Novakia Strobl

Kerr, Peter H., 2007, Revision of the Holarctic genus Novakia Strobl (Diptera: Mycetophilidae), Zootaxa 1554, pp. 27-40 : 28-29

Novakia Strobl

Novakia Strobl 1893:162 .

Type species. Novakia scatopsiformis Strobl 1893:162 , by monotypy.

Diagnosis.

Three ocelli, arranged in lateral line. Frons setose or bare on upper half. Antennae separated or touching at their base; length of antennae considerably shorter in female than in male. Mesopleuron, laterotergite and mediotergite all bare. Wing membrane with small, irregularly arranged trichia only, all wing veins with setae or wing veins mostly bare on the dorsal surface. Costa produced beyond tip of R5. Subcosta bare, ending free. Wing veins sc-r, Rs, and R4 absent; r-m longer than R1; R1 and R5 touch or fuse for short distance at their base. Point of furcation of cubital fork before level of point of furcation of media; both cubital branches straight or nearly so. Female tergites 9 and 10 bearing distinctive fringe of setae on posterior margin, setae of tergite 10 light-colored.

Description.

Body length 1.6–2.2 mm. Wing length 1.6–2.3 mm.

Coloration. Head dark brown to black. Eye color brown when preserved fresh in alcohol, typically eyes gray when dried. Antenna brown to dark brown. Scutum and scutellum dark brown to black. Thoracic sclerites brown to black. Coxae, femora, tibiae, pale brown to black or whitish to pale yellow brown as in N. lisae sp. n.; tibial spurs yellowish brown to brown; tarsi pale brown to black. Wing hyaline or with brownish tint over most of surface, without markings; haltere stem and knob pale to dark brown. Abdominal segments brown to black. Terminalia brown to dark brown in male; in female, cercus pale yellowish to pale yellowish brown.

Head. Flattened anteriorly, surface often somewhat shiny, with short microtrichia. Three ocelli, arranged in lateral line, relative size of ocelli variable, distance of lateral ocellus to eye margin variable. Occiput rounded. Postocular region and top of head with evenly distributed, short brown setae; frons with short setae or bare. Frontal furrow present, frontal tubercle simple and pointed, positioned between or above antennal bases; face with short setae; transverse frontoclypeal membrane developed or incomplete; clypeus with short setae. Labellum appressed against head, with short dark setae. Lacinia short, inconspicuous. Palp consisting of five palpomeres, about as long or longer than head, sensilla cochleariformis of palpomere 3 small and inconspicuous. Antennal length in male 2–3.5 times the length of head; in female, antennal length 0.8–2.2 times the length of head; antennal bases separated by approximately 1/3 width of scape or touching; scape wider than long; pedicel wider than long; 14 flagellomeres; length of flagellum considerably shorter in female than in male, terminal flagellomere longer than wide.

Thorax. Surface somewhat shiny with short pubescence throughout. Scutum with short, dark, appressed, evenly distributed setae; lateral prealar setae thicker and longer. Antepronotum and proepisternum setose; other lateral thoracic sclerites bare. Posterior margin of scutellum bearing two pairs of long black setae. Mediotergite bare. Setae of legs short, variable; fore tibia with ovate anteroapical depressed area bearing several rows of small setae. Tarsal claws may be modified into comb-like form; empodium pulvilliform. Wing membrane clear (or mostly brown tinted in N. lisae ), with small, irregularly arranged trichia over cell membrane surface; costal vein extends approximately 0.3–0.5 of distance between R5 and M1; subcostal vein short, thick, bare, and ending free; radial veins with setulae on upper surface; vein R1 shorter or longer than half length of r-m; crossvein r-m close to R along length, touching or becoming fused for short distance before R1 arises; Rs absent; stem of medial fork very faint to mostly absent or readily apparent; length of medial fork relative to medial stem variable; medial veins setulose on upper surface or mostly bare; cubitus and its anterior branches (CuA1 and CuA2) with setulae on upper surface or bare in; anal vein short, setulose or bare on upper surface.

Male Terminalia. Tergite 8 reduced and terminalia rotated so that dorsal surface of terminalia is directed anteriorly, appressed against abdomen, and ventral surface is directed posteriorly. Epandrium (= tergite 9) posterior margin emarginate or not. Gonocoxites with lateral lobes bearing gonostyli; gonocoxal lobes ( Søli, 1997) separated widely or by small notch medially; gonostylus of variable form, sometimes bearing thin, elongate internal lobe.

Female Terminalia. Tergite 8 incomplete dorsally, split into two lateral sclerites; tergite 9 entire, posterior margin with row of black setae, of two lengths, interspersed; tergite 10 entire, with fringe of long light-colored setae; cerci two-segmented; two spermathecae.