Glyphiulus, GERVAIS, 1847 OF THE

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group, Zoosystema 29 (1), pp. 7-49 : 42

publication ID

https://doi.org/ 10.5281/zenodo.5392079

persistent identifier

https://treatment.plazi.org/id/03F4C378-A20C-306C-FE97-0785FB88FE9C

treatment provided by

Marcus

scientific name

Glyphiulus
status

 

KEY TO SPECIES OF GLYPHIULUS GERVAIS, 1847 OF THE View in CoL GRANULATUS- GROUP

1. Carinotaxy patterns of collum and body segments typical, all crests well-developed: carinotaxy formula of collum: I-VI+7a+pc+ma+pc+7a+VI-I, i.e. middle part of collum with two abbreviated paramedian crests over front 2/3 collum, as well as three abbreviated crests medially in caudal 1/3 collum ( Figs 2A View FIG ; 10B View FIG ; 13B View FIG ; 15B View FIG ; 17B View FIG ); carinotaxy of midbody segments 3(2)+I+4+I+3(2) and 3(2)+i+3+i+3(2) ( Figs 2 View FIG B-F; 3B; 7C-F; 10C, D; 13C, D; 17C, D; 19A-C, E; 21; 22D; 27D), rarely even with a third row of 3(2)+i+3+i+3(2) ( Fig. 15C, D View FIG ), but median crest always doubled anteriorly. Relatively small species:length of adults ≤ 25 mm, width ≤ 1.3 mm ........................................................................... 2

— Carinotaxy patterns of collum and body segments deviating from the typical ones, i.e. crests on collum either at least partly obliterated ( Figs 7B View FIG ; 19A, B View FIG ; 21B View FIG ; 24A, B View FIG ; 27A, B View FIG ) or, more seldom, showing a different formula, or some crests hypertrophied and differentiated, whereas midbody segments with either a typical (2+I+4+I+2 and 2+i+3+i+2) or an atypical formula, e.g., 3(2)+I+3+I+3(2) ( Fig. 24 View FIG ). Often larger species .............. 7

2. Midbody segments in cross-section slightly compressed laterally ( Fig. 13C View FIG ). Legs slightly elongate, about as long as midbody height. Yunnan Prov., China............................... 3

— Midbody segments round in cross-section ( Figs 3B View FIG ; 19E View FIG ; 22D View FIG ; 25B View FIG ), not compressed laterally. Legs usually slightly shorter, rarely evidently longer, than midbody height .... 4

3. Antennae elongate ( Fig. 12A View FIG ). Spine at base of claw more than half as long as claw ( Figs 11D View FIG ; 12D, E View FIG ) ................................................................. G. paragranulatus n. sp.

— Antennae stout ( Fig. 14A View FIG ). Spine at base of claw less than half as long as claw ( Fig. 14D View FIG ) ................................................................................................. G. semigranulatus n. sp.

4. Crests on midbody segments divided into three tubercles ( Fig. 15D, E View FIG ). Epiproct with a small but evident middorsal tubercle ( Fig. 15E View FIG ). Posterior gonopods with a plumose and subflagelliform distal process ( Fig. 16E, F View FIG ) ........................................... G. beroni n. sp.

— Crests on midbody segments divided into two tubercles ( Figs 2D View FIG ; 7C, D View FIG ; 17C, D View FIG ; 21C, D View FIG ). Epiproct without a middorsal tubercle ( Figs 2E View FIG ; 17E View FIG ). Posterior gonopods with a plumose but subspatuliform distal process ( Fig. 18H View FIG ) ................................................ 5

5. Paraprocts evidently elevated, ridge-like along midline ( Fig. 17F View FIG ). Anterior gonopods with thicker telopodites, distomedial corners subunciform ( Fig. 18G View FIG ). Northern Laos.......... ........................................................................................................... G. bedosae n. sp.

— Paraprocts virtually flat and smooth at midline ( Figs 3C, D View FIG ; 8B View FIG ; 25D View FIG ). Anterior gonopods with telopodites more slender, distomedial corners different ....................................... 6

6. Legs evidently shorter than midbody height ( Fig. 3B View FIG ). Antennae short and stout ( Figs 3A View FIG ; 4A View FIG ). Anterior gonopods with characteristic, very high, distomedial processes ( Figs 3E View FIG ; 5A, B View FIG ; 6C View FIG ). Pantropical ........................................................................... G. granulatus

— Legs only slightly shorter than midbody height. Antennae slightly longer and more slender (more like in Figure 26A View FIG ). Anterior gonopods much like in Figure 18G View FIG , distomedial processes low and broadly rounded. Hubei Prov., China............................ G. lipsorum

7. Collum conspicuously hypertrophied due to crests I-IV, but paramedian crests complete and contrastingly low in anterior 1/3 collum. Vestigial 2-segmented leg remains on male sternum 1. Anterior gonopods much as in G. granulatus ( Fig. 5A, B View FIG ), distomedial processes similarly high but much wider. Posterior gonopods strongly constricted in distal 1/3. Mountains of central Vietnam.................................................................. G. capucinus

— Collum not hypertrophied, often some of its crests more or less obliterated. Male legs 1 either completely reduced to 1-segmented tubercular (usually) or leg-like. Anterior gonopods usually with much lower distomedial processes, if any. Posterior gonopods not so constricted in distal part ( Fig. 7F, G View FIG ) ...................................................................... 8

Darwin Core Archive (for parent article) View in SIBiLS Plain XML RDF