Glyphiulus bedosae, Golovatch & Geoffroy & Mauriès & Spiegel, 2007

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group, Zoosystema 29 (1), pp. 7-49 : 30-32

Glyphiulus bedosae n. sp.

( Figs 17 ; 18 )

TYPE MATERIAL. — Laos. Luang Prabang Prov., Nong Kiaw (= Muang Ngoy), Tham Pha Kouang Cave , 29.XII.2000, leg. A. Bedos & L. Deharveng (LAO-099), holotype ♂ ( MNHN GA 039 ) ; paratypes 1 ♂ ( MNHN GA 039 ), 1 ♀ ( SEM). ETYMOLOGY. — Honours Anne Bedos, one of the collectors .

DIAGNOSIS. — Differs from other species by the paraprocts showing a clear median marginal ridge, combined with a plumose spatuliform distal process of the posterior gonopods and a subunciform distomedian corner of the anterior gonopods.

DESCRIPTION

Length of adults of both sexes 16-20 mm, width 1.0- 1.1 mm.

Holotype 16 mm long, 1.0 mm wide, with 40p+3a+T and 11 ocelli; paratype ♂ about 20 mm long, 1.1 mm wide, with 54p+2a+T and 14 ocelli.

Coloration, antennae ( Figs 17A ; 18A ), gnathochilarium (mentum divided, n = 1) ( Fig. 18B ), carinotaxy ( Fig. 17 A-E), segment shape, legs ( Fig. 18 C-F), etc., generally as in G. granulatus , but paraprocts with a low, but evident, median marginal ridge ( Fig. 17F ).

Anterior gonopods like those of G. semigranulatus n. sp. ( Fig. 18G ), but posterior gonopods with a plumose spatuliform distal process ( Fig. 18H ), i.e. much as in G. quadrohamatus or G. lipsorum .

REMARKS

In the absence of any troglomorphic traits, this species can only be considered troglophilic at most.

Figures

FIG. 17. — Glyphiulus bedosae n. sp., ♀ paratype: A, anterior part of body, lateral view; B, same, dorsal view; C, middle part of body, lateral view; D, same, dorsal view; E, caudal part of body, lateral view; F, telson, ventral view. Scale bars: A, C, 0.5 mm; B, D, E, 0.2 mm; F, 0.1 mm.

FIG. 18. — Glyphiulus bedosae n.sp., ♂ paratype:A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view; E, leg 3; F, midbody leg; G, anterior gonopods, caudal view; H, posterior gonopod, subcaudal view. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm.