Globivalvulina retroseptata, sensu Filimonova, 2008

Vachard, D, Krainer, K & Lucas, SG, 2015, Late Early Permian (late Leonardian; Kungurian) algae, microproblematica, and smaller foraminifers from the Yeso Group and San Andres Formation (New Mexico; USA), Palaeontologia Electronica (English ed.) 3 (8), pp. 1-77 : 30-32

Globivalvulina retroseptata n. sp.

Figures 21.1-21.8 , 27.1, 27.4, 27.9, 27.20 , 28.1- 28.3, 28.7 , 29.1-29.4, 29.7 , 30.2, 30.3, 30.11, 30.13, 30.16 , 31.9-31.12 , 32.1-32.3, 32.6

zoobank.org/ BD9C7144-0795-47E1-84FD-6CCA76E1E64E

v. 2003 Globivalvulina spp. (partim); Krainer, Vachard and Lucas, pl. 8, figs. 2, 12.

v. p. 2013 Globivalvulina ; Brose, Lucas and Krainer, fig. 6.7 (only, non fig. 6.6 = G. parapiciformis n. sp.).

v. 2013 b Globivalvulina aff. kamensis Reitlinger ; Vachard, Krainer and Lucas, p. 348 (no illustration).

Etymology. With septa turned backwards (relatively similar to those of the genus Retroseptellina Gaillot and Vachard, 2007 ).

Holotype. NMMNH P-67553; Figure 21.4 ; sample SAR 13-2 .

Paratypes. NMMNH P-67554-67560; Figure 21.1- 3, 21.5-8 .

Material. 35 sections.

Locus Typicus. NMMNH locality 8898; San Andres Formation, reference section, San Andres Mountains near Rhodes Canyon (New Mexico).

Stratum Typicum. Late? Kungurian (latest Leonardian).

Diagnosis. Test medium-sized, often ovoid, with septa curved backward, funnel wide and shallow, and thin microgranular wall.

Description. Test ovoid to rounded trapezoidal; D = (0.23)- 0.30-0.60 mm; w = 0.28-0.55 mm; w/D = 0.63-1.25. Planispiral to trochospiral axis of biseriality. Proloculus spherical, small to medium-sized (0.03-0.08 mm; mainly 0.06-0.07 mm), moderately large, centred. Dorsal side gently curved, strongly convex with weak sutures, to irregular and humped. Sutures moderately deep to deep. Whorl: 1- 1.5. Peripheral margin semicircular to crescentic. Lateral sides of spire relatively flat. Umbilicus absent. Chambers: (4)-5-6-(rarely 8) pairs, semicircular to rounded, eventually deformed; the two last chambers are disproportionate with a strong increase in height as in Globivalvulina graeca Reichel, 1946 (h = (rarely 0.11)- 0.13-0.30 mm). Septa thin and curved backwards at their distal extremity. Ventral side flat to slightly concave. Aperture is a basal slit. Funnel wide and shallow. Oral valvula variously developed, from faint to developed, as in G. kamensis Reitlinger, 1950 . Wall thin, dark, undifferentiated, unilayered, and microgranular (s = 0.01-0.02 mm).

Remarks. The new species differs from all other species of Globivalvulina (except for those assigned to Retroseptellina ) by the curvature of septa. Some transverse sections are difficult to distinguish from G. kamensis (nevertheless, the axial sections of this species are poorly known), generally smaller with a diameter of 0.43-0.50 mm for Reitlinger (1950) and 0.43 mm according to Bogush (1963, p. 60). Although synonymized with Globivalvulina bulloides by Groves (1988), G. kamensis differs by its smaller dimensions, stronger valvular projections, and a microgranular wall that is never differentiated.

Occurrence. Kungurian (late Leonardian) of New Mexico (Yeso Group, Los Vallos Formation, Torres Member, Massacre Gap (MG): samples MG 10-9, MG(2) 10-12; and San Andres Formation, McLeod Hills (MLY) and reference section in the San Andres Mountains (SAR): samples MLY 5-3, MLY 5-23, MLY(2) 5a-5a, MLY(2) 6-13, MLY(2) 6-31, MLY 8-2a, SAR 3-1, SAR 3-4, SAR 8a-19, SAR 10- 3, SAR(2) 12-6, SAR 13-2, SAR 13-5b, SAR 13-5c, SAR 13-7, SAR 13-10, SAR(2) 13a-1, SAR(2) 13a- 2, SAR 13a-7, SAR 13a-8, SAR 13a-9, SAR 13a- 10, SAR 13a-11, SAR 19-2, SAR 19-3, SAR 19-4, SAR 20-1, SAR 20-6, SAR (A-1§)).

Figures

FIGURE 21. 1-8. Globivalvulina retroseptata n. sp. 1. NMMNH P-67554, paratype. Axial section showing the siphon formed by a curved backwards septum and an oral valvula. San Andres Fm. Sample SAR 10-3. 2. NMMNH P-67555, paratype. Transverse section. San Andres Fm. Sample SAR 12-6. 3. NMMNH P-67556, paratype. Subtransverse section. San Andres Fm. Sample SAR 12-7. 4. NMMNH P-67553, holotype. Transverse section. San Andres Fm. Sample SAR 13-2. 5. NMMNH P-67557, paratype. Subtransverse section. San Andres Fm. Sample SAR 13a-3a. 6. NMMNH P-67558, paratype. Oblique section. San Andres Fm. Sample SAR 13a-4. 7. NMMNH P-67559, paratype. Subtransverse section. San Andres Fm. Sample SAR 13a-6. 8. NMMNH P-67560, paratype. Oblique subtransverse section. San Andres Fm. Sample SAR 13a-3. 9-14. Globivalvulina praegraeca n. sp. 9. NMMNH P-67565, paratype. Subtransverse section. San Andres Fm. Sample SAR 7-1. 10, NMMNH P-67566, paratype. Subtransverse section. San Andres Fm. Sample SAR 7-2. 11, NMMNH P-67567, paratype. Oblique subtransverse section. San Andres Fm. Sample SAR 7-6. 12. NMMNH P-67564, holotype. Subtransverse section. San Andres Fm. Sample SAR 7-7. 13. NMMNH P-67568, paratype. Subaxial section. San Andres Fm. Sample MLY 3-3. 14. NMMNH P-67569, paratype. Axial section. San Andres Fm. Sample MLY 6-18. 15. Globivalvulina novamexicana n. sp., NMMNH P-67562, paratype. Oblique subaxial section. San Andres Fm. Sample MLY 5-22. Scale bars equal 0.10 mm.

FIGURE 27. 1, 4, 9, 20. Globivalvulina retroseptata n. sp. 1. Subaxial section. San Andres Fm. Sample MLY 8-2a. 4. Subtransverse section. San Andres Fm. Sample MLY 5-23. 9. Subaxial section. San Andres Fm. Sample MLY 5-3. 20. Transverse section. Yeso Group. Sample MG 10-9. 2, 5-8, 12-19. Globivalvulina novamexicana n. sp. 2. Subtransverse section. San Andres Fm. Sample MLY 4-13. 5. Oblique subaxial section. San Andres Fm. Sample MLY 4- 9. 6. Oblique section. San Andres Fm. Sample MLY4-1. 7. Axial section. San Andres Fm. Sample MLY 4-2. 8. Subaxial section. San Andres Fm. Sample MLY 4-6. 12. Axial section. San Andres Fm. Sample MLY 3-8. 13. Subtransverse section. Yeso Group. Sample MG 10-8 (already illustrated in Lucas et al., 2013b, figure 8G). 14. Subaxial section. Yeso Group. Sample MG 10-14. 15. Oblique subtransverse section. Yeso Group. Sample MG 10-17a. 16. Subtransverse section. Yeso Group. Sample MG 10-17. 17. Subaxial section. Yeso Group. Sample MG 10-7. 18. Subtransverse section. Yeso Group. Sample MG10 6. 19. Subaxial section. Yeso Group. Sample MG 10-6a. 3, 10- 11. Globivalvulina parapiciformis n. sp. 3. Axial section. San Andres Fm. Sample MLY 6-16. 10. Axial section. San Andres. Sample MLY 5-20a. 11. Axial section. San Andres Fm. Sample MLY 5-7c. Scale bars equal 0.10 mm.

FIGURE 28. 1-3, 7. Globivalvulina retroseptata n. sp. 1. Transverse section. San Andres Fm. Sample SAR 13-5. 2. Transverse section. San Andres Fm. Sample SAR 13-5b. 3. Transverse section. San Andres Fm. Sample SAR 13- 5c. 7. Transverse section. San Andres Fm. Sample SAR 8-5. 4-6, 8-12, 14-16. Globivalvulina praegraeca n. sp. 4. Transverse section. San Andres Fm. Sample SAR 13-7. 5. Transverse section. San Andres Fm. Sample SAR 8-2. 6. Subtransverse section. San Andres Fm. Sample SAR 8-3. 8. Subaxial section. San Andres Fm. Sample SAR 8-7a. 9. Subaxial section. San Andres Fm. Sample SAR 8a-2. 10. Subaxial section. San Andres Fm. Sample SAR 8a-5. 11. Axial section. San Andres Fm. Sample SAR 8a-5a. 12. Transverse section. San Andres Fm. Sample SAR 8a-18. 14. Subtransverse section. San Andres Fm. Sample SAR 8a-9. 15. Oblique subtransverse section. San Andres Fm. Sample SAR 8a-10. 16. Subtransverse section. San Andres Fm. Sample SAR 8a-12. 13. Globivalvulina parapiciformis n. sp. Transverse section. San Andres Fm. Sample SAR 8a-8a. Scale bars equal 0.10 mm.

FIGURE 29. 1-4, 7. Globivalvulina retroseptata n. sp. 1. Transverse section. San Andres Fm. Sample SAR 19-2. 2. Transverse section. San Andres Fm. Sample SAR 19-3. 3. Transverse section. San Andres Fm. Sample SAR 19-4. 4. Subtransverse section with developed oral valvula. San Andres Fm. Sample SAR 20-1. 7. Teratogenic specimen. San Andres Fm. Sample SAR 3-1. 5, 8, 10-13. Globivalvulina praegraeca n. sp. 5. Transverse section. San Andres Fm. Sample SAR 20-3. 8. Axial section. San Andres Fm. Sample SAR 3-4. 10. Subaxial section. San Andres Fm. Sample SAR 8-1. 11-12. Two transverse sections. San Andres Fm. Sample SAR 7-4. 13. Subaxial section. San Andres Fm. Sample SAR 7-5. 6, 9. Globivalvulina novamexicana n. sp. 6. Subtransverse section. San Andres Fm. Sample SAR 20-4. 9. Axial section. San Andres Fm. Sample SAR 7-3. Scale bars equal 0.10 mm.

FIGURE 30. 1, 4-6, 8-10, 12, 14-15. Globivalvulina praegraeca n. sp. 1. Subtransverse section. San Andres Fm. Sample SAR 8a-13a. 4. Transverse section. San Andres Fm. Sample SAR 9-1. 6. Transverse section. San Andres Fm. Sample SAR 13-10. 8. Subaxial section. San Andres Fm. Sample SAR(2) 13-2. 9. Subtransverse section. San Andres Fm. Sample SAR(2) 13-6. 10. Transverse section. San Andres Fm. SAR 13-9. 12. Axial section showing the proloculus and the first whorl. San Andres Fm. Sample SAR(2) 8-21. 14. Transverse section. San Andres Fm. Sample MLY(2) 4-9. 15. Transverse section. San Andres Fm. Sample SAR 8-18. 2-3, 11, 13, 16. Globivalvulina retroseptata n. sp. 2. Subtransverse section. San Andres Fm. Sample SAR 8a-19. 3. Transverse section. San Andres Fm. Sample SAR 8a-24. 11. Transverse section. San Andres Fm. Sample SAR 13a-7. 13. Transverse section. San Andres Fm. Sample MLY(2) 6-13 (already illustrated by Brose et al., 2013, figure 6.7). 16. Axial section. San Andres Fm. Sample SAR(2) 12-6. 5. Globivalvulina parapiciformis n. sp. Subtransverse section. San Andres Fm. Sample SAR 13-9. 7. Globivalvulina novamexicana n. sp. Transverse section. San Andres Fm. Sample SAR 13-12. Scale bars equal 0.10 mm.

FIGURE 31. 1. Globivalvulina novamexicana n. sp. Axial section. San Andres Fm. Sample SAR 8-4. 2-8. Globivalvulina praegraeca n. sp. 2. Subtransverse section. San Andres Fm. Sample SAR 8-6. 3. Axial section. San Andres Fm. Sample SAR 8-7. 4. Subaxial section. San Andres Fm. Sample SAR 8a-10a. 5. Paratype. Oblique section. San Andres Fm. Sample SAR 8a-20. 6. Subaxial section. San Andres Fm. Sample SAR 8a-21. 7. Subaxial section. San Andres Fm. Sample SAR 12-2. 8. Subaxial section. San Andres Fm. Sample SAR 13-1. 9-12. Globivalvulina retroseptata n. sp. 9. Oblique subaxial section. San Andres Fm. Sample SAR 13a-7. 10. Subaxial section. San Andres Fm. Sample SAR 13a-1. 11. Oblique subaxial section. San Andres Fm. Sample SAR 13a-8. 12. Axial section. San Andres Fm. Sample SAR 13a-9. Scale bars equal 0.10 mm.

FIGURE 32. 1-3, 6. Globivalvulina retroseptata n. sp. 1. Subaxial section. San Andres Fm. Sample SAR 13a-10. 2. Oblique subaxial section. San Andres Fm. Sample SAR 13a-11. 3. Subaxial section. San Andres Fm. Sample SAR 20-6. 6. Axial section. San Andres Fm. Sample SAR (A-1§). 4-5. Globivalvulina novamexicana n. sp. 4. Oblique subaxial section. San Andres Fm. Sample SAR 20-8. 5. Subaxial section. San Andres Fm. Sample SAR 20-9. 7-10. Hemigordius? sp. Four sections. San Andres Fm Sample SAR 1. Scale bars equal 0.10 mm.