Tetragona clavipes (Fabricius, 1804) , Det. Camargo, 2001

Nogueira, David Silva, Oliveira, Favízia Freitas De & Oliveira, Marcio Luiz De, 2022, Revision of the Tetragona clavipes (Fabricius, 1804) species-group (Hymenoptera: Apidae: Meliponini), Zootaxa 5119 (1), pp. 1-64 : 6-59

publication ID	https://doi.org/ 10.11646/zootaxa.5119.1.1
publication LSID	lsid:zoobank.org:pub:F5425B3C-1877-4C3F-AE76-0CE37998437D
DOI	https://doi.org/10.5281/zenodo.6378563
persistent identifier	https://treatment.plazi.org/id/03EF87D4-FF90-FF9A-FF2D-FED3F083FE90
treatment provided by	Plazi
scientific name	Tetragona clavipes
status

Treatment

Tetragona clavipes View in CoL species-group

Diagnosis (worker): clypeus yellow, with or without brown maculations, and sometimes divided by a longitudinal brown line. Basal triangle of the clypeus, when present, from dark brown to black. Paraocular area with yellow maculation, not extending above middle tangent of the antennal socket (except in T. perangulata ). Forewing with apical half of membrane hyaline (with whitish microtrichia) and basal half slightly yellow (with yellowish-brown microtrichia). Setae on the sixth tergum about 1.5x–2x or more the length of the setae of fourth tergum, more abundant than in fifth tergum (except in T. korotaii sp. nov.). T2–T6 black with contrasting yellow band on apical third.

Key of T. clavipes species-group species (workers)

1. Hairs or setae inconspicuous and sparse on T2–T5 ( Fig. 20C View FIGURE 20 ), being longer on the apical area of the terga; central area of T6 only with short branched hairs, smaller than the diameter of the flagellum; mandibular condyle yellowish brown ( Fig. 20B View FIGURE 20 ); malar area and contour of the clypeus yellow ( Fig. 20B View FIGURE 20 ); only known from Suriname.......... Tetragona korotaii sp. nov.

- Hairs or setae conspicuous and abundant on T2–T5 ( Fig. 13C View FIGURE 13 ); central area of T6 with elongated simple setae, larger than the diameter of the flagellum; mandibular condyle black ( Fig. 13B View FIGURE 13 ); malar area and contour of the clypeus black ( Fig. 13B View FIGURE 13 ); Neotropical region and beyond.......................................................................... 2

2(1). Paraocular maculation blurry (smoky) yellow ( Fig. 18B View FIGURE 18 ) or not superiorly ( Figs. 16B View FIGURE 16 ), from the vicinity of the epistomal suture obliquely upwards towards compound eye in line or below the inferior tangent of the antennal socket.................. 3

- Paraocular maculation yellow from the vicinity of tentorial pit obliquely upwards towards compound eye in line or below the inferior tangent of the antennal socket.................................................................... 4

3(2). Paraocular maculation yellow, from the upper portion of the epistomal suture to compound eye to the middle tangent of the antennal socket ( Fig. 16B View FIGURE 16 ); T2 with a yellow longitudinal central line wider or equal to the diameter of the flagellum ( Fig. 16C View FIGURE 16 ); T3 with a thin longitudinal central yellow line; metatibia yellow with brown or black areas; found in Central America, Colombia and Ecuador.............................................................. Tetragona perangulata View in CoL

- Paraocular maculation smoky yellow superiorly, from the upper portion of the epistomal suture to compound eye to the inferior tangent of the antennal socket ( Fig. 18B View FIGURE 18 ); T2 without yellow longitudinal central line, or with a yellow line less wide than the diameter of the flagellum ( Fig. 18C View FIGURE 18 ); T3 without a thin longitudinal central yellow line; metatibia completely yellow; found in Brazilian Cerrado................................................................ Tetragona mourei sp. nov.

4(2). Metafemur, metatibia and metabasitarsus yellow ( Fig. 13A, D View FIGURE 13 ); nest entrance usually in the form of a narrow vertical gap with landing area made of rigid cerumen around ( Fig. 15A View FIGURE 15 ); found in eastern and southeastern Amazon, central and southeastern Brazil............................................................................ Tetragona quadrangula View in CoL

- Metafemur, metatibia and metabasitarsus brown with at least one black marking ( Fig. 1A, D View FIGURE 1 ); nest entrance very variable, usually in form of slit in tree trunk without projections ( Fig. 6A, B, D View FIGURE 6 ); with wide distribution from the eastern Andes to Uruguay................................................................................................................................................................................. Tetragona clavipes View in CoL