Pacuvia castanea Curtis, 1844

Smith, Andrew B. T. & Cherman, Mariana, 2021, Review of the genus Pacuvia Curtis, 1844 (Coleoptera: Scarabaeidae: Melolonthinae: Diplotaxini), Revista Chilena de Entomología (Rev. Chil. Entomol.) 47 (4), pp. 955-963 : 958-960

Pacuvia castanea Curtis, 1844

Pacuvia castanea Curtis 1844: 199 . Type locality: “Valparaiso.” Type series: lectotype male at BMNH labeled 1) “Type” (typeset round label with red border), “Valparaiso / 63.49” (handwritten), “ Pacuvia / castanea, Curt. ” (handwritten), “86 Pacuvia / castanea / Curt” (handwritten), “ PACUVIA / CASTANEA / CURTIS ♂ / LECTOTYPE / A.B.T. SMITH” (handwritten and typeset red label), “Southern Neotropical Scarabs / database # AS 2615667 / Pacuvia castanea / Curtis, 1844 ♂ / DET: A.B.T.SMITH 2008” (typeset). Lectotype here designated.

Synonym. Liogenys gayana Solier 1851: 100 . Type locality: “ Santiago, Santa Rosa, Illapel, etc.” Type series: lectotype male at MNHN labeled “ L. Gayanus / Cat. Mus. / Chili. / M. Gay.” (handwritten green label), “15 / 43” (handwritten on underside, green on upperside), “MUSEUM PARIS / Chili / Gay” (typeset and handwritten), “ LIOGENYS GAYANA / SOLIER ♂ / LECTOTYPE / A.B.T. SMITH” (handwritten and typeset red label), “Southern Neotropical Scarabs / database # AS 2614768 / Pacuvia castanea / Curtis, 1844 ♂ / DET: A.B.T.SMITH 2008” (typeset). Lectotype here designated.

Paralectotype male at MNHN labeled “15 / 43” (handwritten on underside, green on upperside), “MUSEUM PARIS / CHILI / GAY 1843” (typeset), “ Liogenys / gayanus / DET. / L. W. SAYLOR” (handwritten and typeset), “ LIOGENYS GAYANA / SOLIER ♂ / PARALECTOTYPE / A.B.T. SMITH” (handwritten and typeset yellow label), “ Southern Neotropical Scarabs / database # AS 2614769 / Pacuvia castanea / Curtis, 1844 ♂ / DET: A.B.T.SMITH 2008” (typeset) .

Material examined. Chile: Coquimbo: Agua Dulce , (31°33’39.32’’S, 71°32’57.34’’W), 22 May 1986, A. Henry — 1 male ( UMCE) ; Canela Baja , (31°24’2’’S, 71°27’16’’W), 13–24 October 1961, L.E. Peña — 1 male ( FMNH) ; Illapel-Canelillo , 20 May 2003, D. Jackson — 5 males ( UMCE) ; Los Vilos , (31°54’42’’S, 71°30’47’’W), 1–31 May 1984, L.E. Peña — 2 males, 1 female ( CMNC) , 2 males, 1 female ( LEMQ) ; N Pichidangui , (32°8’.89’’S, 71°32’.38’’W), 6–12 August 1960 — 1 male ( FMNH) . Valparaíso: Costa Pucalán , (32°45’16.68’’S, 71°20’50.98’’W), 5 October 1967, L.E. Peña — 1 female ( CMNC) ; El Sauce ( 15 km E San Antonio ), (33°36’24.84’’S, 71°28’13.9’’W), 1–31 May 1946, L. Peña Guzman — 9 males ( FMNH) ; ValparaÍso , (33°2’43.83’’S, 71°37’12.78’’W), 1 male ( MNHN) ; ValparaÍso , (33°2’43.83’’S, 71°37’12.78’’W), C. Darwin — 1 male ( BMNH) ; ValparaÍso , (33°2’43.83’’S, 71°37’12.78’’W), 1–31 October 1926 — 1 male ( UMCE) ; Villa Alemana , (33°2’51.08’’S, 71°22’22.89’’W), 8 August 1970, J. Solervicens — 1 male ( UMCE) . Metropolitana: 20 km W Rungue , (33°3’43.05’’S, 70°59’46.98’’W), 26 November 1967, L. O’Brien, C.W. O’Brien — 1 female ( EMEC) ; Apoquindo , (33°24’S, 70°31’59’’W), 1–31 December 1979, L.E. Peña — 2 males ( CMNC) , 1 male ( FMNH) , 1 male ( LEMQ) ; Caleu , (32°59’52.69’’S, 70°58’38.68’’W), 17 September 1986, M. Cerda — 1 male ( UMCE) ; Cerro Calera , (33°39’50.4’’S, 70°48’18’’W), 25 June 2005, A. Vera — 17 males, 1 female ( MNNC) ; Colina , (33°12’S, 70°40’59’’W), 10 July 1979, L.E. Peña — 2 males ( CMNC) , 2 males ( LEMQ) ; Colina , (33°12’S, 70°40’60’’W), 10 August 1979, L.E. Peña — 27 males ( LEMQ) ; Colina, El Portezuelo , (33°12’0.62’’S, 70°40’10.16’’W), 31 July 1979 — 15 males, 1 female ( FMNH) ; Colina, El Portezuelo , (33°12’0.62’’S, 70°40’10.16’’W), 12 July 1985, L.E. Peña — 1 male, 1 female ( CMNC) , 33 males ( LEMQ) ; El Arrayán , (33°20’33.06’’S, 70°28’18.64’’W), 1–30 September 1979, L.E. Peña — 6 males ( LEMQ) ; El Canelo , (33°34’59’’S, 70°27’0’’W), 1–31 October 1951 — 1 male ( CMNC) , 1 male ( FMNH) ; El Canelo , (33°34’59’’S, 70°27’0’’W), 1–31 October 1952 — 5 males ( UMRM) ; El Canelo , (33°34’59’’S, 70°27’0’’W), 1–30 November 1952, L.E. Peña — 2 males ( FMNH) ; El Canelo , (33°34’59’’S, 70°27’0’’W), 1 October–31 December 1978, L.E. Peña — 4 males ( CMNC) , 29 males ( CMNH) ; El Manzano , (33°35’S, 70°24’W), 10 October 1993, R.D. Ward — 1 male ( CMNH) ; El Manzano, (33°34’59.99’’S, 70°24’0’’W), 1–31 October 2003, Barrera Medina — 1 male ( PKLC) ; El Portezuelo , (33°12’S, 70°40’12’’W), 1–31 August 1979, L.E. Peña — 1 male ( CMNC) , 3 male ( LEMQ) ; El Roble , 1900 m (32°58’33.6’’S, 71°0’36’’W), 1–30 November 1992, L.E. Peña — 4 male ( CMNC) ; Lampa , (33°17’S, 70°54’0’’W), Zapata — 2 male ( UMCE) ; Las Condes , (33°22’S, 70°31’0’’W), 4 May 2004, M. Guerrero — 5 male ( MNNC) , 1 male, 1 female ( CMNC) ; Las Condes , (33°22’S, 70°31’0’’W), 12 May 2005, M. Guerrero — 2 males ( MNNC) ; Lo Curro , (33°22’S, 70°34’59’’W), 17–18 August 1968, L. Alfaro — 1 female ( CMNC) ; Maipu , (33°30’40.96’’S, 70°48’6.24’’W), 1–28 February 1906, F.V. Ibarro — 1 male ( USNM) ; Peñalolen, Camino Las Pircas , (33°29’55.87’’S, 70°31’44.69’’W), 31 July 2015, V.M. Diéguez — 1 male ( VMDM) ; RÍo Clarillo , (33°39’S, 70°37’60’’W), 20 August–3 October 1991, J. Solervicans — 1 female ( UMCE) ; Santiago , (33°27’S, 70°40’0’’W) — 1 male ( BMNH) , 6 males ( CASC) ; Santiago , (33°27’S, 70°40’0’’W), 1 October–31 December 1978, L.E. Peña — 3 males ( CMNH) . O’Higgins: Los Arrayanes , NW Rangacua, 1500 m (34°3’37.86’’S, 70°52’46.78’’W), 1–30 November 1981, M. Marin — 1 male ( FMNH) . Maule: Linares , (35°51’S, 71°36’0’’W), 1–31 December 1946 — 5 male ( UMRM) ; Los Cipreses , (35°3’S, 70°36’0’’W), 14 January 1968, L.E. Peña — 1 male ( FMNH) . Ñuble: Cordillera Ñuble , 1–28 February 1983, L.E. Peña — 1 male ( CMNC) , 8 males ( LEMQ) ; Biobío: Concepción , (36°49’60’’S, 73°3’0’’W), 15 December 1962, O. Fetes — 2 males ( CNCI) ; Contulmo , (38°0’36’’S, 73°13’48’’W), 9 December 1967, T. Cekalovic — 5 females, 1 male ( CNCI) ; La Laja , (37°16’S, 72°42’0’’W), 25 January 1987 — 3 females ( UNSM) ; Penco , (36°43’60’’S, 72°58’60’’W), 1–31 May 1967, Fierro — 2 males, 2 females ( CNCI) ; no data — 1 male ( BMNH) . No data: 8 males, 1 female ( BMNH) , 2 males ( MNHN) , 1 male ( MNNC) ; Reed — 3 females ( BMNH) .

Diagnosis. Length 8.5–12.5 mm (males), 8.5–12.0 mm (females); width 4.0– 6.5 mm (males), 4.5–6.5 mm (females). Antennae with 9–10 antennomeres, male and female antennal club distinctly shorter than the pedicel. Pronotum with small, dense punctures ( Figs. 1–2 ). Males and females not brachypterous and fully extended wing longer than elytra.

Distribution. Chile: Coquimbo, ValparaÍso, Metropolitana de Santiago, O’Higgins, Maule, Ñuble, BiobÍo ( Fig. 17 ).

Comments. Gutiérrez (1951) stated that he only observed two specimens out of 220 that had 10 antennomeres and characterized them as aberrant. We observed a far higher percentage of specimens we examined with 10 antennomeres, so clearly the number of antennomeres in this species is variable and an unreliable diagnostic character.

Figures

Figure 17. Distributional map for Pacuvia castanea (black circles) andPacuvia philippiana (red triangles) in central Chile.

Figures 1–5.Pacuvia castanea Curtis,1844.1. Dorsal habitus.2. Obliquehabitus.3. Head andpronotum. 4. Parameres in dorsal view. 5. Parameres and phallobase in lateral view.