Scaphisoma carinatum, Löbl, 2018

Löbl, Ivan, 2018, A review of Scaphisomatini from Sulawesi, with descriptions of ten new species (Coleoptera: Staphylinidae: Scaphidiinae), Acta Entomologica Musei Nationalis Pragae 58 (1), pp. 151-165 : 160-162

Scaphisoma carinatum sp. nov.

( Figs 40–46 )

Type locality. Indonesia, South Sulawesi, Maros District,Taman Nasional Bantimurung Bulusaraung [National Park], ca. 05°01.0 ′ S, 119°41.1 ′ E.

Type material. HOLOTYPE: ( NMPC), INDONESIA, S SULAWESI: Maros Distr. / Bantimurung Bulusaraung NP / karst; lowland tropical forest / 05°01.0 ′ S, 119°41.1 ′ E, 140 m / J. Hájek & J. Šumpich leg., 15.ii.2015.

Description. Length 2.24 mm, width 1.58 mm. Head, pronotum an hypomera ochreous, elytra on overwhelming surface reddish-brown, yellowish on area entirely covering apical fourth and extending somewhat onto posterior part of middle third; ventral side of mesothorax and metathorax reddish-brown, darker than hypomera, abdominal ventrites I–IV lighter than metaventrite, following ventrites yellowish, appendages ochreous to yellowish. Antennae long, length/width ratios of antennomeres as: III 15/10: IV 55/8: V 75/9: VI 64/9: VII 64/15: VIII 57/9: IX 73/13: X 67/12: XI 75/13. Pronotum and elytra not microsculptured. Pronotum strongly narrowed anteriad, with regularly rounded lateral margins, lateral margins carinae entirely visible; punctation on and around pronotal lobe fine, dense, visible at magnification 20×, consisting of well delimited punctures smaller than puncture intervals, punctures on remaining discal surface very fine, hardly visible at 20× magnification, dense and very shallow, punctation along lateral carinae distinct, dense. Tip of scutellum exposed. Elytra weakly narrowed apically, lateral margins almost regularly rounded, lateral margin carinae exposed throughout in dorsal view, apical margin weakly rounded, inner apical angles situated posterior level of outer angles, apical serration distinct, sutural margin not raised, adsutural areas flat, narrow, each with single puncture row. Sutural striae shortened, starting posterior level of scutellar tip, parallel in anterior two thirds, converging apically. Punctures along lateral margins coarse and dense. Elytral disc with punctures much larger than those on pronotal disc, sharply delimited, puncture intervals mostly about as large to two times as large as puncture diameters. Hind wings fully developed. Hypomera smooth, not microsculptured. Mesepimera as long as intervals between them and mesocoxae and five times as long as wide. Metaventrite with strigulate microsculpture; centre of metaventrite convex, with two shallow impressions near apical process and with mesal carina; punctation dense and coarse in impressed area, very fine and sparse on remaining surface; apical intercoxal margin concave. Lateral parts of metaventrite sparsely and very finely punctate; antecoxal punctures rows present in impressed striae. Submesocoxal lines convex, distinctly punctate; submesocoxal areas 0.05 mm, about as long as fourth of shortest intervals between them and metacoxae. Metanepisterna somewhat convex, slightly narrowed anteriad, with rounded angles. Protibiae straight. Mesotibiae and metatibiae bent. Abdomen with strigulate microsculpture and extremely fine punctation; ventrite I with submetacoxal lines convex, finely punctate; submetacoxal areas 0.07 mm, as long as third of shortest intervals between them and apical margin of ventrite.

Male characters. Prolegs and mid-legs with tarsomeres I strongly enlarged, almost as wide as apices of tibiae, tarsomeres II narrower but distinctly widened, tarsomeres III slightly widened.Abdominal ventrite VI extended by broad apical lobe. Tergite IX with one macroseta on each plate, stalk evenly narrow, broadly rounded ( Fig. 40 ). Aedeagus ( Figs 41–46 ) 1.50 mm long, asymmetrical. Basal bulb weakly sclerotized, not overlapping apical process, with large mesal carina.Apical process of median lobe strongly sclerotized, with dorsal branch triangular in dorsal view, almost obsolete in lateral view. Ventral branch large, with curved and blunt tip in dorsal view, straight and tapering in lateral view, bearing ventral tubercles near tip. Parameres enlarged and overlapping. Internal sac complex, with curved basal tooth-like sclerite, two long denticles, and

dense spine-like structures filling tubular apical section.

Female. Unknown.

Differential diagnosis. This species is a member of the S. tricolor species group as defined in LÖBL & OGAWA (2016). It is similar to the Philippine S. tricolor Heller, 1917 by its external characters, in particular the colour pattern, but may be distinguished by the metaventrite with a mesal carina and by the ventral branch of the apical process bearing subapical tubercles.

Etymology. The species epithet is a Latin adjective carinatus, - a, - um, referring to the carinate metaventrite.

Distribution. Indonesia, South Sulawesi.

Figures

Figs 35–49. 35–39 – Scaphisoma versicoloreum sp. nov.: 35, 36 – aedeagus in dorsal and lateral views; 37 – internal sac of aedeagus; 38 – male tergite 9; 39 – gonocoxite.40–46 – S. carinatum sp. nov.: 40 – male tergite 9; 41 – apical process of aedeagal median lobe in lateral view; 42 – tip of apical process in ventral view; 43, 44 – aedeagus in dorsal and lateral views; 45 – parameres in ventral view; 46 –internal sac of aedeagus. 47–49 – Scaphobaeocera jirkai sp. nov.: 47, 48 – aedeagus in dorsal and lateral views; 49 – gonocoxite. Scale bars: 0.1 mm (Figs 37, 39, 41–42, 46–49); 0.2 mm (Figs 35–36, 43–45); 0.3 mm (Figs 38, 40).