Tomarus rostratus Dupuis, 2014

López-García, Margarita M. & Deloya, Cuauhtémoc, 2018, Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 72 (2), pp. 331-338 : 332-333

Tomarus rostratus Dupuis, 2014 ( Figs. 1–13 )

Tomarus rostratus Dupuis 2014: 1

Description. Female (n = 15). Habitus as in Fig. 1 . Length 17.4–20.0 mm; humeral width 8.3–9.9 mm. Color reddish brown. Frons coarsely and irregularly rugose. Head: Frontoclypeal region with 2 transverse tubercles separated by about 3 tubercle diameters. Clypeus trapezoidal, not strongly narrowed toward apex ( Fig. 2 ); apex with 2 triangular teeth separated by about 1.5 tooth diameters. Mandibles with 2 teeth and a basal lobe, dorsal surface slightly concave. Antennal club subequal in length to antennomeres 2–7. Pronotum: Surface punctate, punctures round, smaller than those on elytra, denser over apical and lateral margins. Apical margin with a small tubercle visible in lateral view ( Fig. 3 ). Pronotal fovea shallow, elongate, narrow (about 1/8 as wide as distance between eyes), surface smooth. Scutellum: Surface without punctures. Elytra: Sutural stria present and complete. Elytral punctures dense and deep, punctures mainly ocellate, 3 pairs of distinct double rows. Legs: Protibia tridentate with additional small, basal tooth.Protarsus simple, not enlarged.Apex of metatibia with small crenulation and 14–15 spinules. Venter: Prosternal process long, thick, apex transversely oval to round and bordered with long yellow setae. Abdomen: Pygidium flat in lateral view, surface homogeneously rugopunctate, apical margin with short setae ( Fig. 10 ). Apical margin of sternite VII slightly emarginated, notch broad and shallow ( Fig. 11 ). Apical margin of sternite VIII emarginate, notch narrow and deep ( Fig. 11 ). Subcoxite subquadrate, inner border with long, yellow setae. Coxite subtriangular, strongly concave, apical border with long, yellow setae ( Fig. 12 ).

Male (n = 2). The examined specimens differ from the female and extend the variation of the species as follows: Abdomen: Pygidium convex in lateral view, basal third deeply rugopunctate, disc smooth, apical margin truncate, without setae ( Fig. 8 ). Apical margin of sternite VII straight.Sternite VIII widely emarginated ( Fig. 9 ). Parameres: Widened at apical fourth, elongated from basal fourth to apex; sides parallel, without teeth or lateral projections ( Fig. 4 ), uniformly curved at lateral view ( Fig. 6 ). Spiculum gastrale: Shape symmetrical, thickened; base with 2 attached sclerites; apex recurved ( Fig. 13 ).

Variation. Body length 17.4–27.5 mm. The number of spinules on the apex of the metatibia is 8–15. The first interval of the elytra can be densely punctate or nearly impunctate (as in T. laevicollis ). The parameres are evenly curved in lateral view ( Fig. 6 ) or with an anterior slight depression or concavity ( Fig. 7 ).

Diagnosis. Tomarus rostratus is unique in the genus because of its extremely elongate parameres ( Figs. 4–7 ). It can be differentiated by the following combination of characters: Clypeus trapezoidal, pronotum with small tubercle and shallow fovea, protibia with a small fourth basal tooth, apical border of pygidium strongly truncate in males, apical margin of sternite VIII emarginated in females.

Specimens Examined. CHILE: Arica, Cuya, 16.ii. 1989, 100 m, R. Miller & R. Stange Coll. [16, 1♀, FSCA]. Arica , Valle Azapa, 6 km E Arica, 16.ii. 1989, 100 m, R. Miller & R. Stange Coll. [8♀♀, FSCA] . PERU: Lima, xii.1922, D.S. Bullock Coll. [5♀♀, USNM]. Lima, 2.xi.1930, Luis Pro Castillo Coll. [162♀, USNM] .

Distribution. Peru ( Dupuis 2014) and Chile NEW COUNTRY RECORD.

Comments. The only previously known specimen of T. rostratus was the male holotype deposited in MNHN. The type locality was also unknown as only “ Peru ” was indicated on its labels. However , the additional records from Lima and the new records in Chile provide more precise locality data, so we now can conclude that the species occurs along the coastal deserts of Peru and Chile.

Figures

Figs. 1–3. Tomarus rostratus, female. 1) Habitus; 2) Head, dorsal view; 3) Head and pronotum, lateral view.

Figs. 4–13. Tomarus rostratus. 4, 5) Parameres, frontal view; 6, 7) Parameres, lateral view; 8) Male pygidium; 9) Male sternite VIII; 10) Female pygidium; 11) Female sternite VIII; 12) Female genital plates; 13) Male spiculum gastrale. Figs. 5 and 7 photographed by A. Mantilleri.