. Varady-Szabo leg. ( LEMQ 0039383); Gaspésie, near Routhierville, 48°10.9'N 67°08.8'W, sweep picnic area, 4.viii.2001, 1 ♂ 2 ♀♀, S. Boucher leg. ( LEMQ 0039184, -82, -83); Gatineau Pk., Harrington Lk., 10.vi.1954, 1 ♀, E. E. Sterns leg., 12.vi.1954, 1 ♂ 1 ♀, 16.vi.1954, 1 ♀, 22.vi.1954, 1 ♀, H. J. Huckel leg., 3.vii.1963, 17 ♂♂ 10 ♀♀, J. R. Vockeroth leg.; Gat[ineau] Nat. Pk., Ramsey Lake, 16.vii.1982, 4 ♂♂ 3 ♀♀ (1 ♂ genit. prep.); Gat[ineau] Nat. Pk., Ridge Road, 16.vii.1982, 1 ♀, all H. C. W.Walther leg. (all CNCI); Godbout, dunes near ferry dock, swept, eclector, (Universität Bielefeld, X 988), 24.viii.1994, 1 ♀, M. v. Tschirnhaus leg. ( ZSMC, in ethanol); Ile Bonaventure, 3 km from Côte de Percé, 48°30'N 64°10'W, sweep grass, 28.vii.2000, 21 ♂♂ 17 ♀♀, A. Thibault leg. ( LEMQ 0039205, -206, -209, -210, -244, -264–283, -285–288, -290–295, -303–305), 15 ♂♂ 10 ♀♀, H. Varady-Szabo leg. ( LEMQ 0039224, -225, -234–240, -243, -245, -247–253, -255, -256, -258, -259, -335, -337, -340); Îles de la Madeleine, Cap-aux-Meules, 47°22.04'N 61°52.26'W, sweep picnic area on coast of Cap-aux-Meules, 8.viii.2004, 1 ♀, S. Boucher leg. ( LEMQ 0040300); Îles de la Madeleine, Cap-aux-Meules, Chemin Chiasson, 47°21.7'N 61°55.86'W, sweep alfalfa neld by road, 6.viii.2004, 2 ♂♂, V. Dion leg. ( LEMQ 0040297, -99); Îles de la Madeleine, Cap-auxMeules, Chemin du Chiasson, 47°21.7'N 61°55.86'W, sweep neld along road, 6.viii.2004, 6 ♂♂ 4 ♀♀ ( LEMQ 0040511–20); Îles de la Madeleine, Cap-aux-Meules, L. Aucoin nr. Baie du Cap-Vert, 47°24.57'N 61°52.44'W, sweep neld, 9.viii.2004, 1 ♀ ( LEMQ 0040509); Îles de la Madeleine, Cap-aux-Meules, Havre-Aubert, “collines de la Demoiselle ”, 47°14.17'N 61°51.43'W, sweep, 5.viii.2004, 5 ♀♀ ( LEMQ 0040282, -283, -527, -528, -530), all S. Boucher leg.; Îles de la Madeleine, Havre-aux-Maisons, Chemin de la Pointe Basse, 47°23.93'N 61°48.65'W, sweep grass slope, 8.viii.2004, 1 ♂, V. Dion leg. ( LEMQ 0040508), 1 ♀, S. Boucher leg. ( LEMQ 0040309); Îles de la Madeleine, road 199 between Havre-aux-Maisons and Point-aux Loups, 47°30.36'N 61°44.14'W, sweep Ammophila at dunes, #6, 9.viii.2004, 1 ♀, S. Boucher leg. ( LEMQ 0040521); Îles de la Madeleine, Île de la Grande Entrée, Chemin du Bassin Ouest, 47°32.96'N 61°32.64'W, sweep neld at road, 9.viii.2004, 4 ♂♂ 7 ♀♀, V. Dion leg. ( LEMQ 0040479, -80, -82–90), sweep neld along road, 9.viii.2004, 3 ♂♂ 8 ♀♀, S. Boucher leg. ( LEMQ 0040492–496, -498, -499, -501, -504, -506, -507); Îles de la Madeleine, Sentiers au Havre-Aubert, Parc des bois brulés, 47°14.03'N 61°54.99'W, 5.viii.2004, 2 ♀♀ ( LEMQ 0040281, -535), S. Boucher leg., 3 ♂♂ 6 ♀♀ ( LEMQ 0040268, -87–94), V. Dion leg.; Lac Brulé, swept from “rugosa”, 25.vii.1947, 1 ♀, O. Peck leg.; Lac Phillipe, 45°37'N 76°W, 7.vii.1965, 3 ♂♂ 5 ♀♀, J. R. Vockeroth leg. (all CNCI); Lac Roddic, 16 km S Maniwaki, 22.vi.1991, 3 ♂♂ 3 ♀♀, M. Barták leg. ( MBPC, 1 ♂ genit. prep.); Gatineau Co., 3 mi SE Lake Duncan, near Masham, 8.vii.1979, 1 ♂, A. C. Pont leg. ( BMNH); Laniel, 1.vii.1944, 2 ♂♂ 2 ♀♀, A. R. Brooks leg., 1–3.vii.1963, 1 ♂, W. Gagne leg. ( CNCI); Laurentide Pk., 6.viii.1956, 1 ♀, A. H. Sturtevant leg. ( USNM); La Verendrye P. Pk., mi.139, Rte.58, 28.vi.1965, 1 ♂ 2 ♀♀, D. M. Wood leg. ( CNCI); Gatineau Co., Masham Township, 27.vii.1995, 2 ♂♂ 2 ♀♀, E. Ikeda leg. ( LEMQ 0039361, -62, -90, -91); Meach [sic Meech] Brook, Old Chelsea, 24.vi.1980, 3 ♂♂ 2 ♀♀, K. N. Barber leg. ( DEBU, 1 ♂ genit. prep.); Newport, 14.viii.1983, 2 ♂♂, B. M. Bissett leg. ( CNCI); Old Chelsea, 14.vi.1980, 1 ♂, K. N. Barber leg. ( DEBU); Old Chelsea, 24.vi.1956, 1 ♀; Old Chelsea, Summit King Mt., 1150', 18.vii.1961, 1 ♀, both J. R. Vockeroth leg. (both CNCI); Terrebonne Co., Lac Carre, Lot-31, Range 8, 19–23.viii.1968, 1 ♂, W. Boyle & R. Lalonde leg. ( LEMQ); Saguenay, Monts Valin, trail to Pic de la Tête de Chien, 48°34.89'N 70°52.23'W, sweep, 20.viii.2004, 1 ♀, M. Giroux leg. ( LEMQ 0040473). SASKATCHEWAN: Christopher Lake, 11.vii.1959, 1 ♀, A. & J. Brooks leg. ( CNCI); 50 mi E Regina, 1.vii.1980, 1 ♂, S. A. Marshall leg. ( DEBU, genit. prep.); Sturgis, 1.vii.1955, 51°56'N 102°32'W, 1 ♀, J. R. Vockeroth leg. ( CNCI). UNITED STATES OF AMERICA: GEORGIA: Rabun Co., Addie Branch, E. Fork Chattooga River, 2400', 1.viii.1957, 1 ♂; Rabun Co., Pine Mt., 1500', 1.viii.1957, 1 ♂, both J. G. Chillcott leg.; Rabun Bald, 9.viii.1957, 1 ♂ 1 ♀, W. R. Richards leg. (all CNCI). MAINE: Hancock Co., 800 m from Bass Harbor Head lighthouse, sweep wet clearing, 6.viii.2006, 1 ♀, J. Mlynarek leg. ( LEMQ 0040525); Caratunk, 2.viii.1950, 1 ♂ 1 ♀; Chebeag[ue] Is., 17.vii.1962, 1 ♂ 4 ♀♀, all A. H. Sturtevant leg. (all USNM); Lincoln Co., East Boothbay, 43°51'N 69°35'W, sweep stream at roadside, 9.viii.2000, 1 ♂ 3 ♀♀, T.A. Wheeler leg. ( LEMQ 0039341, -46–48), 2 ♀♀, J. Savage leg. ( LEMQ 0039349, -50), 1 ♀, J. Forrest leg. ( LEMQ 0039406); Eddington, 11.viii.1956, 1 ♂ 1 ♀; Mt. Desert, 19.vii.1962, 2 ♀♀, all A. H. Sturtevant leg.; Pittston, 3.viii.1930, 1 ♂; Seal Harbor, 29.vii.1930, 1 ♀, both A. L. Melander leg. (all USNM); Seal Harbor, Mt. Desert, 13.viii.1958, 3 ♀♀, J. R. Vockeroth leg. ( CNCI); Weld, 15–17.viii.1940, 2 ♂♂ 1 ♀, A. Stone leg. ( USNM, 1 ♀ with Sabrosky det. as “ gracilis auctt.”); Hancock Co., Winter Harbor–Schoodic, Diptera Blitz, Lot #32, 15–16.vii.2006, 1 ♀, [no collector] ( CLEV). MASSACHUSETTS: Middlesex Co., Belmont, Rock Meadow, 24.vii.1981, 2 ♀♀, N. E. Woodley leg.; Middlesex Co., Lincoln, Malaise trap, 10.viii.1982, 1 ♂ 1 ♀, E. T.Armstrong leg.; Pelham, 10.vii.1954, 2 ♀♀ (1 ♀ with det. as Anthomyza gracilis ), A. H. Sturtevant leg. (all USNM). MICHIGAN: Brevort, 2.viii.1936, 1 ♀; Champion, 27.vi.1940, 3 ♂♂ 1 ♀, all C. W. Sabrosky leg.; Cusino, 26–27.vi.1940, 2 ♀♀, T. F. Boyce leg. (all USNM, all with S. W. Frost det. as Anthomyza gracilis ); Cheboygan Co., 25.vi.1934, 1 ♂, Dobrovolny leg. ( USNM, with Steyskal det. as Anthomyza gracilis ); Cheboygan Co., 24.vii.1955, 1 ♂, W. J. Hanson leg., 1.viii.1933, 1 ♀, 29. vi.1936, 1 ♀, H. B. Hungerford leg., 1 ♀, H. Leighton leg., 26.vii.1935, 1 ♀, L. R. Penner leg., 11.vii.1933, 2 ♀♀, H. Peters leg. ( SEMC), 6.vii.1933, 1 ♀, H. Peters leg. ( USNM), 4.vii.1940, 1 ♂ 2 ♀♀ (1 ♂ genit. prep.), 22.vii.1940, 1 ♀, R. I. Sailer leg. ( SEMC); Cheboygan Co., Douglas Lake, 1.viii.1950, 1 ♀, R. I. Sailer leg. ( USNM); Douglas Lake, 9.vii.1931, 2 ♀♀, C. W. S[abrosky] leg. (with S. W. Frost det. as Anthomyza gracilis ), 18.vii.1943, 1 ♂, C. W. Sabrosky leg.; Houghton Co., 11.viii.1953, 1 ♀, R. R. Dreisbach leg.; Hunt Ck. Exp. Sta., nr. Lewiston, 20. vii.1942, 3 ♂♂ 1 ♀, C. W. Sabrosky leg. (all USNM); Ironwood, 46°27.9'N 90°10.1'W, sweeps, grasses /herbs, 22.vii.1999, 2 ♂♂ 16 ♀♀, K. N. Barber leg. ( CNCI); Keweenaw Co., 26.vi.1955, 1 ♂, R. R. Dreisbach leg.; Keweenaw Co., Isle Royale, 15., 17.vii.1938, 1 ♂, 16., 18.vii.1938, 1 ♂, G. Steyskal leg.; Midland Co., 15.vii.1947, 1 ♂; Otsego Co., 24.vii.1955, 1 ♂, both R. R. Dreisbach leg.; Rapid City, 10.vii.1941, 1 ♀, C. W. Sabrosky leg. (all USNM); Sault Ste. Marie, 27.vii.1960, 1 ♀, Kelton & Whitney leg. ( CNCI). MINNESOTA: Eaglesnest, 11.vii.1952, 1 ♀ ( INHS 40,086), 30.vi.1959, 1 ♀ ( INHS 40,090), 4.viii.1958, 1 ♂ 2 ♀♀ ( INHS 40,091, -116, -117), 10.viii.1957, 1 ♀ ( INHS 40,093), 11.vii.1959, 2 ♀♀ ( INHS 40,096, -110), 3.ix.1958, 1 ♀ ( INHS 40,097), 22.vii.1957, 1 ♀ ( INHS 40,101), 6.vii.1959, 1 ♀ ( INHS 40,104), 30.vii.1959, 2 ♀♀ ( INHS 40,114, -115), 13.vii.1957, 2 ♀♀ ( INHS 40,118, -120), 12.vii.1957, 1 ♂ 1 ♀ ( INHS 40,123, -124), 28.vi.1957, 2 ♀♀ ( INHS 40,125, -127), 20.viii.1958, 7 ♀♀ ( INHS 40,132–138), 23.vii.1958, 1 ♂ 1 ♀ ( INHS 40,139, -141), 30.vii.1958, 1 ♂ 4 ♀♀ ( INHS 40,145, -148–150, -151), 1.vii.1957, 1 ♂ 8 ♀♀ ( INHS 40,152–160), all W. V. Balduf leg. (all part of mixed series [with A. furvifrons ] with Sabrosky det. as Anthomyza gracilis ); Stearns Co., 1 mi E Saulk Center, 4.vii.1994, 1 ♀, D. E. Hansen leg. ( CNCI); Clearwater Co., #15, Sucker Lake, 3 mi W Lake Itasca P. O., 1550', 7.vii.1970, 1 ♀, G. W. Byers leg. ( SEMC); Aitkin Co., 14 mi W Willow River, 46.334°N 93.096°W, 8.vii.1993, 1 ♀, 19.vii.1995, 2 ♀♀, 1.vii.1995, 2 ♀♀, 28.vii.1995, 4 ♂♂ 2 ♀♀ (1 ♀ genit. prep.), 29.vi.1997, 1 ♂, 5.vii.1997, 1 ♀, 20.vii.1997, 1 ♀, 3.viii.1997, 1 ♂, D. E. Hansen leg. ( CNCI). NEW HAMPSHIRE: Alstead, 4.viii.1956, 1 ♂, A. H. Sturtevant leg.; Benton, 6.vii.1931, 1 ♂, A. L. Melander leg.; Grafton Co., Easton, 24.vi.1976, 1 ♂, F. C. & B. J. Thompson leg.; Franconia, 6.vii.1936, 1 ♂, A. L. Melander leg. (all USNM); Franconia, [no date], 2 ♀♀, Mrs. Slosson leg. ( USNM, 1 ♀ with det. as Anthomyza gracilis ), 2 ♀♀, Mrs.A. T. Slosson leg. ( AMNH Ac. 26226, with dets. as Anthomyza gracilis and A. ssavipes ); Gorham, 15.vii.1957, 2 ♀♀, J. R. Vockeroth leg.; Mt. Madison, Dolly Copp Campground, 8–9.vii.1986, 1 ♂ 2 ♀♀ (1 ♀ genit. prep.), J. M. & S. H. Cumming leg.; Mt. Washington, el. 5000', 13.viii.1951, 1 ♀, G. S. Walley leg. (all CNCI); Mt. Washington, tundra at summit, 29.vii.1961, 1 ♀, W. W. Wirth leg. ( USNM); Mt. Washington, Summit Flats, 5900–6200', 2.viii.1954, 1 ♀, Becker, Munroe & Mason leg. ( CNCI); Peterborough, 10.vi.1950, 2 ♂♂, E. C. Zimmermann leg. ( BMNH, 1 ♂ genit. prep.); Rindge, 27.vi.1979, 1 ♂, B. Mather leg. ( MEMU); Coos Co., 1 km E Stark, 44°36'N 71°24'W, sweep along Ammonoosuc R., 8.viii.2000, 1 ♀, J. Savage leg. ( LEMQ 0039408); Grafton Co., White Mountain N. Forest, Kancamagus Pass, 44°01'N 71°29'W, sweep, 10.viii.2000, 1 ♂ 1 ♀, J. Forrest leg. ( LEMQ 0039400, -407); White Mts., Dolly Copp, 11.vii.1931, 4 ♂♂ 2 ♀♀, A. L. Melander leg. ( USNM). NEW YORK: Adirondacks, Avalanche Trail, 30.vii.1929, 1 ♀, A. L. Melander leg. ( USNM); Adirondack Pk., Blue Mountain, 43°52.5'N 74°25.8'W, sweep grasses at summit, 31.vii.2002, 1 ♂, J. Forrest & T. A. Wheeler leg. ( LEMQ 0040379); Lake Placid, 28.vii.1929, 2 ♂♂, A. L. Melander leg. ( USNM); Lake Placid, 2000', 19.vii.1962, 2 ♂♂ 2 ♀♀; Franklin Co., Paul Smiths, 20.vii.1962, 1 ♀, all J. R. Vockeroth leg.; St. Lawrence Co., ~ 25 km SE S. Colton, 44°19.07'N 74°43.42'W, sweeps, mostly Carex , edge of wetland, 27.vii.2006, 6 ♂♂ 4 ♀♀ (2 ♂♂ genit. prep.), K. N. Barber leg. (all CNCI); Schoharie Co., 32, 6.vii.1974, 1 ♀, P. Gargiullo leg.( UGCA); Whiteface Mt., 4600–4872', 19.vii.1962, 1 ♂ 1 ♀, J. R. Vockeroth leg.; Whiteface Mt., summit, 5700', 19.vii.1962, 1 ♀, J. G. Chillcott leg. (all CNCI); Wilmington, 30.vi.1922, 1 ♂, J. M.Aldrich leg. ( USNM). NORTH CAROLINA: Clingman’s Dome, 5.viii.1957, 1 ♂; Highlands, 31.v.1957, 1 ♂, 15.vi.1957, 1 ♂, 12.viii.1957, 1 ♀, 13.viii.1957, 2 ♂♂ 1 ♀, 21.viii.1957, 1 ♂ 1 ♀, 22.viii.1957, 2 ♀♀, all W. R. Richards leg.; Highlands, 3800', 17.viii.1957, 1 ♂ 1 ♀, 21.viii.1957, 1 ♀; Highlands, Horse Cove Bog, 3000', 18.vii.1957, 2 ♂♂, all J. G. Chillcott leg.; Highlands, Whiteside Mt., 21.viii.1957, 1 ♀, C. J. Durden leg.; Highlands, Whiteside Cove, 18.viii.1957, 6 ♂♂ 19 ♀♀ (1 ♂ genit. prep.), W. R. Richards leg.; Highlands, Whiteside Cove, 2600', 17.viii.1957, 1 ♀, 2800', 11.viii.1957, 1 ♂ 1 ♀; Mollie Gap, Blue Ridge Pkwy., Heintooga Road, 5350', on blossoms Smilacina sp., 8.vi.1965, 6 ♂♂ 2 ♀♀ (1 ♀ genit. prep.), all J. G. Chillcott leg.; Haywood Co., GSMNP [Great Smoky Mountains National Park], Purchase Knob, 17S 312178E 3939620N, 1463 m, forest edge, hillside between house & Ferguson Cabin, 11.vi.2008, 1 ♀, B. J. Sinclair leg. (all CNCI); Avery Co., Pisgah N. Forest, Grassy Ridge Bald, 1863 m, 36.10246°N 82.08066°W, sweep grassy bald, ( GRAS S 3), 26.v.2008, 1 ♀, T. A. Wheeler leg. ( LEMQ). PENNSYLVANIA: Bradford Co.m Wilawana, 21.vii.1991, 1 ♀, R. H. Crandall leg. ( LACM ENT 329117). TENNESSEE: Blount Co., SW corner of Cades Cove, Great Smoky Mts. N. Pk., 23.v.2001, 1 ♂ 6 ♀♀ (1 ♀ genit. prep.), swept from marsh vegetation, 24.v.2001, 2 ♀♀ (1 ♀ genit. prep.), swept from Eleocharis , 24.v.2001, 1 ♂ (genit. prep.), all B. A. Foote leg. (all CMNH); Gregory Bald, GSMNP [Great Smoky Mts. N. Pk.], sweeps, 1.vi.1979, 1 ♂ 1 ♀, M. J. Sharkey leg. ( DEBU); Colke [sic Cocke] Co., Hartford, 24.vi.1986, 1 ♀, J. D. Pinto leg. ( CSCA); Indian Gap, 24.vii.1957, 1 ♀, W.R. Richards leg. ( CNCI). VERMONT: Island Pond, 17.vi.1975, 1 ♂, H. J. Teskey leg. ( CNCI). WEST VIRGINIA: Blackwater St. Pk., daisy sweep, 3.vii.1986, 1 ♂ 2 ♀♀, Davis leg. ( DEBU, 1 ♂ genit. prep.). WISCONSIN: Columbia Co., Swan Lake, 4 mi E Portage, taken on Andropogon sp., 16.vii.1963, 3 ♂♂ 2 ♀♀, Stannard & Smith leg. ( INHS 40,220–224); Washburn Co., T39N R12W B28, 28.vi.1953, 2 ♂♂, T39N R12W B32, 25.vi.1953, 1 ♂ 2 ♀♀, T39N R12W B32, 2.vii.1953, 1 ♂ 5 ♀♀, T39N R12W B32, 3. vii.1953, 1 ♀, T39N R12W B32, 4.vii.1953, 1 ♂, T39N R12W B32, 10.vii.1953, 1 ♀, T39N R13W B30, 4.vii.1953, 1 ♂, R. H. Jones leg. ( USNM).

Other material examined (not included in type series). CANADA: MANITOBA: Aweme, 8.viii.1918, 1 ♀ 1 spec., N. Criddle leg. ( CNCI, 1 ♀ missing wings, 1 spec. missing abdomen and most legs);Aweme, Criddle farm, 49°42.5'N 99°36.1'W, sweep grasses near house, 8.vii.2000, 1 spec., V. Crecco leg. ( LEMQ 0039503, abdomen missing); 5 mi SW Shilo, swept from open prairie, 11.vii.1958, 1 ♂, J. G. Chillcott leg. ( CNCI, headless). ONTARIO: Belwood, 23.vi.1968, 1 ♂ 1 spec., D. H. Pengelly leg. ( DEBU, 1 ♂ bleached, 1 spec. tip of abdomen missing); Blackburn, 12.vii.1954, 1 ♂ (headless); Blackburn, Mer Bleue, 27.vi.1955, 2 ♀♀ (both headless) all W. R. M. Mason leg. (all CNCI); Bruce Peninsula N. P., Singing Sands, 45°11.6'N 81°34.7'W, sweeps, Agropyron / Calamagrostis , 5.vii.1998, 1 ♀, K. N. Barber leg. ( DEBU, T 7+ S 7 deformed); ~ 66 km NNW Elliot Lake, Rocky Island Lake, 46°50.82'N 83°08.76'W, 405 m, sweeping Scirpus sp.on dried shoreline, 4.vii.2010, 1 ♂, J. Roháček leg.( SMOC, without wings); Kenora Co., Kenora, UTM 183789, bur oak prairie sweep, 22.vi.1998, 1 spec. (abdomen missing), M. J. Oldham & W. D. Bakowsky leg.; Midland, 25.vi.1970, 1 ♀ (left wing missing, overglued), J. T. Huber leg. (both DEBU); Hald[imand]-Norfolk Reg., Manester Tract, 6 km NNW St. Williams, sandy neld, sweep, 8.vi.2001, 1 ♀, M. Buck leg. ( DEBU 00158331, wings missing); Pancake Bay Prov. Park, 46°57.74'N 84°42.63'W, sweeping Ammophila breviligulata on beach sand, 9.vii.2010, 1 ♂ 1 ♀, J. Roháček leg. ( SMOC, both headless); Sault Ste. Marie, Finn Hill, 46°31.63'N 84°17.33'W, sweeping boggy meadows, mostly Carex stipata stipata , 7.vii.2010, 3 ♂♂, J. Roháček leg. ( SMOC, used for molecular work, 1 ♂ genit. prep.); S[ault] S[te.] Marie, Sault Coll[ege] Outdoor Lab, 46°32.1'N 84°18.2'W, sweeps, mixed meadow veg., 18.vi.1998, 1 ♂, K. N. Barber leg. ( CNCI, epandrium incompletely developed on left side); ~ 10 km W Sault Ste. Marie, Airport Rd., 46°29.72'N 84°28.96'W, natural gas r[igh]t-of-way, sweeping graminoids, composites, Equisetum , Rubus , ferns, 12.vii.2010, 1 ♀, J. Roháček leg. ( SMOC, headless); Waubamic [sic Waubamik], Parry S[oun]d, 7.vii.1915, 1 ♀, J. M. Aldrich leg. ( USNM, head & thorax crushed). QUEBEC: Beechgrove, 27.vi.1984, 1 ♂, B. M. Bissett leg. ( CNCI, head crushed, left wing missing); Gaspesie, Parc Forillon, “Le castor ”, 48°50'30"N 64°14'25"W, sweep veg. near marsh, 16.viii.2006, 1 ♂, S. Boucher leg. ( LEMQ 0040307, abdomen and wings dirty); Gatineau Pk., Harrington Lk., 3.vii.1963, 1 ♂, J. R. Vockeroth leg. ( CNCI, headless); Grandes-Bergeronnes, 167 Route 138, 29.vi.1997, 1 ♂, N. Luth leg. ( DEBU, dirty, wings tattered, genit. prep.). UNITED STATES OF AMERICA: MAINE: SW Harbor, 13.vii.1918, 1 spec. (abdomen missing, with det. as Anthomyza gracilis ), C. W. Johnson leg.; SW Harbor, 14.vii.1918, 1 spec. (abdomen missing), [C. W. Johnson?] leg.; Weld, 21.viii.1952, 1 ♂ (missing legs), A. Stone leg. (all USNM). MICHIGAN: Cheboygan Co., 4.vii.1940, 1 ♀ (dirty), R. I. Sailer leg.; Cheboygan Co., 6.vii.1933, 1 ♀ (head and thorax damaged), H. Peters leg. (both SEMC); Cusino, 26–27.vi.1940, 1 ♀, T. F. Boyce leg. ( USNM, left wing missing with other damage, abdomen glued to point, with S. W. Frost det. as Anthomyza gracilis ). MINNESOTA: Clearwater Co., #15, Sucker Lake, 3 mi W Lake Itasca P. O., 1550', 7.vii.1970, 1 ♀, G. W. Byers leg. ( SEMC, bleached, right wing missing). NEW HAMPSHIRE: Alstead, 4.viii.1956, 1 ♀ (headless) 1 spec. (missing head & abdomen), A. H. Sturtevant leg.; White Mts., Dolly Copp, 11.vii.1931, 1 spec. (abdomen missing), A. L. Melander leg. (all USNM). NEW YORK: Adirondacks, Connery Pond, 15.vii.1938, 1 ♀ (headless), A. L. Melander leg.; Lake Placid, 29.vii.1922, 1 ♂, J. M. Aldrich leg. (both USNM, dirty, wings missing or wrapped on pin). TENNESSEE: Gregory Bald, GSMNP [Great Smoky Mts. N. Pk.], sweeps, 1.vi.1979, 1 ♀, M. J. Sharkey leg. ( DEBU, dirty, abraded).

Description. Male. Total body length 2.06–2.54 mm; general colour as in A. furvifrons but more densely and paler grey microtomentose ( Fig. 520 ). Head more elongate than in all relatives, distinctly longer than high ( Figs 520 , 547 ), subquadrangular, with face strongly receding in pronle and frons projecting in front of eye ( Fig. 520 ), dark brown and yellow. Occiput dorsomedially slightly but distinctly concave, dark brown to black with distinct greyish microtomentum. Frons relatively narrow ( Figs 521 , 550 ), longer than in relatives, medially dull blackish brown (at most up to anterior third of frons), only frontal triangle dark grey microtomentose; anterior third or more of frons orange-yellow; orbits largely yellow and silvery white microtomentose, more posteriorly (behind posterior ors) gradually brownishdarkened and with silvery grey microtomentum. Frontal triangle narrow, reaching at most to midpoint of frons. Frontal lunule yellow, whitish microtomentose. Face, parafacialia and gena yellow to orange-yellow and mostly dull. Face very narrow, medially weakly sclerotized, more or less furrowed and somewhat darkened. Parafacialia relatively broad and (like gena) with silvery white microtomentum (most dense on vibrissal angle). Both parafacialia and gena with rather broad but relatively pale (dark yellow to ochreous) marginal stripe. Postgena ventrally yellow and whitish microtomentose, dorsally brown and greyish microtomentose merging with similar vestiture on occiput ( Fig. 520 ). Mouthparts (including palpi) similarly coloured to those of A. furvifrons . Cephalic chaetotaxy (all macrosetae relatively short): pvt small, convergent, with apices meeting or crossed; vti, vte (sometimes shorter), oc (sometimes longer than others) and posterior ors subequal, longest of cephalic setae; 3 ors but only 2 long, widely spaced, middle ors slightly to distinctly shorter than posterior, and anterior ors reduced to setula (rarely somewhat enlarged); 1–2 pairs of medial microsetulae in the anterior third of frons; small setula behind vte not longer than postoculars, the latter (7–8) small, in single row; postgena with a few setulae and 2 (1 longer) usual posteroventral setae; vi about as long as middle ors but nner; subvibrissa very small, weak, slightly or not longer than foremost peristomal setula; only 3–4 nne peristomal setulae. Palpus slightly less clavate than that of A. furvifrons but similarly setose. Eye moderately convex, more elongate than in relatives, of almost pyriform outline (see Fig. 520 ), anteroventrally most widened, with longest diameter elongately oblique and 1.4–1.5 times as long as the shortest. Smallest genal height about 0.15–0.16 times as long as shortest eye diameter. Antenna with all segments yellow; 1st nagellomere nattened laterally, with very short white pilosity. Arista brown to dark brown including basal segments, 1.9–2.0 times as long as antenna, with cilia slightly longer than those on nrst nagellomere.

Thorax hardly narrower than head; mesonotum entirely blackish brown, pleural part of thorax usually brown except propleuron rarely yellow at least ventrally, sternal midline area also rarely yellow; mesonotum and pleurae (mostly) very densely pale grey (often with some bluish tinge) microtomentose and dull ( Figs 520 , 547 ). Thoracic chaetotaxy (all macrosetae relatively weak): 1 hu (slightly shorter than anterior npl) and 1–2 microsetae on humeral callus; 2 npl (anterior longer); 1 small prs (shorter than hu); 1 sa (shorter than pa); 1 moderate pa; 2 postsutural dc (the shorter anterior about as long as or longer than anterior npl) and 5–8 dc microsetae in front of them, the hindmost dc microseta often longer than others; ac microsetae sparse, in 3–4 rows on suture, in 2 rows between anterior dc and usually no ac microseta beyond them; 2 sc, laterobasal sc very weak, about as long as but thinner than prs; apical sc subequal to posterior dc (both longest thoracic setae); 1 minute ppl; 2 long stpl (anterior usually slightly shorter) and 0–2 setulae in dorsal half of sternopleuron; its ventral part with 5–6 longer but pale setae. Scutellum rounded triangular, distinctly convex dorsally. Legs almost entirely brightly to light yellow, sometimes with shiny brownish darkening on base of cx 1, only 1–2 terminal segments of all tarsi pale brown to brown. f 1 with ctenidial spine usually distinctly longer than (rarely equal to) maximum width of t 1. Other pedal chaetotaxies as described for A. furvifrons . Wing ( Fig. 525 ) long and narrow (usually slightly narrower than those of relatives), with pale brown veins and hyaline unicolourous (ochreous to brownish) membrane. Wing venation mostly as in A. furvifrons only R 4+5 usually very slightly bent (recurved) and running closer to M. Discal (dm) cell often narrower and less widened distally, with r-m situated somewhat in front of the middle of dm cell. Apical portion of CuA 1 usually longer than dm-cu and alula narrower. Wing measurements: length 2.14–2.62 mm, width 0.61–0.78 mm, Cs 3: Cs 4 = 1.16–1.68, rm\dm-cu: dm-cu = 2.44–3.53. Haltere pale yellow to whitish yellow, sometimes with partial ochreous darkenings.

Abdomen slender, elongate ( Fig. 520 ), generally formed, coloured and setose as that of A. furvifrons unless mentioned otherwise. Preabdominal sterna lighter than those of A. furvifrons , pale ochreous yellow, only S1, S2 (darkest) and S5 (partly, laterally) brownish. T6 yet larger (longer) than in A. furvifrons , usually more than half length of T5, largely brown but with a narrow unpigmented area anteromedially or medially across sclerite (hence sometimes seemingly divided into two sclerites), often with a few setae at posterior margin. S6 (paler) and S7 (darker) brown, both with more sclerotized and blackish brown anterior margin, each with 2 (rarely 3) setae; S8 as in A. furvifrons .

Genitalia most similar to those of A. equiseti . Epandrium ( Figs 551, 552 ) about as broad as high thus slightly higher than that of A. equiseti , blackish brown and somewhat more densely setose than in latter species, with 1 dorsolateral pair of (not much) longer and thicker setae; anal nssure distinctly narrower ( Fig. 551 ) than in A. equiseti . Cercus moderately narrow, with apex more or less pointed; setae nne and numerous, the longest apical slightly longer than subapical. Medandrium ( Figs 551 ) wider and not as high as that of A. furvifrons , in pronle somewhat constricted in the middle ( Fig. 552 ), dorsolateral corners small. Gonostylus ( Figs 551, 552, 559 , 563 ) small, most similar to that of A. equiseti thus moderately bent medially, with simple acute apex, micropubescence on most of outer side and setae on inner side of both anterior (there more than in A. equiseti ) and posterior margins ( Figs 559 , 563 ) but somewhat more elongate and with anterior margin more concave than in the latter species. Hypandrium ( Fig. 553 ) relatively elongate, with anterior internal lobes reduced, membranous; its posterior arms nrmly fused with transandrium. Transandrium ( Fig. 554 ) relatively strong and well sclerotized, posteromedially with broad, nat but dorsomedially weakly sclerotized caudal process terminating in distinctive trough-like appendage ( Figs 553–555 , 561 ). This appendage smaller and narrower than in A. equiseti and with teeth on ventrolateral margins reduced or absent ( Figs 553 , 561 ), only those on posterior corners ( Figs 554, 555 ) well developed. Pregonite ( Figs 553 , 560 ) with anterior part angularly projecting medioventrally (this process distinctly larger than in A. equiseti ) and bearing 3 setae (as in relatives), 1 long, 1 shorter and 1 minute; posterior part of pregonite reduced and hairless. Postgonite ( Fig. 553 ) slender, more elongate and bent than that of A. equiseti , with moderately pointed apex and 1 anterior setula in distal fourth in addition to granular sensillae; its proximal part attached to large basal sclerite provided with several (5–8) ventral spines. Armature of basal membrane incorporated into ventral appendage of caudal process of transandrium. Aedeagal part of folding apparatus with similarly spinose walls as those of A. equiseti : right wall dorsally with a large cluster of strong dark spines, ventrally with only a number of spine-like tubercles ( Fig. 558 ); left wall with only 2–3 long and dark spines in dorsal cluster, ventrally there are only smaller spines and tubercles ( Fig. 556 ). Connecting sclerite slender, only dorsally dilated and with granulous structure, well sclerotized on apex armed with 1–2 dark spines surrounded by a few pale tubercles ( Fig. 556 ). Phallapodeme slender with basal part dilated and deeply forked, fulcrum rather long but relatively slender, apex with small lateroventral corners. Aedeagus with short, frame-like phallophore fused to distiphallus as in relatives. Distiphallus basally with dorsal and ventral elongate sclerites; bind from about two-nfths. Saccus proximally well sclerotized (more on right side, Fig. 556 ) and its dilated distal membranous part provided with rather variable armature of 2 ( Fig. 562 ) to 4 robust black spines ( Fig. 556 ). Filum formed by single sclerite, only partly pigmented, very nnely spinulose in proximal and middle part, subterminally only somewhat widened without corner or projection, with terminal part slender; apical part gradually tapered into sinuate, very slender and acute point with 3–4 spinulae ( Fig. 557 ). Ejacapodeme small, slender and pale, with elongate digitiform projection ( Fig. 556 ).

Female. Similar to male unless mentioned otherwise. Total body length 2.26–3.18 mm. Antenna with 1st nagellomere anterodorsally (on both sides) more or less extensively ochreous to brownish-darkened ( Figs 545 , 547 ), sometimes leaving only narrow proximal and ventral parts yellow. Palpus usually darkened to brown or grey apically. Marginal stripe of parafacialia and particularly gena darker ochreous to brown. Yellow areas on thoracic sternal midline and propleuron more common and more extensive in some populations, and may include margins of the humerus and margins and isolated spots on the mesopleuron. Wing measurements: length 2.46–3.26 mm, width 0.69–1.02 mm, Cs 3: Cs 4 = 1.23–1.47, rm\dm-cu: dm-cu = 2.25–3.27. Abdomen ( Fig. 547 ) similarly formed, coloured and setose to that of A. furvifrons but T3–T5 not longer than T2, subequal in length but T4 slightly wider than T3 or T5. Preabdominal sterna S2–S5 somewhat narrower than in male; only S1 (wider than long) and S2 (as long as wide) brownish, S3–S5 (and S6) light ochreous yellow. S2–S4 becoming only slightly larger and wider posteriorly, S3–S5 as long as broad or slightly longer than broad, S5 usually narrower than S4.

Postabdomen ( Figs 566–568 ) closely resembling that of A. equiseti , elongate, slender, gradually tapered posteriorly. T6 large, broad but narrower than T5, brown with posterior margin more or less extensively unpigmented, setose in posterior three-fourths. S6 slightly transverse, wider posteriorly, with anterior corners rounded, pale ochreous thus lighter than in A. equiseti , setose in posterior half. Tergosternum T7+S7 conically ring-shaped, blackish brown laterally ( Figs 564 , 566 ) and dorsally with narrow posteromedial incision ( Fig. 567 ). Ventral part of T7+S7 with relatively large desclerotized area ( Figs 565 , 568 ) anteriorly delimited by short transverse (longitudinally grooved, see Figs 565 , 570 , arrows), pale brown strip connecting both blackish brown lateral parts of synsclerite (both yellowish brown in palest specimens); the groove often producing a chordate anterior margin to the membranous area. The ventral part of T7+S7 differing from that of A. equiseti in having desclerotized area with more setae and, particularly, in lacking narrow yellowish brown anterolateral areas reaching above 7th spiracle. Posterior to the ventral desclerotized area there is an additional pale brown, nat and distinctively micropubescent posteromedial appendage ( Figs 564–566, 568, 570 , often at least partly invaginated) shared with A. equiseti . 8th segment ( Figs 566, 568 ), narrow, long, largely membranous, micropubescent. T8 narrow, elongate, posteriorly parallel-sided, with nne exclinate lateral setulae and anterior corners laterally wing-like dilated and separated by a deep narrow incision ( Fig. 567 ); S8 ( Fig. 568 ) much shorter than T8, medially divided to form 2 narrow, nnely setose sclerites with posterior end curved and invaginated (as in relatives). Genital chamber (uterus) with internal sclerotization ( Figs 571, 573 ) formed by 1 pair of long, nat, but rather crooked and complex sclerites with suboblong pronle (cf. Fig. 571 , in contrast to a more regularly oblong outline in A. equiseti ) and 1 anterior annular sclerite that is wider anteriorly and posteriorly, down-curved posteriorly (usually more than in A. equiseti ). Ventral receptacle ( Fig. 571 ) very long, slender, tubuliform, hyaline and multiply twisted, distinctly longer than that of A. equiseti ; accessory gland hyaline, elongate, with duct slightly dilated distally. Spermathecae (1+1) suboval, with long digitiform invagination ( Figs 569, 572 ), indistinctly wrinkled surface and dark spinulae (more than in A. equiseti ) near duct insertion; the latter situated close to opening of invagination; duct ending in very short, pigmented cervix. T10 ( Fig. 567 ) small, dark, transversely oval, with 2–4 pairs of setae including 1 long pair and with sparse micropubescence. S10 ( Fig. 568 ) small, longer than T10, somewhat tapered and rounded anteriorly, wider posteriorly, nnely setulose and micropubescent. Cercus slender, relatively long, with numerous nne setae, apical and dorsopreapical longest, subequal in length ( Figs 566–568 ).

Discussion. Based on a number of shared male and female genitalic characters, A. vulgaris sp. nov. is considered the sister species of A. equiseti sp. nov. Most of the male genitalic features are probably plesiomorphic (e.g. medandrium wider and more robust; gonostylus with simple apex; aedeagal part of folding apparatus without long spines ventrally) but the unique posteromedial ventral appendage of the female T7+S7 (see Figs 564, 565 , 570 , 589, 591 ) is considered a strong synapomorphy of this sister pair.

It is notable that A. vulgaris is variable in some structures of the male genitalia, particularly the form of the gonostylus ( Figs 559 , 563 ), the armature of the caudal process of the transandrium (cf. Figs 553 , 561 ) and the number of spines in the saccus ( Figs 556 , 562 ), which ranges from two to four. This variability may renect the extreme habitat and host-plant tolerance of A. vulgaris (see below) but this assumption should be experimentally tested. Nevertheless, A. vulgaris differs from all Nearctic species of the A. gracilis group in having lighter microtomentum of the thorax and a distinctly elongate head and eye. There are also a number of differences from its closest relative, A. equiseti , not only in the terminalia (epandrium narrower, pregonite more projecting, postgonite more slender, ventral appendage of caudal process with teeth reduced, saccus with fewer spines, female T7+S7 with paler anteroventral strip simple (not extended to 7th spiracle), female T8 with anterolateral wing-like dilations), but also in the form of the head (see above), the colouration of the frons and, particularly, the legs (in A. vulgaris uniformly yellow except for the sometimes shiny brown patch on the base of cx 1 and the 2 brownish terminal tarsal segments).

Etymology. The new species is named vulgaris (= common, a Latin adjective) to renect its frequent occurrence in various communities of graminoid plants.

Biology. The habitat of Anthomyza vulgaris includes a wide range of sites from extremely dry to relatively wet sites with plant communities usually dominated by grasses. In at least Ontario, A. vulgaris is one of the most commonly encountered anthomyzids, particularly in open, disturbed grassy areas.

The driest sites include those collections made from the beach grass Ammophila breviligulata Fernald from three localities, namely Nova Scotia: South Harbour Beach – mixed with Lathyrus sp. ( Fabaceae ) not considered a potential host; Ontario: Pancake Bay P. Pk., Fig. 546 ; and Quebec: Îles de la Madeleine, road 199 between Havre-aux-Maisons and Point-aux Loups. There are also similar records from a very sandy site (Ontario: Bruce Peninsula N. P.–Singing Sands) supporting Calamagrostis and “ Agropyron ” (likely Elymus repens ) and from “dunes” (Nova Scotia: Crescent Beach; Ontario: Manitoulin Island–Sand Bay; Quebec: Godbout). An unusual sandy site was the exposed perimeter of a drained man-made lake (Ontario: ~ 66 km NNW Elliot Lake, Rocky Island Lake) where Scirpus microcarpus grew in isolated clumps ( Fig. 548 ).

Multiple collections made from prairie habitat in Manitoba (Aweme), including tallgrass prairie (Gardenton, Winnipeg) and “ Andropogon-Stipa prairie community” (5 mi SW Shilo), also suggest dry habitats, as does the collection from Andropogon (Wisconsin: Swan Lake– 4 mi E Portage).A particular reference to “tundra at summit” (New Hampshire: Mt. Washington) may be another dry site.

There are several records indicating that roadsides and both dry and wet ditches yield this species. Fens, alvars (e.g. Ontario: Manitoulin Is., nr. Misery Bay [Prov.] Nature Reserve, Little Lake Huron Road, Fig. 549 ), and alvar savanna are also recorded, along with riparian habitats such as “shoreline” and “riverside”, along with “lakeside” and “marshy lake shore”. References such as “vegetation on boggy trail”, “noodplain community near tamarack bog”, “boggy meadow” and “ Sphagnum bog” also indicate that wetter sites can support populations of this species.

Observations at one roadside ditch site supporting all three species of the A. gracilis group (Ontario: ~ 21 km NNE Smooth Rock Falls; discussed under A. furvifrons ) suggest that A. vulgaris was more commonly collected on the drier slopes of the roadside ditch in a mix of grasses and herbs, though dennitive segregation among the three species could not be made. There are a few specimens recorded from Equisetum ssuviatile , but this host plant for A. equiseti is likely not a host for A. vulgaris and the specimens were likely part of a population emerged from nearby grasses. For example, the Marathon, Ontario site yielded large numbers of A. equiseti from mostly emergent E. ssuviatile , and the only specimens of A. vulgaris were taken about 50 meters away on the other side of the highway among roadside graminoids (mostly grasses).

Specinc graminoid plants that are mentioned include grasses ( Calamagrostis , Phalaris , “ Agropyron ” (likely E. repens )), sedges ( Carex , Dulichium , Eleocharis , Scirpus ) and rushes ( Juncus ). Cattails ( Typha ) are not likely to be a host but indicate a wetter site. Collections that do not include grasses tend to be represented by small numbers of specimens. Unusual references include “ex. Ulmus sp.”, “ex. Betula glandulosa ” Michx. , and “ex. leafy spurge ” ( Euphorbia esula L.) but these are not considered likely hosts.A collection from the blossoms of Smilacina sp. suggests a pollen and nectar source for adults (North Carolina: Mollie Gap). Adults of A. vulgaris have been collected from 23 May (Tennessee: SW corner of Cades Cove) to 4 September (Ontario: Hilton Township).

Distribution. Anthomyza vulgaris ranges transcontinentally from British Columbia to Prince Edward Island (Alberta, British Columbia, Labrador, Manitoba, New Brunswick, Nova Scotia, Ontario, Prince Edward Island, Quebec and Saskatchewan) in Canada. However, in the United States of America, this most commonly collected species of the Anthomyza gracilis group is restricted to the east from Minnesota to Maine and Pennsylvania to Tennessee and Georgia (Georgia, Maine, Massachusetts, Michigan, Minnesota, New Hampshire, New York, North Carolina, Pennsylvania, Tennessee, Vermont, West Virginia and Wisconsin) (see Table 2).

Figures

Figs 518–523. Primary types of the Nearctic species of the Anthomyza gracilis group. 518 – A. furvifrons sp. nov., holotype male, laterally, body length 2.30 mm; 519 – same specimen, head dorsally; 520 – A. vulgaris sp. nov., holotype male, laterally, body length 2.50 mm; 521 – same specimen, head dorsally; 522 – A. equiseti sp. nov., holotype male, laterally, body length 2.35 mm; 523 – same specimen, head dorsally. Photo by J. Roháček.

Figs 524–526. Wings of the Nearctic species of the Anthomyza gracilis group. 524 – A. furvifrons sp. nov., paratype male, wing length 2.4 mm (Canada: Ontario); 525 – A.vulgaris sp. nov., paratype male, wing length 2.5 mm (Canada: Ontario); 526 – A. equiseti sp. nov., paratype male, wing length 2.4 mm (Canada: Ontario). Photo by K. N. Barber.

Figs 544–546. Habitats of Nearctic species of the Anthomyza gracilis group and A. vulgaris sp. nov. 544 – riverbank of Mississagi River (Canada: Ontario) with rich graminoid communities, habitat in the type locality of A. furvifrons sp. nov.; 545 – A. vulgaris sp. nov., female on leaf of sedge, body length ca. 2.8 mm (Canada: Ontario: Sault Ste Marie); 546 – sandy beach with growth of Ammophila breviligulata (Canada:Ontario: Pancake Bay), extreme habitat of A. vulgaris. Photo by K. N. Barber (Fig. 544) and J. Roháček (others).

Figs 547–550. Anthomyza vulgaris sp. nov. and some of its habitats. 547 – bubbling female on Carex leaf laterally, body length ca. 2.8 mm (Canada: Ontario: Sault Ste Marie); 548 – pure growth of Scirpus microcarpus on exposed lake bottom (Canada: Ontario: Rocky Island Lake); 549 – graminoid vegetation on alvar near Misery Bay (Canada: Ontario: Manitoulin I.); 550 – anterior half of the same female as in Fig. 547, dorsally. Photo by J. Roháček.

Figs 551–559. Anthomyza vulgaris sp. nov., paratype male (Canada: Ontario). 551 – external genitalia, caudally; 552 – the same, laterally; 553 – hypandrial complex, laterally; 554 – transandrium, caudally; 555 – caudal process of transandrium, ventrally; 556 – aedeagal complex, laterally; 557 – apex of nlum, subventrally; 558 – armature of right wall of aedeagal part of folding apparatus; 559 – gonostylus, ventrolaterocaudally (widest extension). Scales = 0.05 mm (Figs 557, 559) and 0.1 mm (others).

Figs 560–565. Anthomyza vulgaris sp. nov., paratypes, male and female (Canada: Ontario). 560 – male pregonite, lateroventrally, widest extension; 561 – caudal process of transandrium, laterally; 562 – saccus of distiphallus, laterally; 563 – gonostylus, ventrolaterocaudally (widest extension); 564 – female T7+S7, laterally; 565 – the same, ventrally. Scales = 0.1 mm (Figs 560–562), 0.05 mm (Fig. 563) and 0.2 mm (others).

Figs 566–573. Anthomyza vulgaris sp. nov., paratype female (Canada: Ontario). 566 – postabdomen, laterally; 567 – the same, dorsally; 568 – the same, ventrally; 569 – spermatheca; 570 – T7+S7, caudally (end of postabdomen omitted); 571 – female genital chamber and S8, laterally (pleural micropubescence omitted); 572 – spermatheca; 573 – internal sclerites, ventrally. Scales = 0.2 mm (Figs 566–568, 570), 0.05 mm (Figs 569, 572) and 0.1 mm (others). For abbreviations see p. 11.

Figs 589–596.Anthomyza equiseti sp.nov.,paratype female (Canada:Ontario).589 – postabdomen, laterally;590 – the same, dorsally; 591 – the same, ventrally; 592 – female genital chamber and S8, laterally (pleural micropubescence omitted); 593 – spermatheca; 594 – apex of ventral receptacle, ventrally; 595 – spermatheca; 596 – internal sclerites, ventrally. Scales = 0.2 mm (Figs 589–591), 0.05 mm (Figs 593–595) and 0.1 mm (others).