Tillicera cleroides Gorham, 1892

Gerstmeier, Roland & Bernhard, Nadine, 2010, Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae), Zootaxa 2359 (1), pp. 1-34 : 15-17

Tillicera cleroides Gorham, 1892

( Figs. 5–6 , 26–27 )

Tillicera cleroides Gorham 1892: 731 ; Schenkling 1902: 127; Schenkling 1906: 262; Mawdsley 1994: 414, 415. Tillicera mimomutilla ( Pic, 1926) : 21 ( Thanasimus ). n. syn.

Dasyceroclerus rouyeri Pic 1938: 4 . n. comb., n. syn.

Thanasimus mimomutilla ab. bicoloricollis Pic 1931: 7.

Tillicera mimomutilla ab. bipartita Pic 1951: 21. n. syn.

Specimens examined: Holotype: ♀. Typus, Syntypus, Tillicera cleroides, Gorham, 1892 , cleroides, Gorh., Carin Chebá , 900–1100m, L. Fea V XII-88, Museo Civico di Genova (MCSN).

Lectotype: ♀. Type, Tonkin, Thanasimus mimomutillus n. sp., Museum Paris, Coll. M. Pic, (MNHN) Tillicera mimomutilla n. syn.

Paralectotypes: ♀, same data as Lectotypus (pinned together) (1 ex., MNHN); Thanasimus , mimomutillus, Pic, 1926 , ZMAN type COLE.1796.1, Cotype: Thanasimus mimomutillus Pic 1926 , Tonkin, Lac Thò, Thanasimus mimomutillus Pic (1 ex., ZMAN); (on one pin): Paratype, Thanasimus mimomutillus Pic, Cotypes , Hoa Binh, Tonkin, Brit. Mus., 1951-293 (2 ex., BMNH).

Holotype: ♂. Type, v. Rouyeri, Pic, Mt. Semerou , Volcans Java, (Rouyer), Museum Paris, Coll. M. Pic ( MNHN), Dasyceroclerus rouyeri n. comb, n. syn.

Holotype: ♂. Type, Chapa, Tonkin, v. nov. bicoloricollis, Museum Paris, Coll. M. Pic ( MNHN)

Paratype: Thanasimus mimomutillus bicoloricollis, Pic, 1931 , ZMAN type COLE.1797.1, Cotype: Thanasimus mimomutillus Pic , v. bicoloricollis Pic , Tonkin, Hanoi, 1932, ex Coll., M. Pic ( ZMAN).

2 Types ab. bipartita: ♂. v. bipartita, Pic , Hoa Binh, Museum Paris, Coll. M. Pic ( MNHN) Tillicera mimomutilla ab. bipartita n. syn.

Others: Laos, Vientiane Prov., Vang Vieng env., Umgebung der Ortschaft, 20.7.2006, Bananenplantagen und Reisfelder, an Strauch (1 ex., RGCM); China: S-Yunnan, (Xishuangbanna), 23km NW Jinghong, vic. Na Ban, N22 09.49, E100 39.92, 05.6.2008 leg. A. Weigel, 730m second. forest KF (1 ex., RGCM); Chiang Dao, Chiang Mai Prov., N. Thailand, 24.6.1979, W. Suzuki leg. (1 ex., RGCM); Nord, Vietnam Tonkin, Hoa-Binh, ex. Coll. Oberthur, I.G. 18293 (4 ex., RGCM); Tonkin: Thai-Nien. Banks, of Fleuve Rouge, 1924, H. Stevens (1 ex., RGCM); Hanoi (Vauloger),(Coll. Ph Francois), Coll. L. Bedel 1922 (2 ex., MNHN); D. Nodier, Hanoi, Coll. M. Pic (4 ex., MNHN); Hanoi, 7.02, Coll. M. Pic (1 ex., MNHN); Tonkin, Coll. M. Pic (1 ex., MNHN); Museum Paris, Tonkin, Dap Can, H. Donckier 1907, 14. au 18.5.06 (1 ex., MNHN); Museum Paris, Tonkin, Dap Can, H. Donckier 1906, 4. au 8.5.06 (1 ex., MNHN); Dong Jang, Tonkin, 4.1903, Vauloger, (Coll. Ph Francois), Coll. L. Bedel 1922 (4 ex., MNHN); D. Nodier, Dong-Dang, Coll. M. Pic (1 ex., MNHN); Museum Paris, Tonkin, Rég de Hoa Binh, A. de Cooman 1987 (1 ex., MNHN); Museum Paris, Tonkin, Rég de Hoa Binh, A. de Cooman, ex Coll. R. Oberthur (3 ex., MNHN); Tonkin occ., Env. de Hoa-Binh, R.P.A. de Cooman, 1919 (1 ex., MNHN); Tonkin occ., Rég. de Hoa-Binh, 1919, ex Coll. R. Oberthur (1 ex., MNHN); Tonkin, Région de, Hoa-Binh, Museum Paris, 1934, A. de Cooman (1 ex., MNHN); Tonkin occ., Rég. de Hoa-Binh, 1919, ex Coll. R. Oberthur (1 ex., MNHN); Hoa-Binh, Coll. M. Pic (1 ex., MNHN); Hoa Binh, Coll. M. Pic (3 ex., MNHN); Hoa Binh, Tonkin, Coll. M. Pic (5 ex., MNHN); Museum Paris, Tonkin, Rég de Hoa-Binh, A. de Cooman 1926 (1 ex., MNHN); Lactho, Tonkin, de lineCooman, Coll. M. Pic (1 ex., MNHN); Env. de Lam, ( Tonkin), L. Blaise,(Coll. Ph Francois), Coll. L. Bedel 1922 (4 ex., MNHN); Rég. de Luc-Nam, ( Tonkin), L. Blaise, (Coll. Ph Francois), Coll. L. Bedel 1922 (5 ex., MNHN); Museum Paris, Tonkin Centr., Env. de Tuyen-Quan, A. Weiss 1901 (1 ex., MNHN); Sept-Pagodes, ( Tonkin), L. Blaise, (Coll. Ph Francois), Coll. L. Bedel 1922 (3 ex., MNHN); Museum Paris, Lakhon, Marmand 1878 (1 ex., MNHN); Phuyen Binh, 11.1907, Coll. M. Pic (2 ex., MNHN); Carin Chebà, 900–1100 m, L. Fea V XII.88 (1 ex., MNHN); Samberbajem, Res. Kediri, E.Java, ex Louwerens, Coll. M. Pic (1 ex., MNHN); Coll. M. Pic (2 ex., MNHN); Hainan Is., NamChaChuen, S.W. of Nodoa, July 8, Lingnan, University, 5th. Hainan Is., Exped. 1929 (1 ex., ZMAN); Depok, 4.1948, C. v. Nidek, Collectie, C. v. Nidek, Acq. 1969 (1 ex., ZMAN); C.J. Louwerens O.- Java, Res. Kediri, Sambong (2 ex., ZMAN); C.J. Louwerens, O.-Java, Res. Kediri, Glodok, Metatype: Thanasimus mimomutillus 1937 M. Pic (1 ex., ZMAN); 1 Ex. C.J. Louwerens, O.-Java, Res. Kediri, Plengong Qentir-, Qentiran?, Metatype: Thanasimus mimomutillus, 1937 M. Pic (1 ex., ZMAN); C.J. Louwerens, O.- Java, Res. Kediri, Zuidergebergte, Popoh, Thanasimus mimomutillus, Pic, Metatype : Thanasimus mimomutillus, 1937 M. Pic (1 ex., ZMAN); C.J. Louwerens, Java 84m, Toeloengagoeng (1 ex., ZMAN); Babakan. Z.K.-, Java Drescher, 7.11.1937 (1 ex., ZMAN); Dammerman, O. Soemba, 450 M., Mao Marroe, 5.1925 (1 ex., ZMAN).

Length: 7–10mm (83 specimens).

Collecting period: April – July, November, December.

Head: Black, glossy, anterior part of clypeus, labrum and palpi brown, posterior part of clypeus brown to black; distributed with fine and diffuse punctation; cranium vested with erect, pale setae; head including eyes as broad as anterior width of pronotum.

Antennae: A1 to A4 light brown, from A5 onwards dark brown to black, with brown and golden depressed setae; antennomeres triangularly dilated from A5 or A6 (sometimes from A7) onwards, A10 broader than long.

Pronotum: Black, glossy, anteriorly and posteriorly narrowly brown to red-brown (at sides sometimes posteriorly dilated towards apex); surface finely and diffusely punctate; vested with predominantly anteriorly directed erect, dark setae, towards apex and base also with golden setae; length:width ratio 1.14:1 (broadest in middle).

Scutellum: Brown, finely punctate, vested with pale setae.

Elytra: Basal half red-brown to brown, remainder black, behind middle with a yellowish pigmented fascia which bears dense golden setae (spanning from the lateral margin to mid-way towards suture), and a posterior pale pigmented fascia, bearing dense golden setae, and not reaching the lateral margin; disc deeply punctate in rows, diameter of punctures larger than interstices, the basal red-brown part with tubercles, apex densely and finely punctate, emarginated; basal half with erect, dark and golden setae, black part with posteriroly directed, semi-decumbent, and isolated erect dark setae, apex with golden setae gradually more erect approaching apices; length:width ratio 2.19:1.

Legs: Dark brown to black, pro- and mesofemora paler, mostly with a dark marking dorsally, metafemora paler only basally, metatibiae often with a pale marking, legs with erect, pale setae; tibiae without carinae.

Ventral surface: Abdomen dark brown to black, with few pale setae, thorax brown, with dense, pale setae; penultimate ventrite of males deeply emarginated.

Aedeagus: Conspicuously asymmetrical, with tooth-like structures.

Distribution: China, Myanmar, Thailand, Laos, Vietnam, Indonesia (Java).

Figures

FIGURE 26: Tillicera cleroides, aedeagus ventral aspect, aedeagus lateral aspect, pygidium and ventrite VI.

FIGURES 1–6: Dorsal habitus of Tillicera spp.. Fig. 1. Tillicera auratofasciata. Fig. 2. Tillicera aurivillosa. Fig. 3. Tillicera bibalteata. Fig. 4. Tillicera callosa n. sp.. Fig. 5. Tillicera cleroides. Fig. 6. Tillicera cleroides ab. bicoloricollis.