Antecerococcus Green, 1901

Chris J. Hodgson & Douglas J. Williams, 2016, (Hemiptera: Sternorrhyncha, Coccomorpha) with particular reference to species from the Afrotropical, western Palaearctic and western Oriental Regions, with the revival of Antecerococcus Green and description of a new genus and fifteen new species, and with ten new synonomies, Zootaxa 4091 (1), pp. 1-175 : 19

publication ID	https://doi.org/ 10.11646/zootaxa.4091.1.1
publication LSID	urn:lsid:zoobank.org:pub:76D13D36-682E-4E91-AC91-693CA9D3D465
DOI	https://doi.org/10.5281/zenodo.6081534
persistent identifier	https://treatment.plazi.org/id/03F2FF48-8139-0D2D-24B6-AA99FB79FDB3
treatment provided by	Plazi
scientific name	Antecerococcus Green, 1901
status

Treatment

Antecerococcus Green, 1901 , revived status.

Antecerococcus Green 1901: 560 . Type species: Antecerococcus punctiferus Green , by monotypy.

Phenacobryum Cockerell 1902: 114 . Type species: Planchonia bryoides Maskell , subsequently designated by Borchsenius, 1960d: 110. Synonymy with Cerococcus by Green 1908: 41. Syn. nov.

Amelococcus Marchal 1904: 557 , 560. Type species: Amelococcus alluaudi Marchal , by monotypy and original designation. Synonymy with Cerococcus by Lambdin & Kosztarab 1977: 16. Syn. nov.

Cercococcus Scott 1907: 455 . Type species: Cercococcus eremobius Scott , by monotypy. Synonymy with Cerococcus by Green 1908: 41. Syn. nov.

Coricoccus Mahdihassan 1933: 562 . Type species: Coricoccus ornatus Green. Synonymy with Cerococcus by Lambdin & Kosztarab 1977: 16. Syn. nov.

Note. Anterocerococcus was synonymised with Cerococcus by Green (1908), but here its status is revised and it is removed from synonymy with Cerococcus Comstock. This revival is because the adult females of all species in Antecerococcus were found to share a range of morphological features, including an anteroventral sclerotization on each anal lobe, that are absent from the adult females of all species of Cerococcus as redefined below. With the recognition of Antecerococcus Green , the names Phenacobryum Cockerell , Amelococcus Marchal , Cercococcus Scott and Coricoccus Mahdihassan become synonyms of Antecerococcus rather than of Cerococcus because their type species (listed above) fall within this genus.

Generic description and diagnosis. Adult female. Mounted material. Basic body structure typical of Cerococcidae ( Figs 4–49 View FIGURE 4 View FIGURE 5 View FIGURE 6 View FIGURE 7 View FIGURE 8 View FIGURE 9 View FIGURE 10 View FIGURE 11 View FIGURE 12 View FIGURE 13 View FIGURE 14 View FIGURE 15 View FIGURE 16 View FIGURE 17 View FIGURE 18 View FIGURE 19 View FIGURE 20 View FIGURE 21 View FIGURE 22 View FIGURE 23 View FIGURE 24 View FIGURE 25 View FIGURE 26 View FIGURE 27 View FIGURE 28 View FIGURE 29 View FIGURE 30 View FIGURE 31 View FIGURE 32 View FIGURE 33 View FIGURE 34 View FIGURE 35 View FIGURE 36 View FIGURE 37 View FIGURE 38 View FIGURE 39 View FIGURE 40 View FIGURE 41 View FIGURE 42 View FIGURE 43 View FIGURE 44 View FIGURE 45 View FIGURE 46 View FIGURE 47 View FIGURE 48 View FIGURE 49 ). Anal lobes rarely sclerotized throughout (except in A. asparagi ), otherwise with wellsclerotized inner margins, each occasionally with reticulations or diagonal ridges, each lobe with 1 or 2 small setose seta (at most) on inner margin but with 2 long fleshy setae on dorsal surface, 1 near apex and other about half-way along each lobe, each fleshy seta several times longer than wide and generally with a blunt apex (rarely long and almost setose); ventral surface of each lobe with a setose or rather spinose seta near apex ( Fig. 3 View FIGURE 3 A); also each lobe generally with a medioventral seta and sometimes an anteroventral seta (presence of latter taxonomically significant) plus a small seta on each outer margin ( Fig. 3 View FIGURE 3 B). Median anal plate typical of family. Anal ring with 4 pairs of setae (exception An gallicolus (Mamet) with 3 pairs), each seta narrowing rather abruptly about 2/3rds along length. Dorsum with up to 4 sizes of 8-shaped pore: largest pores (>15 Μm widest) present associated with each stigmatic band but also along margins and sometimes in swirls or bands medially on dorsum; almost always with a line or band of large 8-shaped pores along dorsal margins of posterior abdominal segments; smaller sizes of 8-shaped pore generally most common pore types medially; smallest pores generally present across posterior abdominal segments (posterior to cribriform plates) and in apices of stigmatic pore bands; structure, size and distribution of dorsal 8-shaped pores important taxonomically; 8-shaped pores also frequently present within apex of each stigmatic band, mainly smallest pores but occasionally other sizes. Simple pores typical of family present but rarely showing anything significant. Tubular ducts of only 1 size on dorsum, generally slightly wider and slightly more abundant on dorsum than on venter. Cribriform plates always present in a submedial group on each side of abdominal segment IV (apart from A. rubra that appears to have long ductules), of highly variable size and structure, sometimes also in bands and on segments anterior to abdominal segment IV. Ventrally, with stigmatic pore bands generally extending onto dorsum (rarely possibly not reaching dorsum (e.g., A. sparsiporus Hodgson & Williams , sp. nov.), posterior bands bifurcated or non-bifurcated; each band with quinquelocular disc-pores but many species with more loculi in disc-pores of each apical group; also each apical group often with a small seta nearby; loculate pores (generally 5-locular) also present in a sparse group near each antenna. Small bilocular pores restricted to cephalothorax. Multilocular disc-pores present or absent; occasionally entirely absent but when present, generally each with 10 loculi, in segmental lines across abdominal segments and sometimes also on metathorax; each band normally with a submarginal group and a medial band; multilocular disc-pores on segment VII generally only represented by submarginal groups on each side of vulva but occasionally band complete; those on segments VIII and IX similar, generally in submarginal groups but occasionally forming a band posterior to vulva; multilocular disc-pores also sometimes present mesad to each spiracle. Small convex closed pores sometimes present, usually in a partial or complete line between each antenna and metathoracic leg stubs. Anteroventral sclerotized areas always present on venter laterad to anal ring ( Figs 3 View FIGURE 3 A, 3B). Leg stubs present or absent. Antennae generally unsegmented but occasionally appearing 2 segmented, usually with 7 fleshy setae but some species also with a few setose setae; some species with a distinct cone-shaped extension to apex of each antenna; other species with one or more fleshy setae in a distinct cavity. Spiracles, each with spiracular disc-pores in a group anterior to spiracle but not in a semi-circle laterad to or posterior to peritreme; occasionally with a sclerotized bar extending anteriorly over each spiracle (probably most obvious on older specimens).