Amplirhagada combeana Iredale, 1938

Köhler, Frank, 2010, Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae), Records of the Australian Museum 62 (2), pp. 217-284 : 227-228

publication ID	https://doi.org/ 10.3853/j.0067-1975.62.2010.1554
publication LSID	lsid:zoobank.org:pub:749A271A-6942-4D4C-B0DF-B968462BFF0C
persistent identifier	https://treatment.plazi.org/id/03F987C5-FF9B-F757-01FB-10DE4831C9AD
treatment provided by	Felipe
scientific name	Amplirhagada combeana Iredale, 1938
status

Treatment

Amplirhagada combeana Iredale, 1938 View in CoL

Helix (Hadra) imitata var. cassiniensis Smith, 1894: 92 View in CoL , pl. 7, fig. 16 (Cassini Island). Amplirhagada combeana Iredale, 1938: 113 View in CoL (nomen novum), Solem, 1981a: 310–312, pl. 12b, figs. 71h–i).

Nomenclatural remarks. The original specific epithet “ cassiniensis ” employed by Smith (1894) is preoccupied and has been replaced with “ combeana View in CoL ” by Iredale (1938), who elevated the taxon to the rank of an independent species. Solem (1981a: 310–312) described details of the shell but was not able to describe the anatomy because no preserved material was available.

Material examined. WAM S41450 View Materials , AMS C463725 (Western Australia, northwestern Kimberley, Bonaparte Archipelago, Cassini Island , 13°57'04"S 125°38'39") , WAM S41455 View Materials , AMS C463726 (13°57'22"S 125°37'53"E) ( Fig. 1 View Figure 1 ) GoogleMaps .

Sealing strategy. Rock sealer.

Shell ( Fig. 14A–C View Figure 14 , Pl. 1.7). Broadly conical with low to medium spire; solid (not translucent). Periphery well rounded to slightly angulate. Umbilicus completely concealed by columellar reflection or forming a chink. Background colour horn to yellowish brown; with diffuse to well defined, thin to moderately broad, chestnut brown sub-sutural and mid-whorl bands that are most conspicuous on last whorl; ventral colour horn; outer lip colour same as shell; inner lip translucent. Protoconch c. 1.7 mm in diameter with 1.5 whorls, comparatively strong axial sculpture. Teleoconch sculptured by coarse, curved, in cross-section rounded, irregularly spaced lirae; spaces equal to thickness of lirae; lirae evenly distributed across shell and whorl diameter, height reduced underneath suture. Angle of aperture 30°; outer lip rounded, moderately thick, expanded, slightly reflected; basal node of lip weak, palatal node absent.

Pallial morphology. Pallial cavity moderately deep, extending 3 ⁄ 4 whorl. Pigmentation on mantle consists of sparse greyish patches or spots. Kidney extending not more than half of pallial cavity.

Genital morphology ( Figs. 12–13 View Figure 12 View Figure 13 ). Penis straight; more or less of same length as anterior part of oviduct or longer. Vas deferens forms simple loop before entering penis. Penial retractor muscle shorter than penis. Penial verge short (<1 ⁄ 8 penial chamber), broad with pointed to rounded tip. Penial wall covered by a number of smooth longitudinal pilasters; pustulation absent. Main stimulatory pilaster welldifferentiated, extending apical to median portion of penial chamber, cone-shaped, corrugated by ridges that support little hooks. Vas deferens entering penial sheath in upper third. Vagina of medium thickness, posteriorly inflated. Inner vaginal wall with smooth, longitudinal pilasters. Spermathecal duct moderately thick, inside with smooth longitudinal pilasters. Spermathecal head globular, connected with oviduct by connective tissue, inside entirely smooth with delicate wall. Free oviduct comprising about half of anterior part of oviduct; coiled underneath entrance to spermoviduct. Spermoviduct longer than anterior part of oviduct. Talon embedded in albumen gland at junction with spermoviduct.

Radular morphology ( Fig. 14D–E View Figure 14 ). Rectangular. Tooth formula C+12+2–3+18–20 with 162.5±3.5 rows of teeth, 33.5±1.9 rows per mm (n = 2). Central teeth with bluntly pointed, triangular to ovate mesocones, shorter than base of tooth; ectocones well developed. Lateral teeth with bluntly pointed, triangular to ovate mesocones; length equal to base of tooth; ectocones well developed, endocones vestigial. Marginal multicuspic; ectocones shorter and narrower than mesocones; endocones approximately same size as ectocones; divided into two denticles.

Comparative remarks. This species differs from all previous species by the absence of penial wall pustules and the presence of a well-developed main stimulatory pilaster.Axial sculpture is not as regular as in A. solemiana and A. indistincta but more pronounced as in A. euroa . With respect to penial morphology A. solemiana and A. indistincta differ most markedly by absence of a well-differentiated main stimulatory pilaster. For comparison with A. mckenziei and A. montesquieuana see below, with other species see Solem (1981a: 310).