Cnemaspis monachorum Grismer, Norhayati, Chan, Belabut, Muin, Wood & Grismer, 2009

Grismer, Lee, Wood, Perry L., Anuar, Shahrul, Riyanto, Awal, Ahmad, Norhayati, Muin, Mohd A., Sumontha, Montri, Grismer, Jesse L., Onn, Chan Kin, Quah, Evan S. H. & Pauwels, Olivier S. A., 2014, Systematics and natural history of Southeast Asian Rock Geckos (genus Cnemaspis Strauch, 1887) with descriptions of eight new species from Malaysia, Thailand, and Indonesia, Zootaxa 3880 (1), pp. 1-147 : 27-34

publication ID 10.11646/zootaxa.3880.1.1

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Cnemaspis monachorum Grismer, Norhayati, Chan, Belabut, Muin, Wood & Grismer, 2009


Cnemaspis monachorum Grismer, Norhayati, Chan, Belabut, Muin, Wood & Grismer, 2009

Monks’ Rock Gecko

Figs. 9 View FIGURE 9 , 10 View FIGURE 10

Holotype. ZRC 2.6774 View Materials . Type locality: “ Wat Wanaram , Pulau Langkawi, Kedah, Peninsular Malaysia (06°20.275 N, 99°52.507 E)” at 35 m in elevation. GoogleMaps

Diagnosis. Maximum SVL 35.1 mm; seven or eight supralabials; 5–7 infralabials; ventral scales smooth; three, contiguous, pore-bearing precloacal scales with round pores; 2–20 paravertebral tubercles; body tubercles randomly arranged, absent from flanks and lateral caudal furrows; no ventrolateral caudal tubercles; lateral row of caudal tubercles present anteriorly; caudal tubercles not encircling tail; subcaudals smooth bearing a medial row of enlarged scales; one or two postcloacal tubercles on each side of tail base; no enlarged femoral, subtibial or submetatarsal scales; subtibials smooth; 24–30 subdigital fourth toe lamellae; gular region, throat, and abdomen in males yellow; faint, dark, lineate, mid-gular marking present (Tables 6,7).

Color pattern in life (Figs. 9,10). Dorsal ground color brown; head, body, and limbs overlain with small, irregularly shaped, somewhat randomly arranged, black and cream colored spots varying somewhat in size; light dorsal blotches in general paravertebral arrangement; markings on top of head smaller than those on body; dark markings in caudal region tend to form bands that encircle tail; anterior gular region yellowish, most pronounced on mental scale; faint, dark, midgular stripe present; pectoral region and anterior, abdominal region orangish yellow; remainder of ventral surfaces of body and limbs beige bearing faint, dark, stippling.

Distribution. Cnemaspis monachorum is known only from the Langkawi Archipelago where it inhabits the islands of Langkawi (Grismer et al. 2009) and reported here for the first time from the smaller satellite island of Langgun ( Fig. 4 View FIGURE 4 ).

Natural history. On Pulau Langkawi, Grismer (2011a) noted that Cnemaspis monachorum is a lowland, saxicolous species known only from the karst outcropping of Wat Wanaram near the town of Kuah in a region dominated by a mixture of primary coastal and lowland dipterocarp forest. Cnemaspis monachorum is a small, swift, agile species abroad only during the day, climbing on the fragmented boulders along the periphery of karst formations as well as along the base of karst cliffsides near the edges of cracks. Lizards are wary and will rapidly retreat into a crack while curling their tail above their back and waving it from side to side. During the night, lizards are not abroad and only occasionally observed deep within limestone cracks. Cnemaspis monachorum is the smallest species of Cnemaspis and possibly associated with this small size is that gravid females carry only a single egg whereas females of all other Cnemaspis carry two eggs. Gravid females have been observed in October. On Pulau Langgun, C. monachorum is also diurnal and common throughout the island on the limestone rocky hillsides as well as the cliff faces that edge Tasik (lake) Langgun ( Fig. 10 View FIGURE 10 ). During August, no gravid females were found on Pulau Langgun suggesting that October may be the beginning of the reproductive season.

Relationships. Cnemaspis monachorum is the basal lineage of the Pattani clade ( Fig. 2 View FIGURE 2 ).

Material examined. Malaysia: Kedah; Pulau Langkawi LSUHC 9118 View Materials , ZRC 2.6774 View Materials 76 View Materials (type series). Material examined since Grismer et al. (2009) : Malaysia: Kedah; Pulau Langkawi LSUHC 9115–17 View Materials ; Pulau Langgun LSUHC 10807–11 View Materials .













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