Diplomoela James, Bartz & Brown, 2023

James, Samuel W., Bartz, Marie L. C. & Brown, George G., 2023, New Ocnerodrilidae genera, species and records from Brazil (Annelida: Crassiclitellata), Zootaxa 5255 (1), pp. 235-269 : 264

Diplomoela James, Bartz & Brown , gen. nov.

Type species: Diplomoela browni sp. nov.

Etymology. The genus name is derived from the double gizzard, using the Portuguese word for gizzard, “moela.”

Diagnosis: Setae lumbricine, dorsal pores present or not from 14/15, male pores on xviii, acanthodriline male field with prostatic pores in xvii, xix; female pores paired in xiv, spermathecal pores paired intrasegmental in viii, ix; first setae behind viii. Male sexual system proandric; seminal vesicles multilobed, post-septal in xi; prostates in xvii, xix, ducts muscular. Spermathecae adiverticulate, ectal bulbous muscular duct; ovaries free in xiii. Gizzards large in viii, ix; paired calciferous glands in ix with parallel tubular structure without central lumen; intestinal origin in xiv, no typhlosole; numerous meronephridia commence before gonadal segments.

Remarks: Diplomoela has dorsal pores, two large gizzards in the space of viii and ix, and it is meronephric, with 40 to 80 nephridia per segment in mid body. This combination of characters is unique in the family, and it is the only meronephric genus in the family.