Kutara Distant, 1908

Xu, Deliang, Dietrich, Christopher H., Webb, M. D. & Zhang, Yalin, 2020, A new species of the Oriental leafhopper genus Kutara Distant (Hemiptera: Cicadellidae: Deltocephalinae) from Vietnam, Zootaxa 4772 (3), pp. 585-592 : 585-586

Kutara Distant

Kutara Distant, 1908: 308 . Type species: Kutara brunnescens Distant , by original designation.

Diagnosis. Moderately robust leafhoppers with coloration ochraceous to brown with a black transverse spot ante- riorly on crown; antennal pedicel dark brown. Head wider than pronotum with two transverse carina on anterior margin with a sulcus between; anterior tentorial branches L-shaped; crown of similar length throughout width, posteriorly elevated, anterior and posterior margin nearly parallel; ocelli situated on sulcus on anterior margin of head, distant from eyes; face rather flat and broad; frontoclypeal suture present; anteclypeus with apex slightly broader than base, sides concave; lora conspicuously broader than anteclypeus; gena prominent; antennae long, more than half body length, arising near upper corners of eyes; antennal ledge strong and oblique. Pronotum with transverse striations and dense punctures, anterior margin arched, lateral margin short, hind margin concave. Scutellum similar in length to pronotum with transverse depression distinct. Forewing with four apical cells and three subapical cells, inner subapical cell closed, cross vein present between claval suture and A1 and between A1 and A2, appendix narrow. Fore femur with anteromedial (AM1) seta present and more additional proximal macrosetae; intercalary (IC) row with fine setae apically, anteroventral (AV) row with short or longish microsetae on ventral margin. Protibia dorsal surface rounded with 4+4 or more dorsal setae ( Fig. 1G ). Metafemur with apical setal formula 2+2+1 ( Fig. 1H ).

Male genitalia: pygofer basolateral membranous cleft present; with an internal posterior ledge; lateral lobe caudally truncate or broad to narrowly rounded, without apical process or with a ventrocaudal process, with several macrosetae and microsetae. Valve triangular or pentagonal. Subgenital plate triangular without macrosetae, with fine hair-like microsetae arising from lateral margin, irregularly arranged. Style with apical process of variable length and shape, robust; preapical lobe well developed or not. Connective usually Y-shaped, stem similar in length to arms, sometimes bifid distally, ventral margin angled at mid-length in lateral view. Aedeagus with dorsal apodeme well developed; shaft slender and elongate, curved dorsad with a subapical prolongation arising from ventral margin exceeding apex of shaft, with or without a pair of apical processes, rarely with pair of basal processes ( K. nigrifasciata Kuoh ) or lateral spines ( K. spinifera Zhang & Chen and K. nigrifasciata Kuoh ); gonopore subapical on dorsal side; anal tube large, sclerotized laterally and dorsally.

Female genitalia: pygofer with numerous setae; ovipositor conspicuously surpassing apex of pygofer; first valvulae with strigate texture dorsally; second valvulae with tooth-like prominence at mid-length of dorsal margin, with dorsal teeth restricted to approximately apical 1/3.

Distribution. Palaearctic, Oriental and Australian Region.

Remarks. Kutara , together with Drabescoides Kwon & Lee and Papuakutara Lu, Webb & Zhang , form the Kutara group. The group is distinguished by the head marking with a transverse medial spot or band and shape of the head ( Fig. 1A ) and short vertex with posterior margin elevated ( Fig. 1E ). It can be distinguished from Drabescoides by the narrower stem of the connective and male pygofer side without posterior process and from Papuakutara in having a cross vein between the claval veins of the forewing, and the connective ventral margin angled at mid-length in lateral view and aedeagal shaft usually extended beyond the gonopore and without a basal process in the male genitalia.

Figures

FIGURE 1. Kutara trispinosa Xu & Zhang sp. nov., male; A, Habitus, dorsal view; B, Head, pronotum and scutellum, dorsal view; C, Face, ventral view; D, Head, dorsoanterior view; E, Lateral view; F, Male abdomen, ventral view; G, Fore femur and tibia, anterior view; H, Apex of hind femur, dorsal view.