Ocypus (Pseudocypus) graeseri Eppelsheim, 1887

He, Liang & Zhou, Hong-Zhang, 2017, Taxonomy of the subgenus Pseudocypus Mulsant & Rey (Coleoptera: Staphylinidae: Staphylinini) and eight new species from China, Zootaxa 4339 (1), pp. 1-77 : 43-44

14. Ocypus (Pseudocypus) graeseri Eppelsheim, 1887

(Figs. 14-1, 14-2)

Eppelsheim, 1887: 424 ( Ocypus ; Type locality: Wladiwostok); Bernhauer & Schubert, 1914: 386 ( Staphylinus ; synonym of fuscatus ); J. Müller, 1925: 45 ( Staphylinus ; subgenus Pseudocypus ; characters; Siberia); Scheerpeltz, 1933: 1401 ( Staphylinus ; subspecies of rambouseki ); Coiffait, 1974: 503 ( Pseudocypus ; valid species; characters; Siberia); Smetana & Davies, 2000: 44 ( Ocypus ; subgenus Pseudocypus ); Smetana, 2009b: 15 ( Ocypus ; Neotype designated); Schülke & Smetana, 2015: 1092 ( Ocypus ; subgenus Pseudocypus ; Catalog of Palaearctic Region).

Subspecies and Synonyms:

Staphylinus graeseri brunneopubens J. Müller, 1926 Ocypus graeseri graeseri Eppelsheim, 1887

Ocypus fuscatoides Coiffait, 1964

Staphylinus rambouseki J. Müller, 1925

Material examined. CHINA: Beijing Municipality: 1 female, Dongling Shan, 900 m, secondary shrub, pitfall traps, 12–15.VI.2012, coll. Xiaodong Yu (IZ-CAS) ; 1 male, 2 females, Dongling Shan, 1000 m, broad-leaved mixed forest, pitfall traps, 12–15.VI.2012, coll. Xiaodong Yu (IZ-CAS); 1 male, 2 females, Dongling Shan, 1100 m, broad-leaved forest, pitfall traps, 14–18.VI.2011, coll. Xiaodong Yu (IZ-CAS); 4 males, 2 females, Dongling Shan, 1100 m, broad-leaved mixed forest, pitfall traps, 12–15.VI.2012, coll. Xiaodong Yu (IZ-CAS); 2 females, Dongling Shan, 1140 m, larch forest, pitfall traps, 13–16.VII.2011, coll. Xiaodong Yu (IZ-CAS); 1 female, Dongling Shan, 1140 m, larch forest, pitfall traps, 16–19.VII.2012, coll. Xiaodong Yu (IZ-CAS); 1 male, 2 females, Dongling Shan, 1160 m, Juglans mandshurica forest, pitfall traps, 14–17.VI.2011, coll. Xiaodong Yu (IZ-CAS); 1 female, Dongling Shan, 1160 m, Juglans mandshurica forest, pitfall traps, 13–16.VIII.2011, coll. Xiaodong Yu (IZ-CAS); 3 males, 3 females, Dongling Shan, 1160 m, Juglans mandshurica forest, pitfall traps, 17–20.V.2012, coll. Xiaodong Yu (IZ-CAS); 2 females, Dongling Shan, 1160 m, Juglans mandshurica forest, pitfall traps, 13–16.VII.2012, coll. Xiaodong Yu (IZ-CAS); 1 male, 3 females, Dongling Shan, 1190 m, larch forest, pitfall traps, 14–17.VII.2011, coll. Xiaodong Yu (IZ-CAS); 1 male, 1 female, Dongling Shan, 1190 m, larch forest, pitfall traps, 16–19.V.2012, coll. Xiaodong Yu (IZ-CAS); 1 female, Dongling Shan, 1190 m, larch forest, pitfall traps, 16–19.VII.2012, coll. Xiaodong Yu (IZ-CAS); 1 female, Dongling Shan, 1190 m, larch forest, pitfall traps, 11–14.VIII.2012, coll. Xiaodong Yu (IZ-CAS); 3 males, Dongling Shan, 1200 m, oak-birch forest, pitfall traps, 12–15.VI.2012, coll. Xiaodong Yu (IZ-CAS); 1 male, Dongling Shan, 1225 m, deciduous broad leaved mixed forest, pitfall traps, 17–20.V.2012, coll. Xiaodong Yu (IZ-CAS); 1 male, Dongling Shan, 1250 m, oak forest, pitfall traps, 14–17.VI.2011, coll. Xiaodong Yu (IZ-CAS); 1 male, 1 female, Dongling Shan, 1250 m, oak forest, pitfall traps, 17–20.V.2012, coll. Xiaodong Yu (IZ-CAS); 2 males, Dongling Shan, 1250 m, oak forest, pitfall traps, 15–18.VII.2012, coll. Xiaodong Yu (IZ-CAS); 2 males, 2 females, Dongling Shan, 1300 m, oak-birch forest, pitfall traps, 12–15.VI.2012, coll. Xiaodong Yu (IZ-CAS); 1 male, Dongling Shan, 1400 m, oak-birch forest,, pitfall traps, 12–15.VI.2012, coll. Xiaodong Yu (IZ-CAS); 2 females, Dongling Shan, 1630 m, oak-birch forest, pitfall traps, 16–20.VI.2011, coll. Xiaodong Yu (IZ-CAS); 2 females, Dongling Shan, 1630 m, oak-birch forest, pitfall traps, 14–17.VII.2011, coll. Xiaodong Yu (IZ-CAS); 3 females, Dongling Shan, 1730 m, birch forest, pitfall traps, 13–16.VI.2012, coll. Xiaodong Yu (IZ-CAS); 1 male, Dongling Shan, 1820 m, birch forest, pitfall traps, 16–19.VI.2011, coll. Xiaodong Yu (IZ-CAS); 1 male, Dongling Shan, 1935 m, birch forest, pitfall traps, 16–19.VI.2011, coll. Xiaodong Yu (IZ-CAS); 3 males, 2 females, Dongling Shan, 2000 m, birch forest, pitfall traps, 16–19.VI.2011, coll. Xiaodong Yu (IZ-CAS); 1 female, Dongling Shan, 2100 m, subalpine meadow, pitfall traps, 24–27.VI.1999, coll. Xiaodong Yu (IZ-CAS); 1 female, Dongling Shan, 2100 m, subalpine meadow, pitfall traps, 17–20.VII.1999, coll. Xiaodong Yu (IZ-CAS); 9 males, 1 female, Dongling Shan, 2300 m, pitfall traps, 21–23.IX.1998, coll. Xiaodong Yu, Junjian He (IZ-CAS); 1 male, Xiaolongmen, 1120–1170 m, pitfall traps, 25–30.VI.1998, coll. Haisheng Zhou (IZ-CAS); 4 males, 2 females, Xiaolongmen, 1120–1170 m, pitfall traps, 25.VI–4.VII.1998, coll. Haisheng Zhou (IZ-CAS); 1 male, Yanqing County, Songshan National Nature Reserve, 720–850 m, individually collected, under stone, 22.V.2010, coll. Xinlei Huang (IZ-CAS) ; 1 female, Yanqing County, Songshan National Nature Reserve, 797 m, individually collected, donkey dung, 22.V.2010, coll. Liang Lv, Xi Zhang (IZ-CAS) ; Heilongjiang Province: 1 male, 1 female, Harbin, 30.V.1946 (IZ-CAS) ; 8 males, 11 females, Harbin, 19–22.V.1946 (IZ-CAS); 1 female, Harbin, 30.VIII.1946 (IZ-CAS); Inner Mongolia Autonomous Region [ New Record ]: 1 male, Helan Shan, Shuimogou, 2152 m, 27.VII.2015, coll. Guijun Yang (IZ-CAS) ; Ningxia Hui Autonomous Region [ New Record ]: 8 males, 3 females, Helan Shan, Suyukou, 2164 m, 17.VI.2008, coll. Guijun Yang (IZ-CAS) ; 2 males, 5 females, Helan Shan, Suyukou, 2280 m, 17.VI.2008, coll. Guijun Yang etc. (IZ-CAS); 1 male, 1 female, Xiji County, 24.VII.1989, coll. Guodong Ren (IZ-CAS).

Measurements. Body length: 16.5–19.7 mm.CL: 0.68; EL: 2.50; ELS: 1.28; EW: 2.72; HL: 1.89; HW: 2.16; PO: 0.68; PL: 2.57; PW: 2.40.

Distribution. Ocypus graeseri is widely distributed in Mongolia, southern portions of eastern Siberia, Russian Far East and north of China (Beijing, Hebei, Heilongjiang, Inner Mongolia, Ningxia, Qinghai and Shanxi provinces).