Ulmeritoides angelus Souto, Da-Silva, Nessimian & Gonçalves, 2016

Orlando, Thales Yann, Salles, Frederico Falcão, Boldrini, Rafael & Krolow, Tiago Kütter, 2021, Updated records for Leptophlebiidae (Ephemeroptera) and a new species of Thraulodes Ulmer, 1920 from Tocantins State, Northern Brazil, Zootaxa 5076 (1), pp. 39-55 : 51

Ulmeritoides angelus Souto, Da-Silva, Nessimian & Gonçalves, 2016

Distribution. Brazil (Bahia and Minas Gerais). New Record: Tocantins

Material examined. BRAZIL, Tocantins, Palmas, distrito de Taquaruçu do Porto , vale do Vai Quem Quer, 10°22’0.9”S / 48°8’1”W, 04-06.xii.2018, white sheet light trap, Fernandes, A.S. and team cols. (7♂, CEUFT) .

Remarks. First record for the state. Body size, fore wing, and hind wing are smaller than the specimens from Souto et al. (2016) (6.4 – 8.9 mm; 7.7 – 8.2 mm; 1.2 – 1.4 mm versus. 9.0 mm; 11.0 mm; 2.0 mm, respectively). Penes shape and telopenis, styliger plate, and spots on bullae of veins Sc and RP1 match with the original description; however, the number of cross veins basal to bullae do not (from 8 to 11 in the original description and 6 in the observed individuals). The number of cross veins is widely used to separate species of Leptophlebiidae ; however, Kluge (2020), in his work about Thraulodes , argues that in mayflies this number varies among individuals of the same species. The only species-specific characters related with cross veins are thickness and/or coloration of cross veins and their approximate number or density in certain fields. In that sense, it is plausible to assume that smaller individuals might have a reduced number of cross veins. Based on that, we decided to maintain these specimens identified as U. angelus . It was found in only one site, in an area with Cerrado influence.