BOUEINEAE Shuysky

Vachard, D, Krainer, K & Lucas, SG, 2015, Late Early Permian (late Leonardian; Kungurian) algae, microproblematica, and smaller foraminifers from the Yeso Group and San Andres Formation (New Mexico; USA), Palaeontologia Electronica (English ed.) 3 (8), pp. 1-77 : 12-14

Tribe BOUEINEAE Shuysky in Chuvashov, Luchinina, Shuysky, Shaikin, Berchenko, Ishchenko, Saltovskaya and Shirshova, 1987

Description. Codiaceae with a medullar zone well to poorly preserved. Cylindrical thalli (or segments). Medullar zone with coarse, tangled siphons. Subcortical zone with subvertical siphons, acrophore to slightly phloiophore, more or less euspondyl. Cortical zone with perpendicular, thin, acrophore, finer, ramified siphons first parallel and then linked in tuft-like groups and showing several ramifications with constant diameter up to the outer surface.

Composition. Boueina Toula, 1884; Boueina ? (this study); Maslovina Obrhel, 1968 ; Pseudolitanaia Mamet and Préat, 1994; Bijagodella Chuvashov, Yuferev and Liuchinina, 1985 ; Poncetellina Mamet and Roux, 1984 ; Palaeoporella ? sp. sensu Vachard, 1988; Funiculus Shuysky and Shirshova in Chuvashov et al. (1987); Arabicodium Elliott, 1957 .

Remarks. Boueinae? differs from the other tribes of Palaeozoic Codiaceae listed above, and especially from Litanaiae Shuysky in Chuvashov et al. (1987), by the weak development or absence of inflated parts in the siphons and/or deltoid terminations (utricles). Some Devonian taxa have more affinities with Boueineae than Litanaiae or Palaeoporelleae Shuysky in Chuvashov et al. (1987), such as Maslovina meyenii Obrhel, 1968 ; M. australensis Mamet and Pohler, 2002 ; Pseudolitanaia graecensis [sic; to correct in graeca ] ( Hubmann, 1990) Mamet and Préat, 1994; Paralitanaia baileuxensis Mamet and Préat, 1985; Praelitanaia anirica ( Maslov, 1956) Shuysky in Chuvashov et al. (1987); Bijagodella bijagodensis ( Chuvashov, 1973) Chuvashov et al. (1985) ; etc. (the diagnoses of these taxa are compiled, for example, in Bassoullet et al. (1983)). Arabicodium seems to be morphologically transitional between Boueina and Halimeda Lamouroux, 1812 (see Elliott, 1982; Bassoullet et al., 1983).

Occurrence. FAD in the Late Silurian with Maslovina . First acme in the Middle Devonian with Funiculus . The tribe is present with Palaeoporella ? sp. at the Devonian-Carboniferous boundary ( Vachard, 1988, plate 1, figure 7). Boueina is known from the Late Triassic of Thailand ( Flügel, 1988) to Early Cretaceous ( Toula, 1884). LAD in early Eocene with Arabicodium ( Bassoullet et al., 1983) .