Glomomidiella infrapermica, Vachard, Krainer & Lucas, 2015

Vachard, D, Krainer, K & Lucas, SG, 2015, Late Early Permian (late Leonardian; Kungurian) algae, microproblematica, and smaller foraminifers from the Yeso Group and San Andres Formation (New Mexico; USA), Palaeontologia Electronica (English ed.) 3 (8), pp. 1-77 : 54-56

Glomomidiella infrapermica n. sp.

Figures 22.1, 22.3 , 23.1, 23.3, 23.7, 23.8 , 35.9, 35.11, 35.14 , 36.1-36.12, 36.16

zoobank.org/ D88B37EB-ED93-4EAA-899C-6C338B784706

? 1982 Orthovertella sp. ; Wang, p. 19, pl. 4, fig. 2.

v. 2013b Orthovertellopsis ; Lucas, Krainer and Voigt, fig. 8I.

v. 2013 b Orthovertella ? sp. 1 (partim); Vachard, Krainer and Lucas, p. 348 (no illustration).

Etymology. Latin infrapermicus, from the Lower Permian strata.

Holotype. NMMNH P-67600; Figure 22.1 ; sample MG 10-22 .

Paratypes. NMMNH P-67601-67605; Figures 22.3 , 23.1, 23.3, 23.7-8 .

Material. 35 specimens.

Locus Typicus. NMMNH locality 8904; Yeso Group, Los Vallos Formation, Torres Member, Massacre Gap in the southern Fra Cristobal Mountains (New Mexico).

Stratum Typicum. Early? Kungurian (late Leonardian).

Diagnosis. Large Glomomidiella with weak pseudosepta, numerous whorls and last whorls semi-evolute to evolute.

Description. Test ovoid to circular or triangular (D = (rarely 0.24)- 0.35-0.69 mm). The spherical proloculus is proportionally very large (up to 0.09 mm). The coiling is almost entirely streptospiral, but becomes planispiral and perhaps a little uncoiled at the end of the tube growth. There are (rarely 3.5)- 4-5 involute whorls. Sutures weakly marked between the different whorls of coiling. Lateral sides of spire irregular. Peripheral margin rounded. No umbilicus. The tube is undivided at the beginning but can show some pseudosepta in the two last whorls ( Figures 22.1, 22.3 , 23.1, 23.3, 23.7-8 ). Tube walls moderately thick to thick (0.02-0.05 mm), always and entirely neosparitized. Lumina crescentic (h = 0.06-0.13 mm).

Remarks. Orthovertella sp. sensu Wang, 1982 is remarkably similar but larger (D = 1.25 mm). The new species differs from G. nestellorum by lower lumina, more numerous whorls and occasionally a semi-evolute last whorl. It differs from Glomomidiella americana (Nestell and Nestell, 2006) n. comb. by less developed pseudosepta and higher lumina; from Glomomidiella miscella (Nestell and Nestell in Nestell et al., 2006) n. comb., which is very similar and probably the direct descendent of G. infrapermica n. sp.; the latter species differs by smaller dimensions of the test and the proloculus and especially by lower chambers (h = 0.10-0.18 mm); and from Glomomidiella darvasica n. comb., another similar species, by larger dimensions, a smaller proloculus, and more whorls (5-7).

Neodiscus grandis Lin, Li and Sun, 1990 is relatively similar to G. infrapermica n. sp., although much larger (D = 1.09-1.51 mm), but it is also related to some species of Iran and Armenia that exhibit sporadic pseudosepta and even rare true septa (materials of Mobin Ebrahim Nejad and Gevorg Grigoryan), and consequently are apparently convergent with Baisalina .

Occurrence. Kungurian (late Leonardian) of New Mexico (Yeso Group, Los Vallos Formation, Torres Member, Massacre Gap: samples MG 10-2, MG(2) 10-3, MG(2) 10-4, MG(2) 10-5, MG(2) 10-7b, MG(2) 10-7c, MG(2) 10-8, MG(2) 10-9, MG(2) 10- 10, MG(2) 10-11, MG 10-12, MG(2) 10-13, MG 10- 15, MG 10-16, MG 10-22, MG 10-25, MG(2) 12-7). San Andres Formation, McLeod Hills: samples MLY 6-26, MLY(2) 5-12, MLY(2) 5-19a, MLY(2) 5- 19c, MLY(2) 5-20, MLY(2) 5-24, MLY(2) 5a-5, MLY(2) 5a-6, MLY(2) 5a-7, MLY(2) 5a-15, MLY(2) 5a-22, MLY(2) 6-18, SAR 11-14). Permian of South China.

Figures

FIGURE 22. 1, 3. Glomomidiella infrapermica n. sp. 1. NMMNH P-67600, holotype. Subtransverse section. Yeso Group. Sample MG 10-22. 3. NMMNH P-67601, paratype. Subtransverse section. Yeso Group. Sample MG 10-25. 2. Orthovertellopsis proteaeformis n. gen. n. sp., NMMNH P-67570, holotype. Axial section with relatively well-preserved wall. Yeso Group. Sample MG 10-21. 4, 7-8, 10, 12-14. Praeneodiscus convexus n. gen. n. sp. 4. NMMNH P-67606, holotype. Axial section. Yeso Group. Sample MG 10-24. 7. NMMNH P-67607, paratype. Axial section. San Andres Fm. Sample MLY 5-13 (already illustrated in Brose et al., 2013, figure 6. 11). 8. NMMNH P-67608, paratype. Axial section. San Andres Fm. Sample MLY 5-4. 10. NMMNH P-67609, paratype. Axial section with a relatively preserved wall. San Andres Fm. Sample MLY 5-7. 12. NMMNH P-67610, paratype. Axial section. San Andres Fm. Sample MLY 5-7a. 13. NMMNH P-67611, paratype. Axial section. San Andres Fm. Sample MLY 5-30. 14. NMMNH P-67612, paratype. Axial section. San Andres Fm. Sample MLY 5-21. 5. Tubiphytes epimonellaeformis n.sp. Subtransverse section. San Andres Fm. Sample SAR 8a-22. 6. Globivalvulina parapiciformis n. sp. NMMNH P-67551, holotype. Transverse section. San Andres Fm. Sample MLY 4-3. 9. Pseudospira aff. ishimbaica (Lipina, 1949). Subtransverse section. Yeso Group. Sample MG 10-26. 11. Hemigordiellina? cf. simplex (Harlton, 1928). Axial section. San Andres Fm. Sample MLY 6-28. 15. Eotuberitina reitlingerae Miklukho-Maklay, 1958. Axial section. San Andres Fm. Sample MLY 4-5. Scale bars equal 0.10 mm.

FIGURE 23. 1, 3, 7-8. Glomomidiella infrapermica n. sp. 1. NMMNH P-67602, paratype. Subtransverse section. Yeso Group. Sample MG 10-12. 3. NMMNH P-67603, paratype. Oblique subaxial section. Yeso Group. Sample MG 10-15. 7. NMMNH P-67604, paratype. Two subtransverse sections. Yeso Group. Sample MG 10-2. 8. NMMNH P-67605, paratype. Subtransverse section. Yeso Group. Sample MG 10-16. 2, 4, 6. Orthovertellopsis proteaeformis n. gen. n. sp. 2. NMMNH P-67571, paratype. Subaxial section. Yeso Group. Sample MG 10-23. 4. NMMNH P-67572, paratype. Subaxial section. Yeso Group. Sample MG 10-13a. 6. NMMNH P-67573, two paratypes. Subtransverse section. Yeso Group. Sample MG 10-20. 5. Calcitornella cf. elongata Cushman and Waters, 1928a. Subaxial section. Yeso Group. Sample MG 10-20a. Scale bars equal 0.10 mm.

FIGURE 35. 1-5, 8. Olgaorlovella davydovi n. gen. n. sp. 1. Axial section. San Andres Fm. Sample SAR 11a-11. 2. Axial section. San Andres Fm. Sample SAR 11a-8 (already illustrated in Brose et al., 2013, figure 6. 12). 3. Subaxial section. San Andres Fm. Sample SAR 11a-15. 4. Axial section. San Andres Fm. Sample SAR 11a-14. 5. Axial section. San Andres Fm. Sample SAR 11a-3. 8. Axial section. San Andres Fm. Sample SAR 11-6. 6-7, 13. Orthovertellopsis calcitornellaeformis n. gen. n. sp. 6. Subtransverse section. San Andres Fm. Sample SAR 11a-16. 7. Transverse section. San Andres Fm. Sample SAR 11-16. 13. Oblique section. San Andres Fm. Sample SAR 11a-5. 9, 11, 14. Glomomidiella infrapermica n. sp. 9. Axial section showing the proloculus. San Andres Fm. Sample SAR 11-14. 11. Subtransverse section. Yeso Group. Sample MG(2) 10-7b. 14. Axial section relatively well preserved (dark wall). Yeso Group. Sample MG(2) 12-7. 10, 12. Orthovertellopsis protaeformis n. gen. n. sp. 10. Axial section. Yeso Group. Sample MG(2) 10-14. 12. Subaxial section. Yeso Group. Sample MG(2) 10-6. Scale bars equal 0.10 mm.

FIGURE 36. 1-12, 16. Glomomidiella infrapermica n. sp. 1. Axial section. Yeso Group. Sample MG(2) 10-9. 2. Subaxial section. San Andres Fm. Sample MLY(2) 5a-15. 3. Oblique subaxial section. Yeso Group. Sample MG(2) 10-8. 4. Subtransverse section. Yeso Group. Sample MG 10-7. 5. Broken subaxial section. Yeso Group. Sample MG(2) 10-10. 6. Subaxial section. Yeso Group. Sample MG(2) 10-4. 7. Axial section. Yeso Group. Sample MG(2) 10-5. 8. Oblique subaxial section. Yeso Group. Sample MG(2) 10-3. 9. Subaxial section. Yeso Group. Sample MG(2) 10-11. 10. Axial section. Yeso Group. Sample MG 10-7a. 11. Subtransverse section. Yeso Group. Sample MG(2) 10-7c. 12. Subaxial section. Yeso Group. Sample MG(2) 10-13 (already illustrated in Lucas et al., 2013b, figure 8I). 16. Poorly preserved subaxial section. San Andres Fm. Sample MLY 6-26. 13. Olgaorlovella davydovi n. gen. n. sp. Axial section. San Andres Fm. Sample SAR(2) 11-7. 14-15. Geinitzina indepressa Cherdyntsev, 1914. 14. Oblique subaxial section. San Andres Fm. Sample SAR(2) 11-2. 15. Subaxial section. San Andres Fm. Sample SAR(2) 19-1. 17. “Frondicularia” aff. turae Baryshnikov in Baryshnikov, Zolotova and Kosheleva, 1982. Axial section. Yeso Group. Sample MG 13f.