Kokiriidae, AG McFarlane, 1964

Johanson, Kjell Arne, 2003, Revision of the New Caledonian genus Mecynostomella (Trichoptera, Kokiriidae), Zootaxa 270 (1), pp. 1-24 : 3-4

Key to male Kokiriidae

1 Forewing Dc length less than 1/6x wing length ................................................ Kokiria (monotypic genus, including the New Zealand K. miharo McFarlane, 1964 )

­ Forewing Dc longer than 1/5x wing length.................................................................. 2

2 Spur formula 1,4,4 .................................................................................... Taskiropsyche (monotypic genus, including the Tasmanian T. lacustris Neboiss, 1977 )

­ Spur formula 2,4,4 ....................................................................................................... 3

3 Hind wing M includes 2 distal veins ............................................... Mecynostomella 6

­ Hind wing M includes 1 distal vein ........................................................................... 4

4 Forewing Tc cell about as long as Dc ...................................................... Tanjistomella (monotypic genus, including the SE Australian T. verna Neboiss, 1974 )

­ Forewing Tc longer than Dc ........................................................................................ 5

5 Maxillary palp with 3rd segment shorter than 2nd segment; in genitalia, IXth segment produced above Segment X .......................................................................... Pangullia (monotypic genus, including the Chilean P. faziana Navas, 1934 )

­ Maxillary palp with 3rd segment longer than 2nd segment; in genitalia, IXth segment not produced dorsally ................................................................................. Taskiria 12

6 In genitalia, Segment IX with posteroventral margin produced posteriorly ( Fig. 17 ); Segment IX produced dorsally above Segment X ( Fig. 15 ); lateral branch of Segment X larger than gonocoxite ( Fig. 15 ); central branch of Segment X, dorsal view, tapering apically ( Fig. 9 ); phallus with endothecae originating laterally, midway on phallobasis ( Figs 18, 19 ); phallobase with ventrodistal plate ( Figs 18, 19 ) ...................................... ..................................................................................... ... Mecynostomella spinosa sp.n.

­ In genitalia, Segment IX with posteroventral margin concave ( Figs 25 , 30 ); Xth segment not produced dorsally above Segment X; lateral branch of Segment X smaller than gonocoxite ( Figs 23 , 28 ); central branch of Segment X, dorsal view, bifurcated ( Figs 24 , 47 ); phallus with endothecae originating apicodorsally ( Figs 26 , 44 ), phallobase without additional plate ( Figs 26 , 49 ) .................................................................. 7

7 Forewing with pale or dark, oval area between Cu and A contrasting in color from the surrounding wing membrane ( Figs 10, 12 ) ................................................................. 8

­ Forewing without area contrasting in color ( Figs 8, 9 ) .............................................. 11

8 In genitalia, central branch of Segment X apically obtuse in lateral view ( Fig. 46 ); gonocoxite sub­straight in lateral view ( Fig. 46 ) ........................................................ 9

­ In genitalia, central branch of Segment X apically narrowing in lateral view ( Figs 36 , 41 ); gonocoxite apex produced dorsad ( Figs 36 , 41 ) and widely incised sub­apically .. ..................................................................................................................................... 10

9 In genitalia, lateral branch of Segment X originating at base of central branch ............ ........................................................................... Mecynostomella fusca Kimmins, 1953

­ In genitalia, lateral branch of Segment X originating midway on central branch ( Fig. 46 ) Mecynostomella flinti sp.n.

10 In genitalia, Segment IX with posteroventral margin deeply hyperbolically concave ( Fig. 38 ); lateral branch of Segment X long ( Fig. 36 )..................................................... ..................................................................................... Mecynostomella hollowayi sp.n.

­ In genitalia, Segment IX with posteroventral margin shallow trapezoidly concave ( Fig. 43 ); lateral branch of Segment X short ( Fig. 41 ) ............ Mecynostomella sigma sp.n.

11 In genitalia, posteroventral margin of segment IX shallowly concave ( Fig. 25 ); median branch located between lateral branch and gonocoxite ( Fig. 23 ); gonocoxite with dorsal margin weakly incised before apex ( Fig. 23 ); phallus with ventral spines on endothecae ( Fig. 26 ) .................................................................. Mecynostomella brevis sp.n.

­ In genitalia, posteroventral margin of segment IX deeply concave ( Fig. 30 ); median branch located behind lateral branch ( Fig. 28 ); gonocoxite with dorsal margin widely incised before apex ( Fig. 28 ); phallus without spines on endothecae ............................ ..................................................................................... Mecynostomella caledonia sp.n.

12 In genitalia, phallus short, thick, slightly curving ventrally; segment X with laterally oriented lateral branches; gonocoxite uniformly slender ............................................... ................................................................................. Taskiria otwayensis Neboiss, 1984

­ In genitalia, phallus long, slender, strongly curving ventrally; segment X without laterally oriented lateral branches; gonocoxite proximally thick ...................................... 13

13 In genitalia, lateral branches of segment X well separated from central branch in dorsal view; Xth segment apex curving ventrally .................. Taskiria austera Neboiss, 1977

­ In genitalia, lateral branches of segment X not separated from central branch in dorsal view; Xth segment apex oriented posteriorly along its length ....................................... .................................................................................. Taskiria mccubbini Neboiss, 1977

Figures

FIGURES 15­19. Mecynostomella spinosa sp.n., male holotype. 15, genitalia, lateral view; 16, genitalia, dorsal view; 17, genitalia, ventral view; 18, phallus, lateral view; 19, phallus, ventral view.

FIGURES 6­14. Mecynostomella species, right wings and wing details. 6, Mecynostomella spinosa sp.n. male, right wings and vein terminology; 7, Mecynostomella spinosa sp.n. male, scaloid seta from forewing Cu2, highly magnified; 8, Mecynostomella spinosa sp.n. female, right wings; 9, Mecynostomella brevis sp.n. male, right wings; 10, Mecynostomella hollowayi sp.n. male, right wings, a indicates the position of a pale area in the Cu­ and A­ area of the forewing, and made from minute structures shown in the next figure; 11, details of a single structure forming a pentagon and with a central opening, and which build up a pale area of the forewing as indicated in the previous figure; 12, Mecynostomella fusca Kimmins, 1953, right wings, with area covered by sensillae indicated by a surrounding black line; 13, Mecynostomella flinti sp.n., detail from wing membrane of dark area between forewing Cu and A, showing position of a modified sensilla and sensilla bases (t), dorsal view; 14, same species and sensilla, lateral view.

FIGURES 23­27. Mecynostomella brevis sp.n., male holotype. 23, genitalia, lateral view; 24, genitalia, dorsal view; 25, genitalia, ventral view; 26, phallus, lateral view; 27, phallus, ventral view.

FIGURES 28­32. Mecynostomella caledonia sp.n., male holotype. 28, genitalia, lateral view; 29, genitalia, dorsal view; 30, genitalia, ventral view; 31, phallus, lateral view; 32, phallus, ventral view.

FIGURES 46­50. Mecynostomella flinti sp.n., male holotype. 46, genitalia, lateral view; 47, genitalia, dorsal view; 48, genitalia, ventral view; 49, phallus, lateral view; 50, phallus, ventral view.

FIGURES 41­45. Mecynostomella sigma sp.n., male holotype. 41, genitalia, lateral view; 42, genitalia, dorsal view; 43, genitalia, ventral view; 44, phallus, lateral view; 45, phallus, ventral view.

FIGURES 36­40. Mecynostomella hollowayi sp.n., male holotype. 36, genitalia, lateral view; 37, genitalia, dorsal view; 38, genitalia, ventral view; 39, phallus, lateral view; 40, phallus, ventral view.