ELMIDAE, Curtis, 1830

Hubbard, Todd, 2017, Aquatic Coleoptera (Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae) of Iowa: Distributional Records and Notes, The Coleopterists Bulletin 71 (3), pp. 471-505 : 486-491

ELMIDAE

Ancyronyx variegatus (Germar) . Allamakee: Yellow R., Postville, CTH, 8/17/11; Black Hawk: Black Hawk Cr., Hudson, MWB, 8/8/02; Bremer: East Fk. Wapsipinicon R., Sweet Marsh SWMA, CTH, 8/25/14; Buchanan: Lime Cr., Brandon, DPH, 9/4/07; Butler: Beaver Cr., New Hartford, CTH, 8/14/03; Cedar: Rock Cr., Tipton, CTH, 7/30/01; Chickasaw: Crane Cr., Jerico, CTH, 9/7/94; Clayton: Volga R., Elkport, CTH, 10/20/00; Clinton: Barber Cr., Grand Mound, CTH, 9/9/98; Delaware: Plum Cr., Earlville, CTH, 8/16/06; Dubuque: North Fk. Maquoketa R., Dyersville, MWB, 7/25/05; Fayette: Little Turkey R., Gouldsburg County Park, CTH, 9/10/03; Floyd: Little Cedar R., Colwell County Park, MWB, 8/20/02; Grundy: South Beaver Cr., Parkersburg, MWB, 9/3/08; Howard: Crane Cr., Riceville, CTH, 7/10/06; Jackson: Prairie Cr., Maquoketa, CTH, 8/14/01;; Jones: Maquoketa R., Monticello, MWB, 9/6/ 05; Linn: Buffalo Cr., Central City, MWB, 9/2/ 08; Louisa: Iowa R., Columbus Junction, CTH, 9/10/08; Mitchell: Little Cedar R., New Haven, CTH, 8/17/05, Muscatine: Mud Cr., Wilton, DPH, 9/15/03; Scott: Lost Cr., Princeton, CTH, 8/30/04; Washington: Crooked Cr., Crawfordsville, CTH, 7/14/05. Distribution: Eastern USA, extending west to Kansas and Texas in the south. Iowa forms part of the western edge of its distribution with collections occurring in the eastern half of the state. New state record.

Dubiraphia bivittata (LeConte) . Clayton: Roberts Cr., Postville, CTH, 8/14/08; Dubuque: Little Maquoketa R., Dubuque, CTH, 8/19/08. Distribution: Eastern USA, extending west to North Dakota in the north, Missouri, and Iowa. Habitat preferences are coarse substrate, including wood in streams and rivers. New state record .

Dubiraphia minima Hilsenhoff. Allamakee : Yellow R., Ion, CTH, 9/14/04; Appanoose: Cooper Cr., Centerville, CTH, 8/11/05; Benton: Bear Cr., Shellsburg, CTH, 10/1/01; Black Hawk: Wapsipinicon R., Dunkerton, CTH, 8/2/ 06; Bremer: East Fk. Wapsipinicon R., Sweet Marsh SWMA, CTH, 8/23/11; Buchanan: Lime Cr., Brandon, CTH, 9/4/07; Buena Vista: Brooke Cr., Rembrandt, JML, 10/5/05; Butler: Dry Cr., Clarksville, CTH, 5/29/07; Carroll: Marrowbone Cr., Lanesboro, TMM, 9/13/07; Cass: Sevenmile Cr., Cumberland, TMM, 6/ 28/06; Cedar: Rock Cr., Tipton, CTH, 8/20/ 08; Cherokee: Silver Cr., Cherokee, KLS, 9/20/02; Chickasaw: Wapsipinicon R., New Hampton, CTH, 8/18/04; Clay: Stony Cr., Everly, TMM, 8/30/05; Clayton: Hewitt Cr., Volga, MWB, 9/1/04; Clinton: Elk R., Teeds Grove, CTH, 8/6/07; Dallas: Little Beaver Cr., Woodward, TMM, 7/24/07; Davis: Soap Cr., Drakesville, CTH, 8/16/04; Decatur: Long Cr., Van Wert, CTH, 9/14/95; Delaware: South Fk. Maquoketa R., Dundee, CTH, 7/12/05; Des Moines: Skunk R., Burlington, MWB, 9/7/06; Dickinson: Little Sioux R., Milford, CTH, 9/4/ 96; Dubuque: Catfish Cr., Dubuque, CTH, 8/1/ 07; Fayette: Turkey R., Eldorado, CTH, 10/1/02; Floyd: Little Cedar R., Colwell, CTH, 8/20/02; Franklin: Maynes Cr., Hampton, CTH, 7/18/95; Greene: West Buttrick Cr., Grand Junction, TMM, 9/4/08; Guthrie: Deer Cr., Stuart, CTH, 7/30/98; Hamilton: White Fox Cr., Webster City, CTH, 11/10/97; Hardin: South Fk. Iowa R., Iowa Falls, TMM, 8/1/07; Henry: Big Cr., New London, CTH, 8/30/05; Howard: Crane Cr., Riceville, CTH, 7/10/06; Ida: Battle Cr., Holstein, KLS, 6/29/04; Jackson: Prairie Cr., Maquoketa, CTH, 10/1/08; Jasper: North Skunk R., Newton, CTH, 8/8/01; Jefferson: Cedar Cr., Stockport, CTH, 7/20/05; Johnson: Iowa R., Iowa City, CTH, 8/8/05; Jones: Bear Cr., Wyoming, MWB, 7/13/04; Keokuk: Cedar Cr., Delta, CTH, 8/18/03; Kossuth: Black Cat Cr., Algona, MWB, 8/25/03; Linn: West Otter Cr., Toddville, CTH, 10/11/01; Louisa: Long Cr., Columbus Junction, CTH, 9/26/95; Lucas: Wolf Cr., Chariton, CTH, 8/10/05 Lyon: Mud Cr., Lester, JML, 7/21/04; Madison: North Branch North R., Winterset, TMM, 8/21/07; Mahaska: North Skunk R., Rose Hill, CTH, 8/ 14/02; Marion: English Cr., Knoxville, MWB, 7/ 18/05; Marshall: South Minerva Cr., St. Anthony, TMM, 8/21/08; Mitchell: Rock Cr., Rock Cr., CTH, 8/15/96; Montgomery: Unnamed trib. to Tarkio R., Stanton, TMM, 8/6/13; Muscatine: Pike Run, Nichols, CTH, 7/22/05; O’ Brien: Little Waterman Cr., Hartley, CTH, 7/9/96; Page: West Tarkio Cr., Essex, KLS, 7/7/04; Pocahontas: Big Cedar Cr., Fonda, KBK, 9/19/01; Polk: Four Mile Cr., Ankeny, TMM, 8/9/06; Poweshiek: Walnut Cr., Belle Plaine, CTH, 7/29/03; Ringgold: Lotts Cr., Lamoni, TMM, 7/18/03; Sac: Boyer R., Schaller, JML, 8/4/04; Scott: Lost Cr., Princeton, CTH, 8/30/04; Shelby: East Branch West Nishnabotna R., Avoca, TMM, 8/16/07; Sioux: Deep Cr., Granville, KLS, 9/26/ 06; Story: South Skunk R., Ames, CTH, 9/19/97; Tama: Richland Cr., Haven, CTH, 9/22/99; Van Buren: Fox R., Mt. Sterling, CTH, 7/19/05;

Washington: Crooked Cr., Crawfordsville, CTH, 7/14/05; Wayne: South Fk. Chariton R., Corydon, TMM, 9/12/12; Webster: Lyons Cr., Webster City, TMM, 9/16/08; Winnebago: Winnebago R., Lake Mills, CTH, 7/30/12; Winneshiek: Yellow R., Postville, CTH, 8/1/ 06; Woodbury: West Fk. Little Sioux R., Bronson, TMM, 8/9/05; Worth: Deer Cr., Carpenter, CTH, 8/21/01; Wright: Otter Cr., Goldfield, JML, 8/11/01. Distribution: Northeastern USA, with Iowa forming part of the western border of its range.

Dubiraphia quadrinotata (Say) . Allamakee: Yellow R., Waukon, CTH, 8/8/06; Benton: Mud Cr., Vinton, CTH, 8/29/05; Black Hawk: Wolf Cr., La Porte City, CTH, 8/19/04; Boone: Squaw Cr., Zenorsville, KBK, 7/14/00; Bremer: Quarter Section Run, Waverly, CTH, 8/9/04; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Buena Vista: Unnamed trib. to Maple Cr., Aurelia, TMM, 8/ 27/13; Butler: Boylan Cr., Aredale, MWB, 11/ 11/02; Calhoun: Unnamed trib. to Purgatory Cr., Lohrville, KLS, 8/14/07; Carroll: Marrowbone Cr., Lanesboro, TMM, 9/13/07; Cedar: Rock Cr., Tipton, CTH, 8/20/08; Cerro Gordo: Bailey Cr., Thornton, CTH, 7/23/03; Cherokee: Bear Cr., Cleghorn, CTH, 9/4/01; Chickasaw: East Fk. Wapsipinicon R., New Hampton, CTH, 10/2/ 95; Clay: Prairie Cr., Spencer, JML, 9/4/01; Clayton: Roberts Cr., Postville, CTH, 8/13/08; Clinton: Brophy Cr., McCausland, CTH, 10/7/04; Davis: Soap Cr., Peach Ave., CTH, 7/17/13; Delaware: Twin Springs Cr., Colesburg, CTH, 9/17/07; Des Moines: Brush Cr., Spring Grove, CTH, 9/28/00; Dickinson: Little Sioux R., Lake Park, TMM, 8/11/14; Dubuque: Lytle Cr., Zwingle, CTH, 8/16/99; Emmet: Ditch 151, Wallingford, TMM, 6/17/03; Fayette: Dibble Cr., Clermont, CTH, 8/5/04; Floyd: Cedar R., Charles City, MWB, 9/9/08; Franklin: Maynes Cr., Hampton, CTH, 8/23/01; Grundy: South Beaver Cr., Parkersburg, CTH, 7/30/96; Hamilton: Long Dick Cr., Ellsworth, TMM, 10/6/08; Hancock: Unnamed trib. to East Branch Iowa R., Goodell, TMM, 7/23/12; Hardin: Iowa R., Eldora, CTH, 8/16/05; Henry: Big Cedar Cr., Oakland Mills, CTH, 9/2/11; Howard: Crane Cr., Riceville, 8/ 16/10; Ida: Ashton Cr., Washta, JML, 7/6/06; Iowa: Deep Cr., Guernsey, CTH, 8/25/11; Jackson: Farmers Cr., Fulton, CTH, 8/8/07; Jasper: North Skunk R., Newton, CTH, 8/8/ 01; Johnson: Iowa R., Iowa City, CTH, 8/8/ 05; Jones: Maquoketa R., Monticello, CTH, 9/6/ 05; Keokuk: Cedar Cr., Delta, CTH, 8/18/03; Kossuth: Four Mile Cr., Algona, TMM, 7/11/06; Linn: Buffalo Cr., Central City, CTH, 9/2/08; Louisa: Buffington Cr., Columbus City, CTH, 8/23/10; Lyon: Little Rock R., George, CTH, 9/ 5/96; Marshall: Burnett Cr., Marshalltown, CTH, 8/22/00; Mitchell: Deer Cr., Carpenter, CTH, 8/25/08; Monona: Jordan Cr., Soldier, TMM, 7/5/06; Muscatine: Mad Cr., Muscatine, CTH, 6/30/04; O’ Brien: Little Waterman Cr., Hartley, CTH, 7/9/96; PaloAlto: Prairie Cr., Whittemore, CTH, 10/2/01; Plymouth: Beaver Cr., Akron, TMM, 8/12/13; Pocahontas: Big Cedar Cr., Fonda, JML, 9/27/06; Polk: Four Mile Cr., Ankeny, CTH, 9/29/99; Poweshiek: Walnut Cr., Belle Plaine, CTH, 9/17/08; Sac: North Raccoon R., Sac City, TMM, 8/12/13; Scott: Lost Cr., Princeton, CTH, 8/9/12; Sioux: Sixmile Cr., Hawarden, KBK, 8/3/00; Story: Walnut Cr., Ames, CTH, 7/15/99; Tama: Little Wolf Cr., Gladbrook, CTH, 8/12/04; Van Buren: Chequest Cr., Pittsburgh, CTH, 7/17/97; Webster: Deer Cr., Clare, TMM, 7/28/04; Winnebago: Little Buffalo Cr., Thompson, DPH, 7/27/04; Winneshiek: Yellow R., Postville, CTH, 7/25/07; Worth: Elk Cr., Kensett, CTH, 8/26/08; Wright: Otter Cr., Holmes, KLS, 8/11/00. Distribution: Primarily northeastern USA, extending from New York in the north to South Carolina in the south and west to the Great Plains.

Dubiraphia vittata (Melsheimer) . Allamakee: Yellow R., Ion, CTH, 9/29/98; Benton: Mud Cr., Vinton, CTH, 8/29/05; Black Hawk: Cedar R., Waterloo, CTH, 9/3/13; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Buena Vista: Brooke Cr., Rembrandt, JML, 10/5/05; Cedar: Clear Cr., Lisbon, CTH, 7/26/01; Cerro Gordo: Bailey Cr., Thornton, CTH, 7/23/03; Cherokee: Maple Cr., Aurelia, CTH, 8/6/97; Clay: Ocheyedan River, Spencer, TMM, 8/26/03; Delaware: Plum Cr., Earlville, CTH, 8/16/06; Dickinson: Little Sioux R., Lake Park, CTH, 9/3/96; Floyd: Cedar R., Charles City, MWB, 9/5/00; Franklin: Maynes Cr., Hampton, CTH, 8/18/08; Fremont: Plum Cr., Thurman, KLS, 8/3/05; Greene: East Buttrick Cr., Dana, TMM, 7/30/09; Guthrie: Deer Cr., Stuart, CTH, 7/30/98; Hamilton: Boone R., Stratford, CTH, 9/17/98; Hancock: Unnamed trib. to East Branch Iowa R., Goodell, TMM, 7/ 23/12; Hardin: South Fk. Iowa R., Iowa Falls, CTH, 7/19/95; Henry: Heather Branch, Mt. Pleasant, CTH, 10/14/98; Howard: Little Turkey R., Protivin, CTH, 8/19/02; Jackson: Farmers Cr., Fulton, CTH, 8/8/07; Johnson: ( Wickham 1911); Jones: Maquoketa R., Monticello, CTH, 9/6/05; Kossuth: Plum Cr., Wesley, TMM, 8/15/06; Linn: Crabapple Cr., Springville, CTH, 7/17/01; Lyon: Mud Cr., Alvord, KLS, 8/29/05; Mitchell: Deer Cr., Carpenter, CTH, 8/15/95; Monona: Little Sioux R., Turin, TMM, 8/7/12; O’ Brien: Waterman Cr., Sutherland, TMM, 8/26/08; Page: East Nishnabotna R., Shenandoah, KLS, 10/17/02; Pocahontas: Big Cedar Cr., Fonda, JML, 9/27/06; Pottawattamie: Pony Cr., Council Bluffs, KJS, 9/27/05; Sac: North Raccoon R., Sac City, TMM, 9/17/08; Scott: Lost Cr., Princeton, CTH, 8/9/12; Sioux: Floyd R., Alton, TMM, 8/6/03; Story: Long Dick Cr., Roland, CTH, 9/22/97; Webster: Lizard Cr., Clare, CTH, 9/11/96; Winnebago: Winnebago R., Lake Mills, CTH, 8/16/95; Worth: Willow Cr., Hanlontown, DPH, 9/4/01; Wright: East Branch Iowa R., Belmond, KLS, 8/25/05. Distribution: Primarily eastern USA, extending from New Hampshire in the north to Florida in the south and west to the Great Plains.

Macronychus glabratus Say. Adair : Middle Nodaway R., Bridgewater, CTH, 10/21/98; Allamakee: Upper Iowa R., Dorchester, MWB, 9/11/01; Appanoose: Chariton R., Centerville, JML, 9/19/01; Benton: Bear Cr., Shellsburg, CTH, 8/2/01; Black Hawk: Black Hawk Cr., Hudson, CTH, 8/29/96; Boone: Squaw Cr., Zenorsville, KBK, 7/14/00; Bremer: East Branch Wapsipinicon R., Tripoli, CTH, 9/3/03; Buchanan: Lime Cr., Brandon, CTH, 8/26/97; Butler: Johnson Cr., Aplington, MWB, 9/11/02; Calhoun: Cedar Cr., Lohrville, JML, 7/31/02; Carroll: Marrowbone Cr., Lanesboro, TMM, 9/ 13/07; Cass: West Nodaway R., Grant, CTH, 8/ 19/98; Cedar: Rock Cr., Tipton, CTH, 8/9/95; Cerro Gordo: Willow Cr., Clear Lake, CTH, 7/ 28/04; Cherokee: Willow Cr., Quimby, JML, 8/14/01; Chickasaw: Wapsipinicon R., New Hampton, CTH, 10/6/04; Clarke: South Whitebreast Cr., Weldon, TMM, 9/29/10; Clay: Little Sioux R., Spencer, TMM, 9/6/05; Clayton: Roberts Cr., St. Olaf, CTH, 7/31/00; Clinton: Silver Cr., DeWitt, CTH, 7/21/98; Crawford: Trinkle Cr., Kiron, TMM, 10/5/ 15; Dallas: Mosquito Cr., Panora, CTH, 7/15/ 96; Davis: Soap Cr., Drakesville, CTH, 8/16/04; Decatur: Weldon R., Woodland, KLS, 9/25/03; Delaware: Plum Cr., Hopkinton, CTH, 9/2/03; Des Moines: Brush Cr., Spring Grove, CTH, 9/28/00; Dickinson: Little Sioux R., Lake Park, TMM, 7/23/03; Dubuque: Lytle Cr., Zwingle, CTH, 8/16/99; Emmett: Ditch 151, Wallingford, TMM, 6/17/03; Fayette: Turkey R., Eldorado, CTH, 10/1/02; Floyd: Cedar R., Floyd, CTH, 9/4/ 02; Franklin: Bailey Cr., Sheffield, CTH, 7/24/ 03; Fremont: Plum Cr., Thurman, KLS, 8/3/05; Greene: Buttrick Cr., Grand Junction, CTH, 8/ 30/95; Grundy: South Beaver Cr., Parkersburg, CTH, 9/3/08; Guthrie: South Raccoon R., Stuart, KLS, 9/13/00; Hamilton: White Fox Cr., Webster City, CTH, 11/794; Hancock: East Branch Iowa R., Goodell, TMM, 7/24/12; Hardin: South Fk. Iowa R., Iowa Falls, CTH, 7/19/95; Harrison: Stowe Cr., Pisgah, TMM, 8/26/15; Henry: Big Cr., New London, CTH, 8/30/05; Howard: Crane Cr., Riceville, CTH, 7/10/06; Humboldt: West Fk. Des Moines R., Humboldt, TMM, 8/2/12; Iowa: Big Bear Cr., Victor, CTH, 9/20/99; Jackson: Bear Cr., Baldwin, CTH, 9/3/97; Jasper: Mud Cr., Baxter, CTH, 9/18/98; Jefferson: Cedar Cr., Stockport, CTH, 7/20/05; Johnson: Iowa R., Iowa City, CTH, 9/20/05; Jones: Maquoketa R., Monticello, 9/6/05; Keokuk: Cedar Cr., Delta, CTH, 8/18/03; Kossuth: Little Buffalo Cr., Titonka, JML, 8/ 21/00; Linn: Buffalo Cr., Central City, CTH, 8/2/96; Louisa: Long Cr., Columbus Junction, CTH, 9/26/95; Lucas: Whitebreast Cr., Lacona, TMM, 7/29/14; Lyon: Little Rock R., George, TMM, 10/1/08; Madison: North Branch North R., Winterset, CTH, 8/12/94; Mahaska: Buck Cr., Barnes City, CTH, 8/3/94; Marion: English Cr., Knoxville, MWB, 7/18/05; Marshall: Burnett Cr., Marshalltown, CTH, 8/22/00; Mills: Keg Cr., Mineola, CTH, 7/30/97; Mitchell: Rock Cr., Rock Cr., CTH, 8/15/96; Monona: Soldier R., Moorhead, TMM, 10/6/04; Montgomery: East Nishnabotna R., TMM, Red Oak, 10/12/07; Muscatine: Mud Cr., Durant, CTH, 9/16/96; O’ Brien: Little Floyd R., Sheldon, JML, 9/12/ 01; Page: West Tarkio Cr., Shenandoah, CTH, 10/12/95; Plymouth: Floyd R., LeMars, TMM, 8/24/04; Pocahontas: Des Moines R., West Bend, TMM, 8/17/05; Polk: Walnut Cr., Windsor Heights, CTH, 8/26/98; Pottawattamie: Pidgeon Cr., Neola, CTH, 7/29/97; Poweshiek: Bear Cr., Brooklyn, CTH, 9/25/02; Ringgold: Middle Fk. Grand R., Mount Ayr, TMM, 8/24/11; Sac: North Raccoon R., Sac City, CTH, 10/26/99; Scott: Lost Cr., Princeton, CTH, 9/10/98; Shelby: West Nishnabotna R., Kirkman, CTH, 8/15/94; Sioux: Floyd R., Alton, TMM, 8/6/03; Story: South Skunk R., Ames, CTH, 9/15/95; Tama: Richland Cr., Haven, CTH, 9/8/95; Van Buren: Fox R., Mt. Sterling, CTH, 7/19/05; Warren: North R., Norwalk, CTH, 10/15/98; Washington: English R., Riverside, CTH, 6/9/00; Wayne: Jackson Cr., Corydon, TMM, 9/14/11; Webster: Prairie Cr., Lehigh, CTH, 8/22/96; Winnebago: Winnebago R., Lake Mills, CTH, 9/6/00; Winneshiek: Canoe Cr., Decorah, CTH, 9/9/03; Woodbury: West Fk. Little Sioux R., Bronson, TMM, 8/9/05; Wright: Boone R., Renwick, TMM, 8/20/03. Distribution: Eastern USA, extending west to Texas in the south, Oklahoma, Nebraska, and Minnesota in the north.

Microcylloepus pusillus (LeConte) . (Downie and Arnett 1996). Distribution: Eastern USA, extending west to Utah, Colorado , and Wyoming.

Optioservus fastiditus (LeConte) . Allamakee: Yellow R., Waukon, CTH, 8/8/06; Buchanan: Pine Cr., Quasqeton, CTH, 8/28/97; Cedar: Clear Cr., Lisbon, CTH, 7/26/01; Cerro Gordo: Calmus Cr., Mason City, MWB, 9/24/01; Clayton: Silver Cr., Gunder, CTH, 8/2/00; Clinton: Silver Cr., DeWitt, CTH, 7/14/04; Delaware: Fenchel Cr., Strawberry Point, CTH, 9/17/07; Dubuque: Middle Fk. Little Maquoketa R., Rickardsville, CTH, 9/28/04; Fayette: Dibble Cr., Clermont, CTH, 8/4/98; Floyd: Little Cedar R., Colwell County Park, CTH, 9/9/09; Franklin: Bailey Cr., Sheffield, CTH, 7/24/03; Howard: Unnamed trib. To Bohemian Cr., Protivin, MWB, 6/17/03; Jackson: Farmers Cr., LaMotte, CTH, 8/8/07; Jones: Unnamed trib. To North Fk. Walnut Cr., Morley, CTH, 8/15/02; Linn: Indian Cr., Cedar Rapids, CTH, 9/25/00; Mitchell: Burr Oak Cr., Osage, CTH, 8/14/95; Winneshiek: Coldwater Cr., Bluffton, CTH, 7/10/95; Worth: Willow Cr., Hanlontown, CTH, 8/17/95. Distribution: Northeastern USA, extending west to North Dakota and south to Iowa, Illinois, and Indiana. Habitat preference is gravel and rocky substrate in medium to fast flowing streams.

Stenelmis bicarinata LeConte. Buchanan : ( Wickham 1911); Johnson: ( Wickham 1911). Distribution: Eastern United States, extending west to New Mexico in the south and Minnesota in the north. Iowa, along with Minnesota, forms the northwestern margin of its range .

Stenelmis cheryl Brown. Allamakee : Upper Iowa R., Hanover, MWB, 7/21/07; Benton: Cedar R., Vinton, MWB, 9/14/07; Black Hawk: Wolf Cr., La Porte City, CTH, 8/19/04; Bremer: Cedar R., Janesville, CTH, 9/9/14; Butler: Shell Rock R., Shell Rock, CTH, 9/8/11; Cedar: Cedar R., West Branch, CTH, 9/12/06; Cerro Gordo: Winnebago R., Mason City, CTH, 8/7/12; Clayton: Turkey R., Elkader, JOK, 9/13/88; Davis: Soap Cr., Eldon, CTH, 7/23/02; Fayette: Turkey R., Eldorado, CTH, 10/1/02; Floyd: Cedar R., Floyd, CTH, 9/4/02; Franklin: Beaver Cr., Ackley, CTH, 8/22/13; Hamilton: Boone R., Stratford, TMM, 8/16/12; Hardin: Iowa R., Eldora, CTH, 8/16/05; Howard: Upper Iowa R., Florenceville, CTH, 10/1/02; Jackson: Maquoketa R., Maquoketa, CTH, 9/13/12; Johnson: Iowa R., Iowa City, CTH, 9/22/05; Lee: Des Moines R., Keokuk, CTH, 10/1/03; Linn: Cedar R., Bertram, CTH, 10/ 10/05; Louisa: Iowa R., Wapello, CTH, 10/8/ 13; Mahaska: South Skunk R., Oskaloosa, CTH, 10/7/13; Mitchell: Cedar R., Osage, CTH, 7/6/06; Monona: Little Sioux R., Rodney, TMM, 7/18/06; Muscatine: Cedar

R., Conesville, CTH, 8/29/12; Poweshiek: North Skunk R., Millgrove, CTH, 8/6/01; Scott: Wapsipinicon R., DeWitt, CTH, 9/16/13; Van Buren: Des Moines R., Farmington, CTH, 9/6/ 06; Wapello: Des Moines R., Ottumwa, CTH, 10/7/13; Winneshiek: Upper Iowa R., Decorah, CTH, 7/25/06. Distribution: Northeastern USA, extending southwest to New Mexico in the south and Minnesota in the north, with Iowa forming part of the western limit of its range.

Stenelmis crenata (Say) . Allamakee: Yellow R., Ion CTH, 9/28/98; Appanoose: Chariton R., Centerville, JML, 9/19/01; Benton: Bear Cr., Shellsburg, CTH, 8/8/95; Black Hawk: Black Hawk Cr., Hudson, CTH, 7/31/96; Boone: Squaw Cr., Zenorsville, KBK, 7/14/00; Bremer: Horton Cr., Horton, CTH, 7/24/01; Buchanan: Lime Cr., Brandon, CTH, 6/12/95; Butler: Coldwater Cr., Greene, CTH, 8/29/96; Calhoun: Cedar Cr., Lohrville, JML, 7/31/02; Carroll: Brushy Cr., Dedham, TMM, 9/16/09; Cass: West Nodaway R., Grant, CTH, 8/19/98; Cedar: Rock Cr., Tipton, CTH, 8/9/95; Cerro Gordo: Calmus Cr., Mason City, MWB, 9/24/01; Cherokee: Maple R., Aurelia, CTH, 8/6/97; Chickasaw: Wapsipinicon R. New Hampton, CTH, 10/6/04; Clay: Willow Cr., Royal, CTH, 8/26/02; Clayton: Roberts Cr., St. Olaf, CTH, 7/31/00; Clinton: Silver Cr., DeWitt, CTH, 7/21/98; Crawford: Otter Cr., Deloit, KLS, 8/11/04; Dallas: Mosquito Cr., Panora, CTH, 7/15/96; Davis: Soap Cr., Peach Ave., CTH, 7/17/13; Decatur: Little R., Leon, CTH, 9/23/02; Delaware: Plum Cr., Hopkinton, CTH, 9/2/03; Dickinson: Little Sioux R., Lake Park, CTH, 9/3/ 96; Dubuque: Lytle Cr., Zwingle, CTH, 8/16/99; Fayette: Volga R., Maynard, CTH, 9/20/95; Floyd: Little Cedar R., Colwell, CTH, 10/3/ 95; Franklin: Maynes Cr., Hampton, CTH, 7/ 18/95; Greene: Buttrick Cr., Grand Junction, CTH, 8/30/95; Grundy: Middle Fk. South Beaver Cr., Wellsburg, CTH, 8/14/01; Guthrie: South Raccoon R., Stuart, KLS, 9/13/00; Hamilton: White Fox Cr., Webster City, CTH, 8/22/95; Hancock: East Branch Iowa R., Goodell, JML, 10/17/00; Hardin: South Fk. Iowa R., Iowa Falls, CTH, 7/19/95; Harrison: Soldier R., Pisgah, CTH, 10/27/99; Henry: Big Cedar Cr., Oakland Mills, MWB, 9/2/99; Howard: Little Turkey R., Protivin, CTH, 8/ 19/02; Humboldt: West Fk. Des Moines R., Humboldt, TMM, 9/10/13; Ida: Maple R., Ida Grove, CTH, 8/5/97; Jackson: Bear Cr., Baldwin, CTH, 9/3/97; Jasper: Indian Cr., Colfax, TMM, 10/1/12; Johnson: ( Wickham 1911); Jones: Bear Cr., Wyoming, CTH, 7/13/04; Kossuth: Black Cat Cr., Algona, CTH, 8/13/97; Lee: Lick Cr., Farmington, CTH, 7/14/03; Linn: Buffalo Cr., Central City, CTH, 8/2/96; Louisa: Long Cr., Columbus Junction, CTH, 9/26/95; Lucas: Wolf Cr., Chariton, TMM, 8/26/11; Lyon: Little Rock R., George, TMM, 10/1/08; Madison: North Branch North R., Winterset, CTH, 6/20/95; Marion: Whitebreast Cr., Knoxville, TMM, 8/ 5/13; Marshall: Minerva Cr., St. Anthony, TMM, 8/8/06; Mills: Keg Cr., Mineola, JML, 8/2/01; Mitchell: Deer Cr., Carpenter, CTH, 8/15/95; Monona: Monona-Harrison Ditch, Blencoe, TMM, 7/19/06; Montgomery: East Nishnabotna R., Red Oak, TMM, 10/12/07; Muscatine: Mud Cr., Durant, CTH, 9/16/96; O’ Brien: Floyd R., Sheldon, CTH, 9/6/95; Plymouth: Floyd R., James, JML, 9/18/08 Pocahontas: Big Cedar Cr., Fonda, JML, 9/27/06; Polk: Walnut Cr., Windsor Heights, CTH, 8/26/98; Pottawattamie: Mosquito Cr., Underwood, TMM, 8/17/04; Ringgold: Lotts Cr., Lamoni, TMM, 7/18/03; Sac: North Raccoon R., Sac City, KBK, 9/20/01; Scott: Lost Cr., Princeton, CTH, 8/9/12; Shelby: East Branch West Nishnabotna R., Avoca, JML, 7/18/00; Sioux: Sixmile Cr., Hawarden, JML, 8/2/ 00; Story: South Skunk R., Ames, CTH, 9/15/95; Tama: Little Wolf Cr., Gladbrook, CTH, 7/1/04; Taylor: Honey Cr., Bedford, CTH, 8/18/98; Van Buren: Chequest Cr., Pittsburgh, CTH, 7/17/97; Wapello: Middle Avery Cr., Chillicothe, MWB, 8/27/02; Warren: North R., Norwalk, CTH, 10/ 15/99; Washington: English R., Riverside, CTH, 6/9/00; Wayne: South Fk. Chariton R., Corydon, TMM, 9/12/12; Webster: Prairie Cr., Lehigh, CTH, 8/22/96; Winnebago: Winnebago R., Lake Mills, CTH, 8/16/95; Winneshiek: Middle Bear Cr., Highlandville, CTH, 7/11/95; Woodbury: West Fk. Little Sioux R., Bronson, TMM, 8/9/05; Worth: Willow Cr., Hanlontown, CTH, 9/4/01; Wright: Otter Cr., Goldfield, JML, 8/11/01. Distribution: Eastern USA, exclusive of Florida, extending west to Oklahoma, Colorado, and North Dakota in the north.

Stenelmis decorata Sanderson. Allamakee : Upper Iowa R., Dorchester, CTH, 8/24/05; Appanoose: Chariton R., Centerville, CTH, 10/15/99; Benton: Cedar R., Vinton, MWB, 9/14/07; Black Hawk: Wolf Cr., La Porte City, CTH, 8/19/04; Bremer: Cedar R., Janesville, CTH, 9/8/11; Buchanan: Bear Cr., Brandon, CTH, 9/8/09; Butler: Shell Rock R., Shell Rock, CTH, 9/8/ 14; Carroll: Elk Run Cr., Lanesboro, TMM, 8/ 4/11; Cerro Gordo: Winnebago R., Mason City, CTH, 8/7/12; Chickasaw: Wapsipinicon R., Twin Ponds Park, CTH, 10/14/10; Clayton: Volga R., Elkport, CTH, 9/27/02; Clinton: Silver Cr., DeWitt, CTH, 7/14/04; Davis: Soap Cr., Floris, CTH, 7/24/02; Delaware: Buck Cr., Delhi, MWB, 8/16/01; Des Moines: Skunk R., Burlington, MWB, 9/7/06; Dubuque: North Fk. Maquoketa R., New Vienna, CTH, 8/21/01; Fayette: Little Turkey R., West Union, CTH, 9/ 10/03; Floyd: Cedar R., Floyd, CTH, 9/4/02; Grundy: South Beaver Cr., Parkersburg, CTH, 9/10/14; Guthrie: Middle Raccoon R., Coon Rapids, MWB, 6/16/07; Henry: Skunk R., Oakland Mills, CTH, 9/24/08; Howard: Upper Iowa R., Florenceville, CTH, 10/1/02; Jackson: Bear Cr., Baldwin, CTH, 8/28/03; Johnson: Iowa R., Iowa City, CTH, 8/8/05; Jones: Maquoketa R., Monticello, CTH, 9/6/05; Kossuth: Lotts Cr., West Bend, MWB, 9/25/02; Lee: Des Moines R., Keokuk, CTH, 10/1/03; Linn: Wapsipinicon R., Central City, CTH, 9/27/06; Louisa: Iowa R., Wapello, CTH, 9/26/02; Mahaska: North Skunk R., Rose Hill, CTH, 8/14/02; Mitchell: Wapsipinicon R., McIntyre, MWB, 8/27/08; Muscatine: Pike Run, Nichols, CTH, 7/26/04; O’ Brien: Waterman Cr., Sutherland, JML, 8/15/01; Poweshiek: Walnut Cr., Belle Plaine, CTH, 7/29/03; Scott: Wapsipinicon R., DeWitt, CTH, 9/16/13; Van Buren: Chequest Cr., Pittsburg, CTH, 7/15/03; Wapello: Des Moines R., Eldon, CTH, 9/30/03; Warren: North R., Norwalk, CTH, 10/15/99; Washington: English R., Riverside, MWB, 9/10/08; Wayne: South Fk. Chariton R., Corydon, TMM, 8/8/16; Winneshiek: Ten Mile Cr., Ridgeway, CTH, 8/ 16/11. Distribution: Eastern USA, extending west to the southern Great Plains, with Iowa forming part of the western border.

Stenelmis grossa Sanderson. Adair : Middle Nodaway R., Bridgewater, CTH, 10/21/98; Allamakee: Upper Iowa R., Dorchester, CTH, 9/27/02; Benton: Pratt Cr., Mt. Auburn, CTH, 9/ 5/00; Black Hawk: West Fk. Cedar R., Finchford, CTH, 9/14/01; Boone: Squaw Cr., Zenorsville, KBK, 7/14/00; Bremer: Cedar R., Janesville, CTH, 9/8/11; Buchanan: Lime Cr., Brandon, CTH, 8/9/00; Butler: Beaver Cr., New Hartford, CTH, 8/14/03; Cass: Sevenmile Cr., Grant, KLS, 7/26/06; Cedar: Rock Cr., Tipton, CTH, 8/9/95; Cerro Gordo: Winnebago R., Mason City, CTH, 9/8/14; Cherokee: Mill Cr., Larrabee, CTH, 10/7/98; Chickasaw: Wapsipinicon R., New Hampton, CTH, 8/18/04; Clay: Prairie Cr., Spencer, JML, 9/4/01; Clayton: Roberts Cr., St. Olaf, CTH, 7/31/00; Clinton: Barber Cr., Grand Mound, CTH, 8/30/04; Crawford: Big Cr., Denison, TMM, 8/10/04; Dallas: Mosquito Cr., Panora, TMM, 9/18/09; Davis: Soap Cr., Eldon, CTH, 7/23/02; Delaware: Buck Cr., Delhi, MWB, 8/16/01; Des Moines: Skunk R., Burlington, MWB, 9/7/06; Dickinson: Little Sioux R., Milford, TMM, 7/23/03; Fayette: Dibble Cr., Clermont, CTH, 8/4/98; Floyd: Little Cedar R., Colwell, CTH, 8/20/02; Franklin: Bailey Cr., Sheffield, CTH, 7/24/03; Greene: West Buttrick Cr., Grand Junction, CTH, 8/30/95; Grundy: South Beaver Cr., Parkersburg, CTH, 8/13/01; Hamilton: White Fox Cr., Webster City, CTH, 8/22/95; Hancock: Winnebago R., Forest City, TMM, 9/18/06; Hardin: South Fk. Iowa R., Iowa Falls, CTH, 7/19/95; Henry: Big Cedar Cr., Oakland Mills, CTH, 9/12/00; Howard: Little Turkey R., Protivin, CTH, 8/19/02; Humboldt: West Fk. Des Moines R., Humboldt, TMM, 9/9/14; Ida: Maple R., Galva, CTH, 8/6/97; Jackson: Bear Cr., Baldwin, CTH, 9/3/97; Jasper: North Skunk R., Newton, CTH, 8/8/01; Jefferson: Cedar Cr., Stockport, CTH, 7/20/05; Johnson: Iowa R., Iowa City, CTH, 8/8/05; Jones: Bear Cr., Wyoming, CTH, 7/13/04; Keokuk: North Skunk R., Sigourney, CTH, 10/7/13; Kossuth: Little Buffalo Cr., Titonka, JML, 8/21/00; Lee: Des Moines R., Keokuk, CTH, 10/1/03; Linn: Buffalo Cr., Central City, CTH, 9/2/08; Louisa: Buffington Cr., Columbus City, CTH, 9/25//95; Lucas: Wolf Cr., Chariton, TMM, 8/26/11; Lyon: Little Rock R., George, KLS, 8/5/03; Madison: North Branch North R., Winterset, CTH, 8/12/94; Mahaska: North Skunk R., Rose Hill, CTH, 8/14/02; Marshall: South Minerva Cr., Clemons, TMM, 8/8/07; Mitchell: Deer Cr., Carpenter, CTH, 8/15/ 95; Monona: Maple R., Turin, TMM, 8/26/03; Muscatine: Mud Cr., Durant, CTH, 9/16/96; O’ Brien: Little Floyd R., Sheldon, CTH, 9/13/ 99; Page: West Tarkio Cr., Essex, KLS, 7/7/04; Plymouth: Floyd R., James, CTH, 10/26/99; Pocahontas: North Branch Lizard Cr., Havelock, TMM,7/27/04; Pottawattamie: Boyer R., Missouri Valley, TMM, 8/13/03; Poweshiek: Little Bear Cr., Brooklyn, CTH, 8/18/05; Sac: Boyer R., Early, TMM, 8/19/03; Scott: Wapsipinicon R., DeWitt, CTH, 8/30/12; Sioux: Sixmile Cr., Hawarden, JML, 8/2/00; Story: South Skunk R., Ames, CTH, 9/19/97; Tama: Little Wolf Cr., Gladbrook, CTH, 8/12/04; Van Buren: Chequest Cr., Pittsburg, CTH, 7/15/03; Warren: Middle R., Indianola, TMM, 9/18/13; Washington: Crooked Cr., Crawfordsville, CTH, 7/14/05; Webster: Prairie Cr., Lehigh, CTH, 8/22/96; Winneshiek: Yellow R., Postville, CTH, 7/25/07; Woodbury: Little Sioux R., Smithland, TMM, 8/18/14; Worth: Deer Cr., Carpenter, CTH, 8/21/01. Distribution: Eastern USA, extending west to North Dakota in the north, Nebraska, and Texas in the south.

Stenelmis lignicola (Schmude) . Des Moines: Burlington ( Schmude et al. 1992). Distribution: Southeastern USA, with Iowa forming part of the northwestern border.

Stenelmis musgravei Sanderson. Allamakee : Upper Iowa R., Dorchester , MWB, 8/24/05. Distribution: Eastern USA, extending west to Wisconsin in the north and Texas in the south. Iowa forms part of the western border of its distribution .

Stenelmis sexlineata Sanderson. Cerro Gordo : Calmus Cr., Mason City, MWB, 9/24/01; Cherokee: Little Sioux R., Larrabee, TMM, 8/ 21/13; Clay: Willow Cr., Royal, CTH, 9/13/99; Clayton: Nutting Cr., Clermont, CTH, 9/21/09; Dallas: South Raccoon R., Redfield, TMM, 9/6/ 13; Des Moines: Brush Cr., Spring Grove, CTH, 9/28/00; Dickinson: Little Sioux R., Milford, CTH, 9/4/96; Fayette: Nutting Cr., Clermont, CTH, 9/21/09; Guthrie: Deer Cr., Stuart, TMM, 7/29/04; Henry: Big Cr., New London, CTH, 8/ 30/05; Lee: Lick Cr., Farmington, CTH, 7/14/03; Louisa: Buffington Cr., Columbus City, CTH, 9/ 25/95; Madison: North Branch North R., Winterset, CTH, 8/12/94; Mitchell: Deer Carpenter, CTH, 8/15/95; O’ Brien: Little Waterman Cr., Hartley, JML, 8/27/02; Sioux: Floyd R., Hospers, TMM, 8/16/06; Van Buren: Honey Cr., Bonaparte, CTH, 6/24/03; Warren: North R., Norwalk, CTH, 10/15/98; Washington: Crooked Cr., Crawfordsville, CTH, 7/14/05. Distribution: Central USA, west to South Dakota and Kansas, east to Indiana and Kentucky.