Glandularia J.F. Gmelin (1791

Cardoso, Pedro Henrique, Valério, Vanessa Imaculada Dos Reis, Neto, Luiz Menini & Salimena, Fátima Regina Gonçalves, 2021, Verbenaceae in Espírito Santo, Brazil: richness, patterns of geographic distribution and conservation, Phytotaxa 484 (1), pp. 1-43 : 12

5. Glandularia J.F. Gmelin (1791 [1792]: 886).

Herbs, monoecious, branches without prickles, not aromatic. Leaves petiolate, glands near to the petiole absent. Inflorescences terminal, compound, racemes spiciform, multiflorous; rachis elongated; bracts membranaceous, green; bracteoles absent. Flowers sessile; calyx tubular, shorter than the corolla, membranaceous, green, 5-lobed, accrescent in fruit; corolla zygomorphic, 5-lobed; stamens 4, included, didynamous, superior pair with or without connective appendages, thecae parallel; ovary 2-carpelate, 4-loculate, ovules 1 per locule, style three times longer than the ovary, stigma 2-lobed. Fruit a schizocarp, with 4 cluses, brown, external surface reticulate.

Glandularia is part of the tribe Verbeneae ( Marx et al. 2010) and presents 88 species disjunctly distributed throughout the subtropical and temperate regions of the North and South America ( Peralta & Thode 2010, Thode & Bordignon 2019, Cardoso et al. 2020c, Cardoso et al. in press.). The genus is characterized by the style three or more times longer than the ovary, stem with cortical chlorenchyma in a continuous band, and chromosomic number equal to five and multiples ( O’Leary & Peralta 2007). The main diversity centers of Glandularia are the south region of Brazil and northeastern Argentina ( O’Leary & Thode 2016). In Brazil, there are 34 species recorded, of which 13 are endemic and 26 are found in the Atlantic Forest domain ( O’Leary & Thode 2016, Thode & Bordignon 2019, Cardoso et al. 2020c, O’Leary et al. 2020). One species occurs in the state of Espírito Santo.