Navarretia leucocephala Benth. subsp. suksdorfii (Howell) L.A. Johnson & D. Gowen, 2022

Johnson, Leigh A. & Gowen, David, 2022, Restoring the original taxonomic concept for Navarretia minima (Polemoniaceae): a revised synonymy and new combination, Phytotaxa 576 (1), pp. 101-112 : 105

Navarretia leucocephala Benth. subsp. suksdorfii (Howell) L.A. Johnson & D. Gowen comb. et stat. nov.

≡ Navarretia suksdorfii Howell. Fl. N.W. Amer. 457 (1901) ≡ Navarretia minima Nutt. var. suksdorfii (Howell) Brand. Pflanzenr. 27[IV,250]: 164 (1907). TYPE:—[ U.S.A.] Washington: [Klickitat County], Falcon Valley, low grounds, 25 July 1883, Suksdorf s.n. (lectotype designated here: ORE 96503! [barcode OSC0000797], isolectotypes M barcode M-0185275 image!, NDG 40872 image!, NY barcode 00336965 image!, UC 185255!).

= Navarretia pilosifaucis St. John & Weitman. Proc. Biol. Soc. Washington 41(45): 196–197 (1928). TYPE:— [ U.S.A.] Washington: Whitman County, Rock Lake , rocky sandy soil, 23 June 1927, G . Weitman 430 (holotype: WS 43975 ! [barcode WS001522 ]) .

Misapplied names: Navarretia minima sensu Jepson (1943) , Mason (1951), Cronquist (1959, 1984), and others. Navarretia leucocephala subsp. minima sensu Day (1993a , b), Johnson (2012, 2019), and others.

Notes: Howell did not cite a specific collection of Suksdorf’s or a date, and Suksdorf collected frequently in ‘Falcon Valley’, the swampy plateau near the southeastern base of Mt. Adams in northwestern Klickitat County, Washington ( Weber 1944). We chose the specimen at ORE, which Howell would have worked from, as lectotype, which includes the exact wording found in the protologue, has been generally considered the type based on annotations, and is in good condition. When Brand reduced N. suksdorfii to a subspecies under N. minima , he noted the specimen he worked from was at the Berlin Herbarium. This specimen is presumably destroyed, but a duplicate at M bears an annotation label by Brand in 1905 using his then unpublished combination. The specimens at ORE, M, and NY have printed labels with no collection number, whereas those at NDG and UC are handwritten and include “167” before the species name, which is likely a referencing number rather than Suksdorf collection number. Navarretia pilosifaucis was considered a synonym of N. intertexta by Mason (1951) and of N. propinqua by Cronquist (1959), but has been largely ignored otherwise. Curiously, the holotype of N. pilosifaucis belongs in N. leucocephala subsp. suksdorfii as defined here, having white pollen and elongated, flattened, flexuous outer bracts that exclude it from both N. intertexta and N. propinqua . However, the single cited paratype, collected a year later in the same general vicinity (WS 48906! [barcode WS001942]), is clearly N. intertexta and may have been the source of Mason’s and Cronquist’s determination.