Laccomimus amazonas, Toledo, Mario & Michat, Mariano C., 2015

Toledo, Mario & Michat, Mariano C., 2015, Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae), Zootaxa 3990 (3), pp. 301-354: 315-316

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Laccomimus amazonas

sp. n.

Laccomimus amazonas  sp. n.

( Figs 10View FIGURES 8 – 13; 24–27; 38; 47–48; 62; 75; 88; 110; 124; 137)

Type locality. Venezuela, Amazonas  State, Communidad caño Gato on Río Sipapo, 4 ° 58.838 'N, 67 ° 44.341 'W, 95 m.

Type material. Holotype male ( MIZA), dissected by us; genitalia mounted in DMHF on plastic label pinned under the holotype “ Venezuela, Amazonas  State, 4 ° 58.838 'N, 67 ° 44.341 'W, 95 m, Comunidad caño Gato on Rio Sipapo, 16.I. 2009, leg. Short, Miller, Camacho, Joly & García, VZ09-0116- 01X, along stream” [printed], “SM0843379 KUNHM-ENT ” [printed, together with barcode], “ HOLOTYPE Laccomimus amazonas  Toledo & Michat” [red, rectangular, printed label].

Paratypes (33 exs, all bearing red, printed label “ PARATYPE Laccomimus amazonas  Toledo & Michat”): Venezuela: same data as holotype (30 exs VAIP, MTP, MCMC, NMW); “ Amazonas  State, 4 ° 55.849 'N, 67 ° 44.645 'W, 87 m, stream along Rio Sipapo, 16.I. 2009, leg Short, Garcia, Camacho, Miller & Joly, VZ09-0116- 02X, stream habitats” (2 exs VAIP); “ Amazonas  State, 5 ° 20.497 'N, 67 ° 45.358 'W, 72 m, ca. 12 km N Samariapo, small stream at rd. x-ing, 6.I. 2006, AS-06-013, leg. A.E.Z. Short” (1 ex. VAIP). Each specimen bearing second VAIP label with its own barcode and catalogue number.

Diagnosis. Easily recognised by the following combination of characters: lobes of metacoxal processes ending in long sharp spine in both sexes; elytra dark with distinct pale markings; male with very large stalked suction palettes and outer claw of forelegs scimitar-shaped. Aedeagal features distinctive, median lobe expanded on apical fourth ( Fig. 88View FIGURES 88 – 90 a) and left paramere triangular, gradually tapering apically ( Fig. 88View FIGURES 88 – 90 b).

Description. Habitus ( Fig. 62View FIGURES 59 – 66). TL: 2.2–2.4 mm; MW: 1.15–1.2 mm; (holotype: TL: 2.3 mm; MW: 1.15 mm); A-TL: 2.4 mm; A-MW: 1.2 mm (NUM: 16 exs); drop-shaped, strongly but regularly narrowed posteriorly. Smooth; elytra with faint silky sheen.

Colour. Head testaceous, often darkened behind eyes. Pronotum testaceous, with darker transverse marking along posterior margin, normally divided into two parts. Elytra chestnut-brown; each elytron with three distinct yellow markings: transverse, subbasal band reaching lateral side; submedial faded spot close to lateral side; subapical marking not reaching apex. Underside reddish-yellow. Antennae and mouthparts testaceous. Legs reddish-yellow.

Structures and sculpture. Head shiny, with faint microreticulation and scattered fine punctures on disc.

Pronotum smooth, sculpture as on head, with faint silky sheen. Elytra elongate, with subapical sinuation poorly developed and elytral apex narrow, almost acute; surface smooth, with faint silky sheen; fine and scattered punctures are visible in most specimens but no grooves occur, in contrast to preceding species. Underside shiny, lacking silky sheen. Prosternal process broad, slightly elevated especially close to rounded tip ( Fig. 24–27View FIGURES 20 – 25View FIGURES 26 – 31). Lobes of metacoxal processes ending in long sharp spine in both sexes ( Fig. 38View FIGURES 38 – 43).

Male. Last abdominal ventrite tectiform with lateral sides almost rectilinear. Stalked suction palettes on pro- and mesotarsi very broad, broader than in any other Laccomimus  species ( Figs 47–48View FIGURES 44 – 51; compare with Fig. 44View FIGURES 44 – 51 and Figs 49View FIGURES 44 – 51; 51). Outer claw of forelegs scimitar-shaped ( Fig. 75View FIGURES 67 – 83). Median lobe of aedeagus, in left lateral view ( Fig. 88View FIGURES 88 – 90 a), resembling that of L. variegatus  , with leaf-like expanded apical fourth but visibly narrower, especially before apical expansion. In dorsal view ( Fig. 110View FIGURES 107 – 115) slightly turning to right, gradually tapering from base to apex, with nipple-like tip; dorsal lamina sinuate and ejaculator groove narrower on basal half and wider near apex. Left paramere distinctive ( Fig. 88View FIGURES 88 – 90 b): triangular, shorter than in any other Laccomimus  species, and more gradually tapering apically.

Female. Last abdominal ventrite elongate, with fore margin broadly rounded. Lateral tergites thin and partly membranous. Gonocoxal blade with single medial denticle; ramus with large and spaced dentation; first tooth, following apical hook, very small with respect to second tooth.

Distribution. Venezuela, Amazonas  State ( Fig. 124View FIGURES 122 – 125).

Habitat. Collected in running waters ( Fig. 137View FIGURES 136 – 138).

Etymology. The specific epithet relates to type locality. It is treated as a noun in the nominative singular.

Remarks. See under L. variegatus  .


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