Mecynostomella fusca Kimmins, 1953

Johanson, Kjell Arne, 2003, Revision of the New Caledonian genus Mecynostomella (Trichoptera, Kokiriidae), Zootaxa 270 (1), pp. 1-24 : 21-22

Mecynostomella fusca Kimmins, 1953

( Figs 12 , 5­5 , 54 )

Material examined: Male holotype: N. New Caledonia, Mt. Tinchialit, 2020 ft., 22.viii­ 10.ix.1949, at light [L. E. Cheesman] ( BMNH, mounted on 3 slides) .

Diagnosis: In fusca , the forewings have a dark area densely covered by cone­shaped sensillae as in flinti . The species is easily separated from flinti and congeners by the presence of long lateral branches originating below segment X. The genitalia are unique in the undulated posterior margin of segment IX, a short segment IX, presence of lateral branches below segment X, and apex of segment X that in lateral view is wider than in flinti .

Description of the male and female is given in Kimmins (1953: 255­257). To the original description the following head characters are added: Tentorium ( Fig. 2 ) with long anterior tentorium arms bearing pointed vestigial projections at dorsal margin; tentorial bridge broad, straight, with long, pointed anteromesal protuberance. Maxillary palp ( Fig. 3 ) slightly shorter than labial palp ( Fig. 4 ). Membrane between palp segments densely covered by dark microtrichiae ( Fig. 5 ) also present on the segments. Wings ( Fig. 12 ) with dark area in anal and cubital region made from densely grouped, dark sensillae. Genitalia ( Figs 51­52 ) as in Kimmins (1953) but in lateral view with more setae along posterior margin of segment IX, and gonocoxite with dorsal setae ( Fig. 51 ). Phallus ( Fig. 52 ) similar to that of flinti, except anteriorly narrower and posteriorly wider.

Figures

FIGURES 6­14. Mecynostomella species, right wings and wing details. 6, Mecynostomella spinosa sp.n. male, right wings and vein terminology; 7, Mecynostomella spinosa sp.n. male, scaloid seta from forewing Cu2, highly magnified; 8, Mecynostomella spinosa sp.n. female, right wings; 9, Mecynostomella brevis sp.n. male, right wings; 10, Mecynostomella hollowayi sp.n. male, right wings, a indicates the position of a pale area in the Cu­ and A­ area of the forewing, and made from minute structures shown in the next figure; 11, details of a single structure forming a pentagon and with a central opening, and which build up a pale area of the forewing as indicated in the previous figure; 12, Mecynostomella fusca Kimmins, 1953, right wings, with area covered by sensillae indicated by a surrounding black line; 13, Mecynostomella flinti sp.n., detail from wing membrane of dark area between forewing Cu and A, showing position of a modified sensilla and sensilla bases (t), dorsal view; 14, same species and sensilla, lateral view.

FIGURES 1­5. Head characters of Mecynostomella species. 1. Mecynostomella spinosa sp.n. male, head, lateral view; 2, Mecynostomella fusca Kimmins, 1953, tentorium, frontal view; 3, same species, maxillary palp; 4, same species, labial palp; 5, same species, membranous area between labial palp segments 2 and 3, densely covered by microtrichiae.

FIGURES 53­54. Distribution maps of New Caledonian Mecynostomella. 53, caledonia (circles), hollowayi (rectangles), and sigma (triangle); 54, spinosa (circles), flinti (rectangle), brevis (triangles), and fusca (diamond).

FIGURES 51­52. Mecynostomella fusca Kimmins, 1953, male holotype. 51, genitalia, lateral view; 52, phallus, lateral view.