Dianthus ciliatus subsp. dalmaticus Čelakovský (1885: 189) Hayek (1924: 246) (Celakovsky)

Terlević, Ana, Ljubičić, Ivica, Tokić, Petar, Rešetnik, Ivana, Shuka, Donald, Siracusa, Giuseppe & Bogdanović, Sandro, 2024, Morphological and nomenclatural notes on Dianthus ciliatus subspecies (Caryophyllaceae), Phytotaxa 661 (3), pp. 213-234 : 222

3. Dianthus ciliatus subsp. dalmaticus Čelakovský (1885: 189) Hayek (1924: 246) ≡ Dianthus dalmaticus Čelakovský

(1885: 189)

Lectotype (designated here):— MONTENEGRO. Flora von Dalmatien. In apricis saxoisis prope Cattaro, July 1877, Studniczka s.n. ( PR 836441 !, Fig. 7 )

= Dianthus ciliatus Gussone (1825: 5) var. cymosus Visiani (1852: 162) .

Lectotype (designated here):— MONTENEGRO. In apricis circa Ragusa et Cattaro, s.d., Visiani s.n. ( PAD-HD05071 !).

Notes: —In 1885 Čelakovský described this species based on herbarium specimens collected in 1877 by K. Studniczka in Kotor (Cattaro) from Montenegro ( Čelakovský 1885). Three relevant herbarium sheets were found at PR that are in relation to the name Dianthus dalmaticus . There is no Čelakovský’s identification in the specimens, but only the annotation “non est” crossed out the name Dianthus ciliatus Guss. β cymosus on the sheet PR 836441. This note is in Čelakovský’s handwriting and it is evident that he saw this specimen and cited it in the protologue, so we chose the specimen PR 836441 as lectotype for the name D. dalmaticus ( Fig. 7 ). The second specimen (PR 86229) is a duplicate and here considered as isolectotype, which was however incorporated in PR much later, in the last third of the 20 th century (Otakar Šída, pers. comm.). The third specimen (PR 836442) could have a relation to the specimen or name mentioned by Čelakovský in the protologue, since Čelakovský nearly surely studied this specimen from Ragusa (Dubrovnik in Croatia) as well.

Habitat and phenology: — Dianthus ciliatus subsp. dalmaticus ( Fig. 2C, D ) occurs in the eu-Mediterranean and sub-Mediterranean zone of the eastern Adriatic coast and inland, occupying colder habitats, on screes, walls, rocky places, along roads, in fissures of rocky places and on deciduous forest margins. The flowering period is from mid-July to mid-August, fruiting in late August to September.

General distribution: — Croatia, Montenegro, Bosnia and Herzegovina ( Fig. 1 ).

Other original material examined: — MONTENEGRO. Flora von Dalmatien. In apricis saxosis prope Cattaro, July 1877, Studniczka s.n. (isolectotypes: PR 86229 !, K 000725360 !, BEOU!; BP HNHM-TRA 00183940 !) .

Figures

FIGURE 1. Localities of sampled populations of taxa belonging to the endemic amphi-Adriatic Dianthus ciliatus used for morphometric analyses (n = 27 populations). Type localities are indicated with thick black symbol outlines.

FIGURE 2. Morphological details of corolla and calyx in Dianthus ciliatus subsp. ciliatus (A, B), D. ciliatus subsp. dalmaticus (C, D), and D. ciliatus subsp. medunensis (E, F, G). Habitus of D. ciliatus from Popoli (locus classicus) in Abruzzo, Italy (H), and floral quantitative characters analysed (I). Photographs by S. Bogdanović.

FIGURE 7. Lectotype of the name Dianthus dalmaticus (PR836441!).