Dichotomius (Selenocopris) favi Kohlmann and Solís, 1997

Valois, Marcely C., Silva, Fernando A. B. & Vaz-de-Mello, Fernando Z., 2022, A taxonomic revision of the globulus species group of Dichotomius Hope, 1838 (Coleoptera: Scarabaeidae: Scarabaeinae), Journal of Natural History 56 (1 - 4), pp. 119-147 : 132-133

Dichotomius (Selenocopris) favi Kohlmann and Solís, 1997

( Figures 2 (g–l), 6)

Dichotomius favi Kohlmann and Solís 1997: 345 , 363; Krajcik 2006: 53; 2012: 91; Ariás-Buriticá et al. 2011: 876; Solis and Kohlmann 2012: 6; Vaz-de-Mello and Nunes 2016: 299.

Diagnosis

Dichotomius favi is suggested to be closely related to D. danieli (see the diagnosis of D. danieli ). However, the following combination of characters can be used to distinguish D. favi from its relatives: surface of head with different-sized ocellate punctures, forming transverse rows at clypeal surface; elytral striae thin and deeply punctate, punctures spaced by their diameter; ventral surface of the mesofemora with large coarse punctures at apical third; and paramera rectangular, obliquely truncate at apex in dorsal view ( Figure 2 (j,k)).

Redescription (male)

Body length: 8.55 mm. Dorsum black, barely shiny on head and pronotum, elytra opaque ( Figure 2 (g)). Head: clypeal edge bidentate, teeth acute, separated by a deep V-shaped emargination. Clypeogenal junction rounded. Cephalic process produced into a small central tubercle ( Figure 2 (g)). Antenal club with light brown tumescence. Thorax: pronotum approximately twice wider than long ( Figure 2 (g)). Anteromedian region of the disc lacking knobs, horns or excavations. Surface with dense ocellate punctures, equally spaced. Mid-longitudinal sulcus feebly marked medially. Surface of hypomera covered with dense ocellate punctures, spaced by less than their diameter; some punctures bearing orange setae. Prosternum covered with ocellate punctures. Mesoventrite covered with ocellate punctures, usually bearing short orange setae. Metaventrite surface with ocellate punctures, except on disc; punctures on anterior region usually bearing short setae. Meso-metaventral suture distinct. Mid-longitudinal sulcus of metaventrite feebly marked posteriorly. Lateral lobes of metaventrite with ocellate punctures larger than those on anterior region. Elytra: slightly convex in lateral view ( Figure 2 (g)). Humeral callus distinct. Interstriae I–VII slight convex, barely shiny, with shagreened microsculpture; medial region densely covered with smooth dots. Legs: protibial spur curved at apical fifth. Metatibial spur deeply emarginate apically. Ventral surface of the profemora covered with coarse setiferous punctures along its entire extension. Ventral surface of the metafemora with ocellate punctures concentrated on apical half; basal half smooth. Meso and metafemora bearing a thin longitudinal groove along its entire extension, wider on apical half. Abdomen: ventrites covered with ocellate punctures in lateral regions; punctures in medial region in a single row along anterior edge. Sixth ventrite 3 times longer than fifth. Pygidium twice wider than long; disc strongly convex apically. Surface of pygidial disc with ocellate punctures concentrated along anterior half, and spaced by half their diameter. Apex of the pygidium margined. Tegmen: in dorsal view, paramera rectangular, one-third longer than its width medially ( Figure 2 (j,k)); apex slightly obliquely truncate; inner edges close to each other at basal fifth ( Figure 2 (j,k)). Dorsal surface of paramera smooth ( Figure 2 (j)). Apical half of paramera noticeably narrower than base in lateral view ( Figure 2 (l)). Ventral sclerotised membrane between paramera thinner and curved at basal half; angulated laterally ( Figure 2 (k)). Female: differs from male by the clypeal teeth larger and acute. Clypeal edge with well defined transverse ridges. Sixth abdominal ventrite 3 times longer than fifth. Disc of the pygidium swollen.

Type material examined

Holotype of Dichotomius favi ♂ [ MNCR]: ‘Sector Cerro Cocori. Fca./de E. Rojas, 150 m. Prov./ Limon, Costa Rica. Oct/1992, E. Rojas/L–N 286000. 567,500’ // ‘ Dichotomius favi / Kohlmann and Solís’ // ‘ COSTA RICA /INBIO [barcode] CRI000 931,796’ // ‘HOLOTYPO/ Dichotomius favi /Kohlmann y Solís’. Examined via photographs ( Figure 1 (i)).

Figures

Figure 1. Dorsal habitus of male. (a) Dichotomius benesi; (b) D. berthalutzae sp. nov.; (c) D. femoratus; (d) D. fortepunctatus; (e) D. gamboaensis.

Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels.