Cephennodes (Cephennodes) annapurnaensis, Jałoszyński, 2017

Jałoszyński, Paweł, 2017, Ant-like stone beetles on the roof of the world. Cephenniini of Nepal and Bhutan (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 4349 (1), pp. 1-120 : 51

Cephennodes (Cephennodes) annapurnaensis sp. n.

( Figs 100 , 110 , 136–139 , 147 )

Type material. Holotype: NEPAL (Myagdi District): ♂, " Nepal, Annapurna / South Himal / Chiule - Tatapani / 2400 m 17 V 2001 / J. Schmidt [white, handwritten in black ink] ( MHNG).

Diagnosis. BL 1.35 mm; body moderately stout, with elytra weakly narrowing behind the broadest site; distal portion of metatibia strongly thickened, with oval subapical porous field much longer than broad, lacking one long bristle within setal fringe of apical margin; abdomen in ventral view with sternite II bearing broad microserrate and emarginate at middle carina along its posterior margin, sternite III with broadly rounded median lobe; apical portion of median lobe of aedeagus rounded, its longitudinal axis forming about 30°-angle with long axis of aedeagus; apical projection with strongly concave distal margin and subtriangular apex.

Description. Body of male ( Fig. 100 ) brown, with shallow constriction between pronotum and elytra, strongly convex, covered with yellowish setae; BL 1.35 mm.

Head subtrapezoidal, HL 0.20 mm, HW 0.33 mm; vertex and frons confluent, evenly and weakly convex; supraantennal tubercles small and weakly elevated; eyes large, strongly convex, coarsely faceted. Punctures on frons and vertex small but very distinct, sharply marked, densely and unevenly distributed, sparser at middle and denser on sides, those in middle of frons between eyes separated by spaces 1–2 × as wide as diameters of punctures; setae short, moderately dense, suberect to erect. Antennae relatively short and moderately slender, AnL 0.68 mm, pentamerous club indistinctly delimited; antennomere I 1.4 × as long as broad; II slightly narrower but not shorter than I, about 1.7 × as long as broad; III slightly narrower than II and much shorter, about as long as broad, IV–VI similar in shape, each slightly longer than III, about 1.5 × as long as broad, VII slightly broader and longer than VI, about as long as broad; VIII slightly broader but shorter than VII, distinctly transverse; IX and X each distinctly larger than preceding one, each distinctly transverse, XI clearly broader than X, about as long as IX– X together, about twice as long as broad.

Pronotum in dorsal view nearly semicircular, equally broad between base and anterior third, PL 0.40 mm, PW 0.64 mm; anterior margin broadly and weakly rounded; lateral margins not serrate, strongly rounded in anterior half, straight in posterior half; posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral carinae narrow and fused with lateral margins; antebasal pits small and very shallow, the deepest site of each pit located much closer to posterior than to lateral margin of pronotum. Punctures on disc very small and shallow, inconspicuous; setae dense, short, suberect.

Elytra short, oval, broadest between middle and anterior third, EL 0.75 mm, EW 0.68 mm, EI 1.11. Humeral calli weakly elevated; subhumeral lines distinct, equal to about 0.4 EL, each developed as a groove between more convex humeral region and less convex adsutural area; basal pit on each elytron large and connected to short, slightly arcuate impression extending posteromesally; apices of elytra separately rounded. Punctures on elytra more distinct than those on pronotum but superficial, diffuse and inconspicuous; setae about as dense and short as those on pronotum.

Hind wings not studied.

Metaventrite with large and sharply delimited postmesocoxal impressions, median part covered with very fine punctures.

Hind legs with strongly modified tibiae ( Fig. 110 ); in dorsal view lateral (external) tibial margin weakly recurved and spatulate portion of tibia gradually narrowed distally, with rounded, not angulate external margin; dorsodistal porous field much longer than broad; distal fringe of setae lacking one long bristle.

Abdomen ( Figs 136–137 ) with sternite II bearing a broad microserrate and emarginate at middle carina along its posterior margin, sternite III bearing broadly rounded, short median lobe.

Aedeagus ( Figs 138–139 ) moderately slender; AeL 0.20 mm, apical portion of median lobe rounded, its longitudinal axis forming about 30°-angle with the long axis of aedeagus; apical projection with deeply concave distal margin, its apex subtriangular, not delimited as a well-defined lobe; parameres long and slender, not exceeding apex of median lobe, each bearing one apical seta.

Female. Unknown.

Distribution ( Fig. 147 ). Central Nepal.

Etymology. Locotypical, after Annapurna.

Remarks. This species is most similar to C. thakanus ; see remarks for the latter species.

The locality Tatapani written in the label is more frequently spelled Tatopani.

Figures

FIGURES 94 – 101. Dorsal habitus of Cephennodes (s. str.) dolakhanus s sp. n. (94), C. (s. str.) manangensis sp. n. (95), C. (s. str.) martensi sp. n. (96), C. (s. str.) paramartensi sp. n. (97), C. (s. str.) monolaminatus sp. n. (98), C. (s. str.) thakanus sp. n. (99), C. (s. str.) annapurnaensis sp. n. (100), and C. (s. str.) parbatensis sp. n. (101).

FIGURES 102 – 111. Right metatibia of male in dorsal view. Cephennodes (s. str.) dolakhanus s sp. n. (102, 104), C. (s. str.) manangensis sp. n. (103, 105), C. (s. str.) martensi sp. n. (106), C. (s. str.) paramartensi sp. n. (107), C. (s. str.) monolaminatus sp. n. (108), C. (s. str.) thakanus sp. n. (109), C. (s. str.) annapurnaensis sp. n. (110), and C. (s. str.) parbatensis sp. n. (111). 104 – 111 not to the same scale.

FIGURES 128 – 143. Abdomen of male in right lateral (128, 132, 136, 140) and ventral (129, 133, 137, 141) views and aedeagus in ventral (130, 134, 138, 142) and lateral (131, 135, 139, 143) views. Cephennodes C. (s. str.) monolaminatus sp. n. (128 – 131), C. (s. str.) thakanus sp. n. (132 – 135), C. (s. str.) annapurnaensis sp. n. (136 – 139), and C. (s. str.) parbatensis sp. n. (140 – 143). Abbreviations: aml, apex of median lobe; ap, apical projection.

FIGURES 146 – 147. Distribution of Cephennodes (s. str.) monolaminatus sp. n. (146, circle), C. (s. str.) thakanus sp. n. (144, triangle), C. (s. str.) annapurnaensis sp. n. (147, circle), C. (s. str.) parbatensis sp. n. (147, triangle).