Ketengus Bleeker, 1846

Marceniuk, Alexandre Pires, Oliveira, Claudio & Ferraris Jr, Carl J., 2024, A new classification of the family Ariidae (Osteichthyes: Ostariophysi: Siluriformes) based on combined analyses of morphological and molecular data, Zoological Journal of the Linnean Society 200 (2), pp. 426-476 : 460

Ketengus Bleeker, 1846

( Figs 1–3 , 34 )

Type species: Ketengus typus Bleeker, 1846 .

Diagnosis

Mesethmoid medial notch large and shallow (1, 2> 0); fenestra delimited by mesethmoid and lateral ethmoid present (3, 0> 1); fenestra delimited by mesethmoid and lateral ethmoid large, filled with cartilage (4,?> 0); mesethmoid posterior horn tubular, narrow and elongate (5, 0> 1); posterior branches of mesethmoid narrow (6, 0> 1); posterior branches of mesethmoid parallel throughout their entire extension (7, 0> 1); lateral ethmoid not contact frontal (14, 3> 0); vomer diamond shaped (49, 1> 0); vomer lateral processes absent (51, 1> 0); accessory tooth plates absent (58, 1> 0); three anterior branches in lachrymal-antorbital anterior part (98, 2> 3); teeth on dentary along entire bone (116, 1> 2); dorsal crest of premaxilla present (125, 0> 1); opercle posterior portion not well developed posteriorly (129, 1> 0); ventral crest of hyomandibula absent (142, 0> 1); dorsal crest of urohyal projected anteriorly (156, 0> 1); one basibranchial series (164, 0> 1); first hypobranchial club shaped (171, 0> 1); second hypobranchial club shaped (175, 0> 1); first epibranchial overlaying second epibranchial (179, 0> 1); third pharyngobranchial funnel shaped (192, 1> 0); seventh vertebra free from ventral superficial ossification (214, 0> 1).

Included species

Ketengus typus Bleeker, 1846 .

Habitat and distribution: Brackish waters, South and Southeast Asia ( Fig. 24 ).

Remarks

The close relationship of Ketengus plus Osteogeneiosus established in previous morphological and molecular studies ( Marceniuk and Menezes 2007, Betancur-R. 2009, Marceniuk et al. 2012) is further supported by the results of the total-evidence analysis.

Figures

Figure 1. MP analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support.

Figure 2. ML analysis. Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green). Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support.

Figure 3. BI analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support.

Figure 24. Representative skulls, in dorsal view, and distribution of the subtribe Ariina.A, Arius maculatus, AUM 44875. B, Cephalocassis melanochir, CAS 49426. C, Cryptarius truncatus, AUM 50243. D, Hexanematichthys sagor, AUM 50242. E, Kyataphisa nenga, AUM 46280. F, Osteogeneiosus militaris, AUM 44880. G, Pseudosciades sona, AUM 44861. * marine, ** brackish water, *** freshwater.

Figure 34. Ketengus typus, ANSP 60704, eastern Indian Ocean, type species of the genus.