Glyphiulus semigranulatus, Golovatch & Geoffroy & Mauriès & Spiegel, 2007

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group, Zoosystema 29 (1), pp. 7-49 : 25-28

publication ID	https://doi.org/ 10.5281/zenodo.5392079
persistent identifier	https://treatment.plazi.org/id/03F4C378-A23D-305D-FD6A-0156FDCEFA37
treatment provided by	Marcus
scientific name	Glyphiulus semigranulatus
status	sp. nov.

Treatment

Glyphiulus semigranulatus View in CoL n. sp. ( Figs 13 View FIG ; 14 View FIG )

TYPE MATERIAL. — China. Yunnan Prov., Meele (= Mile) County, Bai Long Dong Cave (= White Dragon Cave),

2.I.1989, leg. P. Beron, holotype ♂ ( NMNHS) ; paratypes 1 ♂, 1 ♀ ( NMNHS) ; 1 ♀ ( SEM) ; 1 ♂ ( MNHN GA 037 About MNHN ). — Yunnan Prov., Jianshui County, Yan Zi Dong Cave , 10.I.1989, leg. P. Beron, paratypes 5 ♂♂, 3 ♀♀, 4 juv. ( NMNHS) ; 4 ♂♂, 2 ♀♀ ( MNHN GA 037 About MNHN ) ; 4 ♂♂, 2 ♀♀ ( ZMUM) ; 2♂♂, 2 ♀♀ ( ZMUC) ; 1 ♂, 1 ♀ ( MCSNV) ; 1 ♂ ( SEM) .

ETYMOLOGY. — To emphasize the obvious similarity to G. granulatus .

DIAGNOSIS. — Differs by a position somewhat intermediate between the most similar congeners G. granulatus and G. paragranulatus n. sp., i.e. coloration, carinotaxy,

antennae, claws, etc., as in G. granulatus , but size, segment shape, leg length and anterior gonopods more like in G. paragranulatus n. sp.

DESCRIPTION

Length of adults of both sexes 12-24 mm, width 0.8-1.2 mm.

Adults of both sexes with 31-64p+5-1a+T; largest juvenile ♂ with 28p+5a+T, largest juvenile ♀ with 27p+5a+T; two smallest juveniles with 15 or 18 haplopodous segments after 6th, plus 2a+T or 1a+T, respectively.Length of holotype about 22 mm, width 1.0 mm, with 45p+3a+T.

Coloration, carinotaxy ( Fig. 13 View FIG A-E), antennae ( Figs 13A View FIG ; 14A View FIG ), gnathochilarium, claws, etc. ( Fig. 14 View FIG B-D, F, G) basically as in G. granulatus , but body size (usually much larger), segment shape (slightly compressed laterally) ( Fig. 13C View FIG ), telson ( Fig. 13E, F View FIG ) with peculiar flaps on paraprocts near hypoproct ( Fig. 13F View FIG ), legs (somewhat longer) and anterior gonopods ( Fig. 14E View FIG ) more like those of G. paragranulatus n. sp. Adults with 11-15 ocelli.

REMARKS

In the absence of any evident troglomorphic traits, this species can only be considered as troglophilic at most. This is supported by G. semigranulatus n. sp. being present in several relatively strongly isolated caves, sometimes in significant numbers.