Cnemaspis nigridia ( Smith, 1925 )

Grismer, Lee, Wood, Perry L., Anuar, Shahrul, Riyanto, Awal, Ahmad, Norhayati, Muin, Mohd A., Sumontha, Montri, Grismer, Jesse L., Onn, Chan Kin, Quah, Evan S. H. & Pauwels, Olivier S. A., 2014, Systematics and natural history of Southeast Asian Rock Geckos (genus Cnemaspis Strauch, 1887) with descriptions of eight new species from Malaysia, Thailand, and Indonesia, Zootaxa 3880 (1), pp. 1-147 : 100

publication ID 10.11646/zootaxa.3880.1.1

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Cnemaspis nigridia ( Smith, 1925 )


Cnemaspis nigridia ( Smith, 1925)

Black Rock Gecko

Fig. 51 View FIGURE 51

Heteronota Kendallii Gray, 1845:174 View in CoL (in part)

Gonatodes affinis Shelford, 1901:49

Gonatodes nigridius Smith, 1925:22

Cnemaspis nigridius Brongersma, 1934:165 ; Manthey & Grossmann, 1997:214; Cox, van Dijk, Nabhitabhata & Thirakhupt, 1998:90; Auliya 2006:180

Cnemaspis cf. nigridia Chan-ard, Grossmann, Gumprecht & Schulz, 1999:104

Holotype. BM 1946.8 .22.90. Type locality: “Mt. Gadin” = Gunung Gading , Sarawak, East Malaysia (01°41.16 N, 109°50.35 E) at approximately 100 m in elevation. GoogleMaps

Diagnosis. Maximum SVL 75.5 mm; 10 or 11 supralabials; 9–11 infralabials; ventral scales weakly to moderately keeled; 10–16 contiguous, pore-bearing, precloacal scales with round pores; 39–43 paravertebral tubercles; tubercles semi-linear to randomly arranged, absent from flanks and lateral caudal furrows; ventrolateral caudal tubercles present; lateral row of caudal tubercles present; caudal tubercles not encircling tail; subcaudals keeled but bearing an enlarged median row of smooth scales; 2–4 postcloacal tubercles on each side of tail base; no enlarged femoral or subtibial scales; submetatarsal scales of first toe enlarged; subtibials keeled; 26–29 subdigital fourth toe lamellae; a pair of large, round, black spots in shoulder region; yellow to white bars on flanks; wide, black and dull-yellow caudal bands (Tables 6,7).

Color pattern in life ( Fig. 51 View FIGURE 51 ). Dorsal coloration of head, body and tail dark brown overlain with large, oval, black spots on anterior portion of body and nape; cream- colored, vertebral markings on nape and dorsum; thin, semi-transversely oriented, yellow bands on body and flanks; limbs bearing wide, dark and yellowish alternating faint bands; yellow caudal bands containing dark pigmentation; all ventral surfaces dark-gray.

Distribution. Cnemaspis nigridia is known from the type locality of Gunung Gading as well as Gunung Pueh, Gunung Beremput, and the Bau Limestone Area ( Smith 1925; Naming & Das 2004; Das 2006), Sarawak, East Malaysia ( Fig. 4 View FIGURE 4 ).

Natural History. We have observed Cnemaspis nigridia only at night on the surfaces of large granite boulders in the vicinity of streams within old lowland secondary forest in forest ( Fig. 51 View FIGURE 51 ). Surveys during the day resulted in finding only C. kendallii in these areas. In all the locations from which this species has been reported except the Bau Limestone Area (which may be erroneous, see below) it is restricted to granite boulders.

Remarks. Naming & Das (2004) report Cnemaspis nigridia as occurring on karst formations in the Bau Limestone Area. However, it is not clear if specimens were actually examined as there is no character support for their identification or voucher specimens listed. Only the habitat generalists C. kendallii and C. peninsularis sp. nov. and C. flavigaster occur on both karst and granite and we suspect the Bau population may not be C. nigridia but the karst-adapted C. paripari which occurs in that region ( Grismer & Chan 2009).

Relationships. Cnemaspis nigridia is the sister species of C. paripari from northwestern Borneo ( Fig. 2 View FIGURE 2 ).

Material examined. East Malaysia: Sarawak, Gunung Gading LSUHC 9167–70.


La Sierra University, Herpetological Collection














Cnemaspis nigridia ( Smith, 1925 )

Grismer, Lee, Wood, Perry L., Anuar, Shahrul, Riyanto, Awal, Ahmad, Norhayati, Muin, Mohd A., Sumontha, Montri, Grismer, Jesse L., Onn, Chan Kin, Quah, Evan S. H. & Pauwels, Olivier S. A. 2014

Cnemaspis nigridius

Brongersma, L. D. 1934: 165

Gonatodes nigridius

Smith, M. A. 1925: 22

Gonatodes affinis

Shelford, R. 1901: 49

Heteronota Kendallii Gray, 1845:174

Gray, J. E. 1845: 174
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