Praeneodiscus convexus, Vachard, Krainer & Lucas, 2015

Vachard, D, Krainer, K & Lucas, SG, 2015, Late Early Permian (late Leonardian; Kungurian) algae, microproblematica, and smaller foraminifers from the Yeso Group and San Andres Formation (New Mexico; USA), Palaeontologia Electronica (English ed.) 3 (8), pp. 1-77 : 56-58

Praeneodiscus convexus n. gen. n. sp.

Figures 22.4, 22.7, 22.8, 22.10, 22.12-22.14 , 33.17, 33.22, 33.26 , 34.14

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? v. 2009 Midiella ? n. sp.; Krainer, Lucas and Vachard, pl. 4, figs. 1, 2, 3.

v. 2013 Midiella ; Brose, Lucas and Krainer, fig. 6.11.

v. 2013 b Neodiscus ? ex gr. ovatus (Grozdilova) ; Vachard, Krainer and Lucas, p. 348 (no illustration).

Etymology. A species with regularly convex flanks.

Holotype. NMMNH P-67606; Figure 22.4 ; sample MG 10-24 .

Paratypes. NMMNH P-67607-67612; Figure 22.7- 8, 22.10, 22.12-14 .

Material. 40 specimens.

Locus Typicus. NMMNH locality 8905; Yeso Group, Los Vallos Formation, Torres Member, Massacre Gap, southern Fra Cristobal Mountains (New Mexico).

Stratum Typicum. Early? Kungurian (late Leonardian).

Diagnosis. Praeneodiscus characterised by an ovoid test, medium to large in diameter, with oscillating coiling and an ogival lumen of the tubular chamber.

Description. Test ovoid. Oscillating involute coiling. No umbilici. No sutures. Wall surface smooth. Lateral sides of spire strongly inflated. Peripheral margin rounded. Proloculus spherical and small to moderate in size, followed by a tubular chamber with a diameter that is equal to the height of the third volution. D = 0.35-0.50-(0.70) mm; w = (0.14)- 0.22-0.35 mm; w/D = 0.50-0.70; whorls: (4)-5-6; p = 0.01-0.02-(0.05-0.06) mm; h = 0.05-0.10 mm; s = 0.01-0.02 mm.

Remarks. The new species differs from Praeneodiscus arcticus n. comb. (formerly Midiella arctica ) by a less sigmoidal coiling and a less inflated test (probably 0.66-0.71 in this latter species; although the ratio D/w indicated by Blazejowski, 2009 is 2.4- 2.5; a probable lapsus calami for 1.4-1.5). It differs from P. compressus ( Lipina, 1949) n. comb. by more whorls and more aligned last whorls. In contrast to P. ovatus ( Grozdilova, 1956) n. comb. which is relatively similar in the shape of the tube lumina (i.e., crescentic, very wide, and well-developed in the lateral sides), the lumina of P. convexus n. gen. n. sp. are smaller, semicircular and limited to the central part of the test; the test, itself, of P. ovatus is much larger with D = 0.490 -0.990 mm; w = 0.340 -0.480 mm (for the same number of whorls: 5-6); h = 0.076 -0.150 mm; s = 0.026 -0.038 mm. Praeneodiscus darvasica n. comb. is similar to P. ovatus ; it differs from P. convexus n. gen. n. sp. by a shorter lumen of the chambers and a glomus more developed by the internal and external whorls. The holotype of P. convexus n. gen. n. sp. ( Figure 19.4 ) is similar to Neodiscus padangensis sensu Kobayashi (2006 , figures 4, 5), but differs by its smaller size (1.00 mm for the Kobayashi’s specimen), its smaller proloculus, and the total similarity of all lumina, whereas Kobayashi’s specimen seems to exhibit a juvenarium, as does the holotype of N. milliloides Miklukho-Maklay, 1953 or Neodiscus cf. N. milliloides sensu Nestell et al., 2009 (plate 1, figures 30, 31). Praeneodiscus convexus n. gen. n. sp. differs from P. longus n. comb. and P. permicus n. comb. by the more inflated test and therefore a larger width; from Midiella zaninettiae ( Altiner, 1978) by larger dimensions and a more oscillating coiling; from Neodiscus quinqlongensis Wang, 1976 by a test less inflated test with more volutions for a similar diameter; and from Neohemigordius zulumartensis Filimonova, 2010 by larger dimensions.

Occurrence. Early?-middle? Kungurian (late Leonardian) of New Mexico (Yeso Group, Los Vallos Formation, Torres Member: sample MG 10-24; San Andres Formation, McLeod Hills: samples MLY 5-4, MLY(2) 5-5, MLY 5-7, MLY 5-7a, MLY(2) 5-8, MLY 5-8a, MLY(2) 5-8a, MLY 5-9, MLY 5-13, MLY(2) 5-19b, MLY 5-21, MLY 5-30, MLY 5-31, MLY 5-32, MLY(2) 5a-14, MLY(2) 5a-16, MLY(2) 5a-17, MLY(2) 5a-18, MLY 8-1).

Figures

FIGURE 22. 1, 3. Glomomidiella infrapermica n. sp. 1. NMMNH P-67600, holotype. Subtransverse section. Yeso Group. Sample MG 10-22. 3. NMMNH P-67601, paratype. Subtransverse section. Yeso Group. Sample MG 10-25. 2. Orthovertellopsis proteaeformis n. gen. n. sp., NMMNH P-67570, holotype. Axial section with relatively well-preserved wall. Yeso Group. Sample MG 10-21. 4, 7-8, 10, 12-14. Praeneodiscus convexus n. gen. n. sp. 4. NMMNH P-67606, holotype. Axial section. Yeso Group. Sample MG 10-24. 7. NMMNH P-67607, paratype. Axial section. San Andres Fm. Sample MLY 5-13 (already illustrated in Brose et al., 2013, figure 6. 11). 8. NMMNH P-67608, paratype. Axial section. San Andres Fm. Sample MLY 5-4. 10. NMMNH P-67609, paratype. Axial section with a relatively preserved wall. San Andres Fm. Sample MLY 5-7. 12. NMMNH P-67610, paratype. Axial section. San Andres Fm. Sample MLY 5-7a. 13. NMMNH P-67611, paratype. Axial section. San Andres Fm. Sample MLY 5-30. 14. NMMNH P-67612, paratype. Axial section. San Andres Fm. Sample MLY 5-21. 5. Tubiphytes epimonellaeformis n.sp. Subtransverse section. San Andres Fm. Sample SAR 8a-22. 6. Globivalvulina parapiciformis n. sp. NMMNH P-67551, holotype. Transverse section. San Andres Fm. Sample MLY 4-3. 9. Pseudospira aff. ishimbaica (Lipina, 1949). Subtransverse section. Yeso Group. Sample MG 10-26. 11. Hemigordiellina? cf. simplex (Harlton, 1928). Axial section. San Andres Fm. Sample MLY 6-28. 15. Eotuberitina reitlingerae Miklukho-Maklay, 1958. Axial section. San Andres Fm. Sample MLY 4-5. Scale bars equal 0.10 mm.

FIGURE 33. 1-14, 16, 18. Hemigordiellina? cf. simplex (Harlton, 1928). 1. Oblique subtransverse section. San Andres Fm. Sample MLY 3-1. 2. Oblique subaxial section. San Andres Fm. Sample MLY 3-11. 3. Subtransverse section. San Andres Fm. Sample MLY 4-8. 4. Subaxial section. San Andres Fm. Sample MLY 4-11. 5. Transverse section. San Andres Fm. Sample MLY 4-15. 6. Subtransverse section. San Andres Fm. Sample MLY 4-16. 7. Transverse section. San Andres Fm. Sample MLY 4-18. 8. Subaxial section. San Andres Fm. Sample MLY 4-19. 9. Subtransverse section. San Andres Fm. Sample MLY 8-7. 10. Subtransverse section. San Andres Fm. Sample MLY 4-20. 11. Subtransverse section. San Andres Fm. Sample MLY 5-15. 12. Subaxial section. San Andres Fm. Sample MLY 5-16a. 13. Transverse section. San Andres Fm. Sample MLY 5-5. 14. Subtranverse section. San Andres Fm. Sample MLY 4-12. 16. Oblique section. San Andres Fm. Sample MLY 6-25a. 18. Transverse section. San Andres Fm. Sample MLY 4-21. 15, 21. Pseudospira aff. ishimbaica (Lipina, 1949). 15. Subaxial section. San Andres Fm. Sample MLY 6-22. 21. Axial section with Hemigordiellina? cf. simplex (left). San Andres Fm. Sample MLY 5-17. 17, 22, 26. Praeneodiscus convexus n. gen. n. sp. 17. Oblique section. San Andres Fm. Sample MLY 5-16. 22. Transverse section. San Andres Fm. Sample MLY 5-8a. 26. Subaxial section. San Andres Fm. Sample MLY 5-9. 19-20. Orthovertellopsis protaeformis n. gen. n. sp. 19. Subaxial section. San Andres Fm. Sample MLY 6-5. 20. Oblique subaxial section. San Andres Fm. Sample MLY 6-5a. 23-24. Hemigordiellina? aff. pseudopusilla (Baryshnikov in Zolotova and Baryshnikov, 1980). 23. Oblique section. San Andres Fm. Sample MLY 5-25. 24. Axial section. San Andres Fm. Sample MLY 5-24. 25. Palaeonubecularia? sp. Axial section. Perhaps also related with Ellesmerella (see Vachard et al., 2013b). San Andres Fm. Sample MLY 5-29. Scale bars equal 0.10 mm.

FIGURE 34. 1-5. Hemigordius aff. saranensis Baryshnikov in Baryshnikov, Zolotova and Kosheleva, 1982. 1. Subaxial section. San Andres Fm. Sample SAR 13-5d. 2. Subaxial section. San Andres Fm. Sample SAR 11-5a. 3. Axial section. San Andres Fm. Sample SAR 20-5. 4. Transverse section. San Andres Fm. Sample SAR 11a-12. 5. Subtransverse section. San Andres Fm. Sample SAR 13-11. 6, 10, 17. Orthovertellopsis calcitornellaeformis n. gen. n. sp. 6. Subtransverse section. San Andres Fm. Sample SAR 19-1. 10. Oblique section. San Andres Fm. Sample MLY 8-6. 17. Transverse section. San Andres Fm. Sample SAR(2) 11-10. 7-8. Palaeonubecularia sp. 7. Subtransverse section. San Andres Fm. Sample MLY 5-19. 8. Subtransverse section encrusting a Boueina? crassundia n. sp. San Andres Fm. Sample MLY(2) 5a-19. 9. Pseudospira aff. ishimbaica (Lipina, 1949). Two random sections. San Andres Fm. Sample MLY 6- 23. 11-12. Hemigordellina cf. elegans (Lipina, 1949). 11. Random section. San Andres Fm. Sample SAR 18-3a. 12. Random section. San Andres Fm. Sample SAR 18-3. 13. Ammovertella sp. Transverse section. San Andres Fm. Sample SAR 13-9a. 14. Praeneodiscus convexus n. gen. n. sp. Subaxial section. San Andres Fm. Sample MLY 5-32. 15, 18. Hemigordiellina? aff. simplex (Harlton, 1928). 15. Random section. San Andres Fm. Sample SAR 18-2. 18. Transverse section. San Andres Fm. Sample MLY(2) 4-2. 16. Hemigordellina aff. elegans (Lipina, 1949). Random section. San Andres Fm. Sample SAR 14. 19. Calcitornella cf. elongata Cushman and Waters, 1928a. Subtransverse section. San Andres Fm. Sample MLY(2) 5a-24. Scale bars 1-6, 10-19 equal 0.10 mm.

FIGURE 19. 1-2, 4-6, 9. Gymnocodium? johnsonii n. sp. 1. Subaxial section. San Andres Fm. Sample SAR 2-1. 2. Tangential section. San Andres Fm. Sample SAR 2-2. 4. Subaxial section. San Andres Fm. Sample SAR 2-3. 5. Tangential section. San Andres Fm. Sample SAR 2-4. 6. Subaxial section. San Andres Fm. Sample SAR 7-9. 9. Subaxial section. San Andres Fm. Sample SAR 8a-25 (already illustrated in Brose et al., 2013, figure 6.9). 3, 7-8, 10-16. Gymnocodium? bacillum n. sp. 3. Oblique subaxial section. San Andres Fm. Sample SAR 2-5. 7. Subaxial section. San Andres Fm. Sample SAR 8a-3. 8. Axial section. San Andres Fm. Sample SAR 8a-26. 10. Axial section. Yeso. Sample MG(2) 12-2 (already illustrated in Lucas et al., 2013b, figure 8D). 11. Axial section. San Andres. Sample SAR 8-8. 12. Axial section. San Andres Fm. Sample SAR 13-6. 13. Subaxial section. San Andres Fm. Sample SAR 8a-16. 14. Subaxial section. San Andres Fm. Sample SAR 8a-15. 15. Two tangential sections. San Andres Fm. Sample SAR 7- 8. 16. Subaxial section. San Andres Fm. Sample SAR 8a-23.