Isohypsibius sattleri ( Richters, 1902 ), 2017

Kaczmarek, Łukasz, 2017, Annotated zoogeography of non-marine Tardigrada. Part IV: Africa, Zootaxa 4284 (1), pp. 1-74 : 39-40

111. Isohypsibius sattleri ( Richters, 1902) sensu lato [T]

Hypsibius sattleri ( Richters) 1902 (da Cunha & do Nascimento 1964) Isohypsibius bakonyiensis (Iharos, 1964) (Binda 1971, 1984) Isohypsibius bakonyiensis Iharos, 1964 ( Pilato & Pennisi 1976) Hypsibius bakonyiensis Iharos (Iharos 1978)

Isohypsibius bakonyiensis , I. sattleri ( Ramløv & Kristensen 1985) Isohypsibius sattleri ( Richters, 1902) (Pilato et al. 2003) Terra typica : Germany (Europe)

Angola:

• 07°47′S, 15°01′E; 650 m asl: Uíge Province, near Carmona [Uíge], Estate at Pumba Loge, mosses. da Cunha & do Nascimento (1964)

Democratic Republic of Congo:

• 00°04′S, 29°18′E; 1,950 m asl: North Kivu Province, outskirts of Lukanga, moss ( Brachythecium velutinum ). Pilato et al. (2003)

Libya:

• 32°29′N, 20°50′E; 350 m asl: Marj District, Barce [Marj], moss. Pilato & Pennisi (1976)

Morocco:

• 32°56′N, 05°40′W; 1,700 m asl [850 m asl]: Meknès-Tafilalet Region, Khénifra, moss. Binda (1971)

• 32°25′N, 05°13′W; 1,750 m asl: Meknès-Tafilalet Region, S of Azrou , cedar forest, moss on basalt and limestone boulders in the forest glade and moss on soil and fallen branches and stems (3 samples). Ramløv & Kristensen (1985)

• Republic of South Africa:

• 34°01′S, 23°55′E; 200 m asl: Eastern Cape Province, Tzitzikama [Tsitsikamma National Park], moss. Binda (1984)

Tunisia:

• 36°14′N, 08°46′E; 1,100 m asl: Kef Governorate, Kef Governorate, 6 km NE of El Kef, Djebel Eddir Mts., surroundings of Ferme Shitta , moss and lichen on rock. Iharos (1978)

• 36°46'36''N, 08°42'45''E; 784 m asl: Jendouba Governorate, Beni M'tir, urban, moss on soil. Gąsiorek et al. (2017)

Record numbers. Angola: 1, Democratic Republic of Congo: 1, Libya: 1, Morocco: 2, Republic of South Africa: 1, Tunisia: 2; total: 8.

Remarks. This member of the tuberculatus -group has a history of major taxonomic confusion with I. bakonyiensis (now the junior synonym), but was re-described from a Richters’ 1905 slide in the Senckenberg Museum ( Germany) by Dastych (1990). Nevertheless, the literature-based, supposed, cosmopolitan distribution ( McInnes 1994), requires major taxonomic re-evaluation of all specimens attributed to I. sattleri .