Dziriblatta (Blattantis) montana Chopard, 1936b, 2019

Bohn, Horst, 2019, Revision of the genus Dziriblatta Chopard, 1936 (Blattodea, Ectobiidae, Ectobiinae) from North Africa, Spain, and the Macaronesian islands. I. The nine subgenera of the genus, Zootaxa 4610 (1), pp. 1-73 : 17-18

Dziriblatta (Blattantis) montana Chopard, 1936b , comb. nov.

Figs. 5 A–R, 6A–K, 22A, 23H, K, 25H, 26, 29.

Dziriblatta montana Chopard, 1936b: 157 , fig. 1; Chopard 1939: 107.

Lobolampra montana: Chopard 1943:34 .

Phyllodromica (Lobolampra) montana: Princis 1971: 1116 .

Material studied.

Type material. Morocco . Type, 1♀, Tachdirt & Djebel Likoumt, Grand-Atlas, Miss. Le Cerf & Talbot, Grand- Atlas, 28.IV.–9.VI.1927 (completely on two slides: Bo 231) [Ma 5]. ( MNHN) .

Additional material. Morocco. 1♀, Dj. Toubkal , Oued Imimène , 2800 m, 15.–31. VIII. (completely on two slides: Bo 235) [Ma 6]. ( MNHN).— Haut Atlas, Tizi-n-Test—TV station, 2100–2400 m, leg. B. & H.Bohn : 9♀; ex L: 6♂, 4♀, 24.VIII.1984 (slides: ♂, Ma 18/10,13,14,17,22; ♀, Ma 18/11,12,15,16); 14♂, 27♀, 4L, 16.V.1997 (slides: ♂, Ma 18a/1,2,4–6; ♀, Ma 18a/3); 2L, 7.X.2000 (Ma 18b); Haut Atlas, Pays Glaoua, E slope of Jbel Ifirouane (S Âït Barka, ca. 70 km ESE Marrakech), 1500–2200 m , 16♂, 41♀, 2O, 27.V.1989, leg. B. & H.Bohn (slides: ♂, Ma 86/3,6,8,9; ♀, Ma 86/10) ; 13♂, 2♀, 20L, Haut Atlas , Pays Glaoua, E slope of Jbel Ifirouane (S Âït Barka, ca. 70 km ESE Marrakech), 1500–2000 m, 20.II.1998, leg. B. & H.Bohn (Ma 86a) ; 4♂, 10♀, Haut Atlas , W above Zinit (N Tizi-n’Tabgourt, SE Imi-n-Tanoute), 1700–1900 m, 11./ 12.V.1997, leg. B. & H.Bohn (slides: ♂: Ma 145/2–4; ♀, Ma 145/5) ; 2♂, 14♀, 2O, Haut Atlas , W above Anchkrir (S Tizi-n’Tabgourt, SE Imi-n-Tanoute), 2100 m, 12.V.1997, leg. B. & H.Bohn (slides: ♂, Ma 146/1,2) ; 1♂, 1♀, Haut Atlas , Islane (near Tasguinnt, NNE Taroudannt), 1900–2100 m, 14.V.1997, leg. B. & H.Bohn (slide: ♂, Ma 148/2) ; Haut Atlas , Ruines Iguer (ca. 8 km NE Tizi-n-Test), 2050 m, leg. B. & H.Bohn : 1♂, 11♀, 16.V.1997 (slides: ♂, Ma 152/1; ♀, Ma 152/2–4); 1L, 7.X.2000 (Ma 152a); Haut Atlas, Tizi-n-Oulaoune (S Ijoukak), 2100 m, leg. B. & H.Bohn : 1♂, 16♀; ex L: 1♀, 2L, 17.V.1997 (slide: ♂, Ma 153/1); 1♀, 10.VIII.1999 (Ma 153a); 1♀, 1L, 6.X.2000 (Ma 153b); Haut Atlas, Tizi-n- Tamatert (E Imlil), 2150–2250 m, leg. B. & H.Bohn: 7♂, 32♀, 1L, 1O, 18.V.1997 (slides: ♂, Ma 156/1–3,5,6,9,10; ♀, Ma 156/4,7,8); 2L, 11.VIII.1999 (Ma 156a); 1L, 6.X.2000 (Ma 156b) ; 1♀, Haut Atlas , ca. 10 km SE Tahanaoute (S Marrakech), 1400 m, 18.V.1997, leg. B. & H.Bohn (Ma 157) ; 1♀, Haut Atlas , ca. 10 km SE Tahanaoute (S Marrakech), 1400 m, 11.VIII.1999, leg. H.Bohn (Ma 157a) ; 5♀, 1L, Haut Atlas , ca. 15 km SE Tahanaoute (S Marrakech), 1650 m, 18.V.1997, leg. B. & H.Bohn (Ma 158) ; 2♂, 6♀, Haut Atlas , near Tadmamt (ca. 25 km SE Tahanaoute), 1750–1950 m, 18./ 19.V.1997, leg. B. & H.Bohn (slides: ♂, Ma 159/2,3) ; Haut Atlas , 2 km NE Oukaïmeden, 2350 m, leg. B. & H.Bohn : 8♂, 26♀, 2L, 19.V.1997 (slide: ♂, Ma 160/1); 6♂, 10♀, 3L, 20.II.1998 (slide: Ma 160a/6); 2♀, 1O, Haut Atlas , Tizi-n’Outfi (ca. 45 km S Demnate), 2100 m, 20.V.1997, leg. B. & H.Bohn (Ma 162) . (C. Bohn, ZSM).

Description. Size. Length of pronotum in male 1.38–1.50 (mean 1.45) mm, in female 1.79–2.11 (mean 1.97) mm.

Male structures. See under characters of the subgenus.

Colouration. Tegmina transparent with a very characteristic colour pattern: dark patches arranged in longitudinal stripes ( Figs. 5J, K ). Body mainly dark ( Figs. 5 A–I, N), with some lighter coloured areas: A narrow interocular stripe on the head, thoracal nota with broad transparent margins, legs for most part ochre, narrow whitish stripe along the lateral and posterior borders of tergites and the lateral borders of sternites. Lightly coloured areas often further extended, especially in females ( Fig. 5O, P ). Disk of pronotum often reddish ( Fig. 5O ), sometimes of all nota.

Distribution. In the center of the High Atlas, between Taroudannt in the West and Demnate in the East, at altitudes between 1400 and 2800 m (Figs. 28, 29).

Remarks. Two of the male characters show locality dependent variation: The size of the glandular pores on T2 ( Figs. 6 G–I) and the presence ( Figs. 6D, E ) or absence ( Figs. 6A, B ) of a row of strong bristles on the preglandular margin of T7 near the opening of the pit. The distribution of the characters is shown on Table 1 . Large glandular pores are usually combined with the absence of large bristles, and small pores with the presence of larger bristles. But the occurrence of intermediate expressions in both characters ( Figs. 6F, C and 6H , respectively) does not allow a clear limitation between the different populations; a further subdivision of the taxon, therefore, appears inappropriate.

Figures

Fig. 5A–R. Outline of the structures of Blattantis montana. Male: A–J, L–N. A: T2; B: T3; C: T4; D: T5; E: T6; F: T7 with gland; G: T8; H: T9; I: T10, ap medio-anterior arm of left paraproct; J: left tegmen; L: hook; M: S9, three scale sclerites present (nos.1,2,4), cs cleft sclerite, R'3 sclerite; N: thoracal nota. Female: K, O–R. K: left tegmen; O: thoracal nota; P: T5; Q: dorsal genital complex, bd, bv dorsal, ventral basivalular sclerites, sp spermathecal plate; R: ventral genital complex, ls laterosternal shelf. – Enlargements: Same scale for A–K, for L,M, for O,P, and for Q,R. – Identifications: A–H,L (Ma 156/2), I (Ma 86/6), J,K (Ma 156/3,4), M (Ma 146/2), N (Ma 160a/6), O (Ma 152/2), P (slide: Bo 235), Q,R (Ma 18/16,15).

Fig. 6A–K. Male structures of Blattantis montana. A–F, K: variation in the shape of the T7 gland, b bottom of glandular pit, gu gutter, p anterior pouch, pm preglandular margin of 7, arrows in D enclose opening of the pit, arrowhead points to a large bristle, white arrow in K (same specimen as in E, but differently focused) points to the posterior pouch; G–I: variation in size and number of glandular pores on T2; J: SEM picture of forceps bristles from the glandular pit. – Enlargements: Same scale for A–I,K. – Identifications: A,G (Ma 156/2), B (Ma 156/10), C (Ma 159/3), D (Ma 18/6), E,H,K (Ma 145/3), F (Ma 86/3), I (Ma 18/22), J (Ma 86/9).