Pseudocypus Mulsant & Rey, 1876

He, Liang & Zhou, Hong-Zhang, 2017, Taxonomy of the subgenus Pseudocypus Mulsant & Rey (Coleoptera: Staphylinidae: Staphylinini) and eight new species from China, Zootaxa 4339 (1), pp. 1-77 : 5-9

Key to species of the subgenus Pseudocypus Mulsant & Rey, 1876 from China

(modified from Smetana, 2007, 2009a, 2009b)

1. Head not wider than pronotal disc. Eyes usually about as long as or slightly shorter than tempora in dorsal view. Apical portion

of aedeagus with complex orifice; no sensory peg setae on underside of paramere................................... 2 - Head about as wide as or wider than pronotal disc. Eyes small, distinctly shorter than tempora in dorsal view. Texture of pronotal disc and elytra almost the same (e.g. Fig. 1-1 A ). Apical portion of aedeagus simple, without complex orifice; underside of paramere usually with sensory peg setae............................................... 10 ( O. semenowi lineage )

2. Texture of pronotal disc and elytra distinctly different (e.g. Figs. 9-1 A, 21-1 A), elytra usually distinctly dull............. 3

- Texture of pronotal disc and elytra almost the same (e.g. Fig. 12-1 A)........................... 4 ( O. lewisius lineage )

3. Pronotum very finely and densely punctate, with irregular longitudinal row of coarser punctures along each side of impunctate midline. Aedeagus as in Figs. 21-2 B–D. Length 11.0– 20.3 mm ...... O. picipennis ( Fabricius, 1793) ( O. picipennis lineage )

- Pronotum coarsely and moderately densely to sparsely, often irregularly punctate, without rows of coarser punctures along each side of impunctate midline.........................................................6 ( O. fuscatus lineage )

4. Tergite 2 (anterior to first visible tergite (visible when abdomen fully stretched or elytra opened) entirely punctate and pubescent. Abdominal tergites 6 and 7 (fourth and fifth visible) almost always with a patch of yellowish tomentose pubescence at middle. Aedeagus of characteristic shape ( Smetana, 2009a: figs.19–21). Length 14.0–18.0 mm............................................................................................. O. (Pseudocypus) jelineki Smetana, 2009

- Tergite 2 (anterior to first visible tergite) with at least basal portion lacking punctation and pubescence. Abdominal tergites 6 and 7 (fourth and fifth visible) without a patch of yellowish tomentose pubescence.................................. 5

5. Tergite 2 (anterior to first visible tergite) punctate and pubescent only along apical margin; tergite 7 (fifth visible) with pale apical seam of palisade setae. Apical portion of median lobe of aedeagus in ventral view ladle-like, with apex extremely elongate on the left (Fig. 12-1 C); paramere of aedeagus extremely narrow, dagger-like (Fig. 12-1 D). Length 16.0– 25.1 mm .................................................................. O. (Pseudocypus) densissimus ( Bernhauer, 1933)

- Tergite 2 (anterior to first visible tergite) with apical third to half punctate and pubescent; tergite 7 (fifth visible) without pale apical seam of palisade setae. Apical portion of median lobe of aedeagus of different shape ( Smetana, 2009a: fig. 7); paramere not dagger-like ( Smetana, 2009a: figs. 8, 9). Length 17.0–19.0 mm................ O. (Pseudocypus) lewisius Sharp, 1874

6. Abdominal tergite 2 (in front of first visible tergite) entirely punctate and pubescent ( Smetana, 2009b: fig. 62). Fifth visible tergite without pale apical seam of palisade setae ( Smetana, 2009b: fig. 65)........................................ 7

- Abdominal tergite 2 (in front of first visible tergite) to variable extent lacking punctures and pubescence ( Smetana, 2009b: figs. 63, 64). Fifth visible tergite with pale apical seam of palisade setae.............................................. 8

7. Right lateral portion of median lobe of aedeagus in ventral view evenly arcuate (Fig. 9-2 F). Pronotum voluminous, punctation of pronotum dense and very fine, particularly along posterior and lateral margins, markedly finer and denser than that of disc of head. Aedeagus as in Figs. 9-1 B, C, 9-2 B–D. Length 12.0–15.0 mm....... O. (Pseudocypus) aenescens Eppelsheim, 1889

- Right lateral portion of median lobe of aedeagus in ventral view conspicuously dilated (Fig. 16-2 F). Pronotum not voluminous, punctation of pronotum no more than slightly finer and denser than that of head. Aedeagus as in Figs. 16-1 B, C, 16-2 B–D. Length 11.0–14.0 mm..................................... O. (Pseudocypus) inexspectatus Eppelsheim, 1887

8. Punctation of pronotum moderately dense, irregular, leaving variably large areas impunctate; or punctation variably dense, with one impunctate area on each side of impunctate midline. Apical portion of median lobe of aedeagus in ventral view narrow, with subacute apex................................................................................ 9

- Punctation of pronotum dense and more or less regular, no impunctate areas present. Apical portion of median lobe of aedeagus in ventral view wide, with obliquely subtruncate apex (Fig. 14-2 F). Aedeagus as in Figs. 14-1 B, C, 14-2 B–D. Length 12.0–15.0 mm..................................................... O. (Pseudocypus) graeseri Eppelsheim, 1887

9. Right lateral portion of median lobe of aedeagus in ventral view gradually dilated ( Smetana, 2009b: fig. 45). Area behind anterior angles of pronotum densely punctate in male, very sparingly punctate or almost impunctate in female. Aedeagus as in Smetana (2009b: figs. 45–48). Length 11.0–15.0 mm.................... O. (Pseudocypus) fuscatus ( Gravenhorst, 1802)

- Right lateral portion of median lobe of aedeagus in ventral view abruptly dilated ( Smetana, 2009b: fig. 52). Area behind anterior angles of pronotum in both sexes sparingly punctate on ground with distinct microsculpture of longitudinal striae. Aedeagus as in Smetana (2009b: figs. 52–56). Length 12.0–15.0 mm................ O. (Pseudocypus) nigroaeneus Sharp, 1889

10. Pronotal hypomeron microsetose, number of microsetae varying from a few to quite numerous ( Smetana, 2007: figs. 213–215)................................................................................................... 11

- Pronotal hypomeron glabrous, lacking microsetae........................................................... 29

11. Legs uniformly rufobrunneous.......................................................................... 12

- Legs not uniformly rufobrunneous, at least partially darkened, or entirely dark, piceous to black...................... 16

12. Visible abdominal tergites 4 and 5 with patch of not dense, golden-yellowish tomentose pubescence covering almost entire tergite. Aedeagus as in Figs. 18-1 B, C, 18-2 B–D. Length 11.0– 17.5 mm .... O. (Pseudocypus) neocles Smetana, 2007 (pars)

- No patches of yellowish tomentose pubescence on abdominal tergites. Aedeagus different........................... 13

13. Male antenna short, segment 4 about as long as wide, outer segments wider than long (Fig. 24-1 A). Underside of paramere without sensory peg setae (Fig. 24-1 D). Aedeagus as in Figs. 24-1 B, C, 24-2 B–D. Length 11.0– 16.7 mm ............................................................................. O. (Pseudocypus) semenowi Reitter, 1887 (pars)

- Male antenna relatively long, segment 4 distinctly longer than wide (ratios 1.27–1.40), outer segments as long as wide as or longer than wide. Underside of paramere with sensory peg setae. Aedeagus different................................ 14

14. Apical portion of paramere narrow, with subacute to acute apex ( Smetana, 2007: figs. 168, 169). Aedeagus as in Smetana (2007: figs. 167–169). Length 14.0–18.0 mm................................ O. (Pseudocypus) glabrio Smetana, 2007

- Apical portion of paramere wide, with apex arcuate to subtruncate.............................................. 15

15. Apex of median lobe narrowly arcuate (Fig. 10-2 F). Aedeagus as in Figs. 167–169. Apical portion of paramere markedly asymmetrical, knife-like (Figs. 10-1 D, 10-2 G). Length 16.0– 21.7 mm ............ O. (Pseudocypus) ballio Smetana, 2007

- Apex of median lobe subtruncate ( Smetana, 2007: fig. 174). Apical portion of paramere slightly asymmetrical ( Smetana, 2007:

figs. 173, 175). Length 13.0–15.0 mm.................................... O. (Pseudocypus) teuthras Smetana, 2007 16. Visible abdominal tergites 4 and 5 each with patch of yellowish tomentose pubescence at middle..................... 17

- No patches of yellowish tomentose pubescence on abdominal tergites........................................... 20

17. Apex of both median lobe and paramere very wide, widely arcuate; paramere large and wide, covering most of median lobe, with fan-shaped apex (Figs. 20-1 C, D, 20-2 B, F). Length 16.0–17.0 mm.......... O. (Pseudocypus) pelias Smetana, 2007

- Apex of both median lobe and paramere moderately wide, variably arcuate, obtusely rounded, subacute or subtruncate, that of median lobe sometimes emarginate in middle; paramere moderately large and wide, never covering most of median lobe.. 18

18. Apex of median lobe narrowly emarginate in middle ( Smetana: 2007, figs. 111, 112). Aedeagus as in Smetana (2007: figs. 111–114). Small species. Length 10.0– 17.1 mm .............................. O. (Pseudocypus) rhinton Smetana, 2007

- Apex of median lobe not emarginate in middle. Aedeagus different............................................. 19

19. Apex of paramere almost reaching apex of median lobe (Figs. 18-1 C, 18-2 C); apex of median lobe obtuse to subtruncate (Fig. 18-2 F). Male sternite 9 subtruncate apically (Fig. 18-2 H). Golden-yellowish tomentose pubescence on visible abdominal tergites 4 and 5 not dense, covering almost entire tergite. Length 11.0– 17.5 mm .................................................................................................... O. (Pseudocypus) neocles Smetana, 2007 (pars)

- Apex of paramere distinctly not reaching apex of median lobe (Figs. 22-1 C, 22-2 C); apex of median lobe narrowly arcuate to subacute (Fig. 22-2 F). Male sternite 9 emarginate apically (Fig. 22-2 H). Golden-yellowish tomentose pubescence on visible abdominal tergites 4 and 5 dense, limited to middle of tergite. Length 12.0– 19.5 mm. O. (Pseudocypus) puer (Smetana, 2005)

20. Paramere markedly elongate, shaped variably (e.g. Smetana, 2007: fig. 129, 136, 142, 153, 157)...................... 21

- Paramere moderately elongate, shaped variably (e.g. Smetana, 2007: figs. 117, 184, 190)............................ 27

21. Paramere with subacute apex markedly exceeding apex of median lobe ( Smetana, 2007: fig. 157). Underside of paramere without sensory peg setae ( Smetana, 2007: fig. 158). Length 15.0 mm................ O. (Pseudocypus) abaris Smetana, 2007

- Paramere with apex of different shape, no more than moderately exceeding apex of median lobe. Underside of paramere with sensory peg setae..................................................................................... 22

22. Apical portion of paramere conspicuously asymmetrical, shaped as in Smetana (2007: figs. 129, 131, 132). Length 15.0–18.0 mm.................................................................... O. (Pseudocypus) itys Smetana, 2007

- Apical portion of paramere of different shape.............................................................. 23

23. Apical portion of median lobe wider, with obtusely arcuate apex............................................... 24

- Apical portion of median lobe narrow, with subacute apex.................................................... 26

24. Apical portion of paramere markedly curved mediad, sensory peg setae on underside of paramere extended markedly posteriad along lateral margins.................................................................................. 25

- Apical portion of paramere straight, not appreciably curved mediad ( Smetana, 2007: fig. 153), sensory peg setae on underside of paramere forming two short lateral groups connected anteriorly ( Smetana, 2007: fig. 154). Length 16.0 mm................................................................................. O. (Pseudocypus) gorgias Smetana, 2007

25. Apical portion of paramere narrow, conspicuously curved mediad, apex of paramere not reaching apex of median lobe ( Smetana, 2007: fig. 142); right margin of paramere (paramere removed from median lobe and viewed underside up) appearing obtusely angulate ( Smetana, 2007: figs. 144, 145). Length 15.0–18.0 mm.......... O. (Pseudocypus) xerxes Smetana, 2007

- Apical portion of paramere moderately wide, less conspicuously curved mediad, apex of paramere about reaching apex of median lobe ( Smetana, 2007: fig. 136); right margin of paramere (paramere removed from median lobe and viewed underside up) appearing almost evenly arcuate ( Smetana, 2007: figs. 137, 138). Length 16.0–18.0 mm............................................................................................... O. (Pseudocypus) elpenor Smetana, 2007

26. Apical portion of paramere narrow, with apex irregularly, narrowly arcuate ( Smetana, 2007: figs. 195, 197, 198). Length 18.3- 23.2 mm ............................................................. O. (Pseudocypus) orodes Smetana, 2007

- Apical portion of paramere wide, with apex widely rounded ( Smetana, 2007: figs. 149, 150). Length 16.0 mm.............................................................................. O. (Pseudocypus) pammenes Smetana, 2007

27. Aedeagus with wide median lobe and narrow, leaf-like paramere; apex of median lobe markedly exceeding apex of paramere ( Smetana, 2007: fig. 190). Aedeagus as in Smetana (2007: figs. 190, 191). Length 19.0–20.0 mm............................................................................................. O. (Pseudocypus) bion Smetana, 2007

- Apex of paramere about reaching apex of median lobe. Aedeagus different....................................... 28

28. Aedeagus moderately large. Apex of median lobe subtruncate with slight protuberance near apex of paramere ( Smetana, 2007: fig. 118). Aedeagus moderately large, shaped as in Smetana (2007: figs. 117–120). Sternite 9 of male genital segment arcuate apically ( Smetana, 2007: fig. 115). Smaller species. Length 16.0– 21.5 mm ......... O. (Pseudocypus) dolon Smetana, 2007

- Apex of median lobe obtuse, with characteristic formation on face adjacent to paramere on left side below apex ( Smetana, 2007: fig. 185). Aedoeagus large, as in Smetana (2007: figs. 184–187). Sternite of male genital segment emarginate apically ( Smetana, 2007: fig. 182). Larger species. Length 21.0–23.0 mm............. O. (Pseudocypus) palamedes Smetana, 2007

29. Legs uniformly rufotestaceous to rufobrunneous............................................................ 30

- Legs not uniformly rufotestaceous to rufobrunneous, at least partially darkened, or entirely dark, piceous to black........ 42

30. Visible abdominal tergites 4 and 5 each with patch of yellowish tomentose pubescence (mostly not conspicuous). Aedeagus as in Smetana (2007: figs. 3–7). Length 13.0–16.0 mm............................ O. (Pseudocypus) zetes Smetana, 2007

- No patches of yellowish tomentose pubescence on abdominal tergites........................................... 31

31. Aedeagus heavily pigmented, appearing black, shaped as in Smetana (2007: figs. 59, 60). Surface of head and pronotum with slight purplish tint. Length 13.0–16.0 mm................................. O. (Pseudocypus) scaevola Smetana, 2007

- Aedeagus not heavily pigmented, appearing brownish. Surface of head and pronotum without appreciable purplish tint.... 32

32. Male antenna short, segment 4 about as long as wide, outer segments wider than long (Fig. 24-1 A). Aedeagus as in Figs. 24-1 B, C, 24-2 B–D. Length 11.0– 16.7 mm .............................. O. (Pseudocypus) semenowi Reitter, 1887 (pars)

- Male antenna relatively long, segment 4 distinctly longer than wide (ratios 1.15–1.30).............................. 33

33. Underside of paramere without sensory peg setae........................................................... 34

- Underside of paramere with sensory peg setae.............................................................. 35

34. Interspaces between punctures on head and pronotum without microsculpture. Aedeagus short and simple, with paramere distinctly shorter than median lobe ( Smetana, 2007: figs. 77–79). Length 13.0–16.0 mm.. O. (Pseudocypus) hyas Smetana, 2007

- Interspaces between punctures on head and pronotum with microsculpture. Aedeagus elongate, with long paramere almost reaching apex of median lobe ( Smetana, 2007: figs. 17–22). Length 15.0–16.0 mm O. (Pseudocypus) sarpedon Smetana, 2007

35. Apex of median lobe wide, widely arcuate ( Smetana, 2007: fig. 12). Aedeagus as in Smetana (2007: figs. 11–14). Length 12.0–16.0 mm........................................................ O. (Pseudocypus) laelaps Smetana, 2007

- Apex of median lobe narrow, subacute to narrowly arcuate.................................................... 36

36. Sternite 9 with narrow basal portion membranous, located almost centrally....................................... 37

- Sternite 9 with narrow basal portion not membranous, located on left side of basal margin (ventral view)............... 38

37. Sternite 8 with deep medioapical emargination (Fig. 7-2 J). Aedeagus distinctly bent in middle portion in lateral view, shaped as in Figs. 7-1 B, C, 7-2, B–D. Length 17.2–19.5 mm .......................... O. (Pseudocypus) nigriculminis sp. nov.

- Sternite 8 with shallow medioapical emargination (Fig. 8-2 J). Aedeagus not distinctly bended in middle portion in lateral view, shaped as in Figs. 8-1, B, C, 8-2, B–D. Length 18.2–22.1 mm ............... O. (Pseudocypus) recticulminis sp. nov.

38. Paramere almost symmetrical; sensory peg setae on underside of paramere arranged along lateral sides, also almost symmetrical (Figs. 6-1 D, 6-2 A). Aedeagus as in Figs. 6-1 B, C, 6-2, B-D. Length 14.7–19.3 mm .................................................................................................. O. (Pseudocypus) fusciculminis sp. nov.

- Aedeagus distinctly different............................................................................ 39

39. Sensory peg setae on underside of paramere not numerous (about 10), forming a small apical field not extended posteriad along lateral margins of paramere (Figs. 17-1 D, 17-2 A). Aedeagus as in Figs. 17-1 B, C, 17-2 B–D. Length 15.0– 21.5 mm .................................................................... O. (Pseudocypus) menander Smetana, 2007

- Sensory peg setae on underside of paramere numerous to very numerous, in general forming two lateral groups connected anteriorly, or an extensive apical field extended posteriad along lateral margins of paramere.......................... 40

40. Aedeagus elongate, paramere long and narrow, with conspicuous longitudinal crest on face away from median lobe, apex of paramere about reaching apex of median lobe ( Smetana, 2007: fig. 34). Sternite 9 and tergite 10 of male genital segment elongate ( Smetana, 2007: figs. 32, 33). Length 15.0–17.0 mm.................. O. (Pseudocypus) quiris Smetana, 2007 (pars)

- Aedeagus less elongate, paramere moderately long and wide, without conspicuous longitudinal crest on face away from median lobe, apex of paramere not reaching apex of median lobe. Sternite 9 and tergite 10 of male genital segment not elongate................................................................................................... 41

41. Apex of paramere situated considerably below apex of median lobe; sensory peg setae on underside of paramere forming two lateral groups connected anteriorly ( Smetana, 2007: figs. 40, 43). Aedeagus as in Smetana (2007: figs. 40–43). Length 13.0–17.0 mm................................................ O. (Pseudocypus) sericeomicans ( Bernhauer, 1931)

- Apex of paramere situated close to apex of median lobe; sensory peg setae forming an extensive apical field extended posteriad along lateral margins of paramere ( Smetana, 2007: figs. 47, 49). Aedeagus as in Smetana (2007: figs. 47–50). Length 14.0–18.0 mm........................................................ O. (Pseudocypus) imurai Smetana, 2005

42. Body bicolored, head and pronotum of a different color than elytra and abdomen.................................. 43

- Body not distinctly bicolored........................................................................... 44

43. Head and pronotum reddish-brown; elytra and abdomen piceous. Aedeagus with apex of median lobe acute and extremely elongate; paramere distinctly not reaching apex of median lobe ( Smetana, 2008: fig. 16). Length 13.0–17.0 mm.............................................................................. O. (Pseudocypus) bicoloris Smetana, 2008

- Head and pronotum blue; elytra and abdomen black. Median lobe of aedeagus with widely arcuate apex; paramere reaching apex of median lobe ( Smetana, 2008: fig. 22). Length 13.0–14.0 mm............ O. (Pseudocypus) caelestis Smetana, 2008

44. Aedeagus heavily pigmented, apical portion of median lobe and paramere almost totally black....................... 45

- Aedeagus not heavily pigmented, appearing brownish........................................................ 47

45. Aedeagus with apical portion of median lobe elongate and subacute, shaped as in Figs. 3-1 B, C, 3-2 B-D. Length 20.6–24.3 mm ................................................................ O. (Pseudocypus) anguliculminis sp. nov.

- Aedeagus robust, with apical portion of median lobe not elongate.............................................. 46

46. Aedeagus with apical portion of both median lobe and paramere widely arcuate; median lobe distinctly asymmetrical (Figs. 11- 1 B, C, 11-2 B–D, F); underside of paramere with sensory peg setae arranged along apical side, not numerous (Figs. 11-1 D, 11-2 A). Length 15.0– 19.1 mm ........................................... O. (Pseudocypus) calamis Smetana, 2007

- Aedeagus with apical portion of median lobe subacute, almost symmetrical (Fig. 4-2 F); paramere with widely arcuate apex; numerous sensory peg setae arranged along apical and lateral sides of underside of paramere (Figs. 4-1 D, 4-2 G). Aedeagus as in Figs. 4-1 B, C, 4-2 B–D. Length 21.9–26.8 mm ............................ O. (Pseudocypus) digiticulminis sp. nov.

47. Aedeagus with apical portion of median lobe hat-like ( Figs. 1-1 C, 1-2 F ); apical portion of paramere almost evenly narrowed toward subacute apex; sensory peg setae arranged along apical and lateral sides of underside of paramere, not numerous ( Figs. 1-1 D, 1-2 G ). Aedeagus as in Figs. 1-1 B, C, 1-2 B–D . Length 22.4–25.3 mm .... O. (Pseudocypus) pileaticulminis sp. nov.

- Aedeagus distinctly different............................................................................ 48

48. Interspaces between punctures on head and pronotum without appreciable microsculpture, surface therefore relatively shiny. Aedeagus as in Smetana (2007: figs. 83–86). Small species. Length 10.0–12.0 mm... O. (Pseudocypus) dryas Smetana, 2007

- Interspaces between punctures on head and pronotum with microsculpture, surface therefore more or less dull. Larger species (except for O. (Pseudocypus) zeuxis ), length exceeding 13.0 mm............................................... 49

49. Sensory peg setae on underside of paramere pale, not pigmented ( Smetana, 2007: fig. 55). Aedeagus as in Smetana (2007: figs.

54, 55). Length 13.0–16.0 mm........................................... O. (Pseudocypus) sadales Smetana, 2007 - Sensory peg setae on underside of paramere pigmented, appearing black. Aedeagus different......................... 50

50. Median lobe with obliquely subtruncate apex............................................................... 51

- Median lobe with apex not obliquely subtruncate........................................................... 52

51. Paramere with arcuate carina on face away from median lobe ( Smetana, 2008: fig. 9). Aedeagus as in Smetana (2008: figs. 9–12). Length 11.0–15.0 mm............................................. O. (Pseudocypus) nigror Smetana, 2008

- Paramere without arcuate carina on face away from median lobe ( Smetana, 2007: fig. 90). Aedeagus as in Smetana (2007: figs. 90–93). Length 11.0–13.0 mm............................................ O. (Pseudocypus) zeuxis Smetana, 2007

52. Pronotal disc gradually becoming wider posteriorly from anterior third (Fig. 5-1 A). Aedeagus and male sternite 9 elongate (Figs. 5-1 B, C, 5-2 B–D, H). Length 24.5– 21.2 mm ............................ O. (Pseudocypus) alticulminis sp. nov.

- Pronotal disc with almost parallel lateral sides, aedeagus and male sternite 9 usually not elongate..................... 53

53. Paramere with sensory peg setae distinctly forming two groups, located on left and right side ( Smetana, 2008: fig. 5). Aedeagus as in Smetana (2008: figs. 3–5). Length 14.0–19.0 mm...................... O. (Pseudocypus) nabis Smetana, 2008

- Paramere with sensory peg setae not distinctly separated into two groups........................................ 54

54. Sensory peg setae on underside of paramere very numerous, extended considerably posteriad along left lateral margin ( Smetana, 2007: fig. 36); paramere on face away from median lobe with conspicuous longitudinal crest ( Smetana, 2007: fig. 34). Length 15.0–17.0 mm.......................................... O. (Pseudocypus) quiris Smetana, 2007 (pars)

- Sensory peg setae on underside of paramere moderately numerous, no more than moderately extended posteriad along right lateral margin; paramere without conspicuous crest on face away from median lobe, but with a fine carina.............. 55

55. Aedeagus with apical portion of paramere widely arcuate ( Fig. 2-1 D ). Aedeagus as in Figs. 2-1 B, C, 2-2 B–D . Length 18.4–21.9 mm ........................................................ O. (Pseudocypus) denticulminis sp. nov.

- Aedeagus different, with apical portion of paramere somewhat narrow.......................................... 56

56. Apical portion of paramere constricted into a narrow, acute apex (Figs. 15-1 D, 15-2 G). Aedeagus as in Figs. 15-1 B, C, 15-2 B–D. Sternite 9 of male genital segment deeply emarginate apically (Fig. 15-2 H). Length 16.0–19.0mm..................................................................................... O. (Pseudocypus) hecato Smetana, 2007

- Apical portion of paramere with subacute, triangular apex ( Smetana, 2007: figs. 26, 29, 30). Sternite 9 of male genital segment only slightly emarginate apically ( Smetana, 2007: fig. 24). Aedeagus as in Smetana (2007: figs. 26–30). Length 15.0–18.0 mm..................................................................... O. (Pseudocypus) vindex Smetana, 2007

Figures

FIGURE 1 - 1. Ocypus (Pseudocypus) pileaticulminis sp. nov.: A. habitus of species (2.0 mm); B. aedeagus, lateral view; (0.5 mm); C. aedeagus, ventral view (0.5 mm); D. underside of apical portion of paramere (0.5 mm); E. second gonocoxite of female genital segment (0.5 mm). Scale in parentheses.

FIGURE 2 - 3. Ocypus (Pseudocypus) denticulminis sp. nov.: A, B and C: variations of teeth on mandibles (1.0 mm); D. habitus of one of the special large specimens collected together with Ocypus (Pseudocypus) persiciculminis sp. nov. (2.0 mm). Scale in parentheses.

FIGURE 1 - 2. Ocypus (Pseudocypus) pileaticulminis sp. nov.: A. male tergite 10; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. female tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. underside of apical portion of paramere; H. male sternite 9; I. second gonocoxite of female genital segment; J. male sternite 8. Scale bars = 0.50 mm.