Veronica prostrata Linnaeus (1762: 17)

Rojas-Andrés, Blanca M. & Martínez-Ortega, Montserrat, 2016, Taxonomic revision of Veronica subsection Pentasepalae (Veronica, Plantaginaceae sensu APG III), Phytotaxa 285 (1), pp. 448-450 : 448-450

9. Veronica prostrata Linnaeus (1762: 17) . ( Fig. 15 )

Stems 5–25 (40) cm long, decumbent, the vegetative ones prostrate forming ± compact lawns, usually densely covered by fine and short eglandular hairs 0.1–0.4 mm long, incurvate, appressed and antrorse, not concentrated in 2 opposite lines along the stem; apical shoot bearing 1–3 pairs of leaves. Leaves (3) 5–25 (32) × (1) 2–8 (15) mm, the basal and medium ones similar, oblanceolate to narrowly elliptic, elliptic or ovate, usually ± truncate or cuneate at the base, crenate or serrate to rarely subentire, often subrevolute, the adaxial face subglabrous or sparsely covered by very short hairs (0.02) 0.05–0.15 (0.2) mm long, principally along the margin and the primary vein, the abaxial face covered by the same type of hairs, tomentulose, shortly petiolate; apical shoot leaves opposite, narrowly elliptic to lanceolate or linear, serrate to subentire. Racemes axillary, opposite, bearing (4) 25–30 (60) flowers usually very densely arranged (compact appearance); peduncles 1–5 cm long, covered by the same type of indument as that of the stem; bracts (1) 1.5–4 (7) mm long, narrowly elliptic, entire, glabrous or subglabrous, commonly with very short hairs (similar to those of the leaves) along the margin; pedicels 1–3 (6.5) mm long. Calyx (2) 2.5–4 (5) mm long; (4) 5 sepals, linear-lanceolate, most usually shorter than the capsule, glabrous or rarely those of the lower flowers with a few very short eglandular hairs. Corolla (4) 6–8 (11) mm in diameter, light blue, pale blue, less frequently white or pink. Capsule (2) 3–5 (5.5) × (2) 2.5–4.5 (5) mm, usually slightly longer than wide, widely obovate to obcordate or suborbicular, rounded at the base, emarginate, sinus up to 0.5 (0.7) mm depth, glabrous or exceptionally puberulent, with eglandular hairs. Style 3–5 mm long. Seeds 0.9–1.3 × 0.6–1.1 mm, ca. 5–16 per capsule ( Kulpa 1968).

Chromosome Number: —2 n = 16.

Habitat: —Arid meadows with little soil material, shrub-steppe, dry forest or along stony slopes, preferably on limestones; 0–1,500 (1,900) m.

Distribution: —C and E Europe and Asia (Siberia); Austria, Bulgaria, Czech Republic, SE France, NE Germany, Hungary, N and C Italy, Poland, Republic of Macedonia, Romania, Russia (Siberia), Serbia, Slovakia, Slovenia, SW Switzerland and Ukraine ( Fig. 16 ). According to Borissova (1955), it also occurs in Lapland and the

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Baltic region, European Russia, Belarus, Moldavia, and Western Siberia (Upper Tobol and Irtysh). According to Kosachev (2003) it occurs in Central Siberia (Altai Krai, Irkutsk Oblast, Krasnoyarsk region, and Republic of Khakassia) and the Transbaikal region.

Phenology: —Flowering from (March) April to June (September).

Specimens Examined: — AUSTRIA. Burgenland: Neusiedlersee, Ostufer, 19 May 1929, Gufodontis s.n. (W); Leithagebirge, entre Donnerskirchen y Kaiser reiche Franz Josef-Warte, prados secos, vegetación panónica, ca. 250 m, 28 May 2001, M. Martínez Ortega MO1055 (SALA124616); Lower Austria: Unterloiben, Strassenboschung, May 1970, Polatschek s.n. (W); auf dem Grossen?anzel der hoher Wand bei Wiener (illegible word), 28 May 1917, Hayek s.n. (B); Braunsberg b. Hainburg, 10 May 1914, Ronniger s.n. (W); Baden, Mitterberg, 5 May 1997, Krendl s.n. (W); Weinviertel, zwischen Gedersdorf und Saubügel, 3–4 Km EN Krems, trockenrasen, Gebüsche, ca. 200–300 m, 5 May 1979, Krendl s.n. (W); Wr. Neustädter, Steinfeld, 1.2 km W Haschendorf an der Geländstufe (“Wagram”) bei Kote, 233 m, Malicky s.n. (WU); in collibus apricis aridis, ad pedem montis Geissberg prope Perchtoldsdorf, solo cacareo, 300–350 m, Pernhoffer s.n. (B, K, PR, W); Voralpen, Losenheim, Kalk, 600–700 m, Krendl s.n. (W); Wien, Leopoldsberg, 23 May 1908, Watzl s.n. (WU); Kalke Weide am Bierhäuselberg bei Rodaun, 16 May 1908, Watzl s.n. (WU); Wien, Wiesen auf dem Gaisberge bei Rodam, 22 May 1900, Krebs s.n. (B); Perchtoldsdorf, June 1923,?eny s.n. (K); Stratzing, S-exp. Weinbergböschung, 350 m, 20 May 1995, Kleesald s.n. (WU); Thermenlinie, Wienersdorf bei Traiskirchen, trockenwiese, 200 m, 30 April 1994, Till s.n. (WU); Wiesen auf den Anhöhen am Kreuz, 19 May 1912, Wettstein s.n. (WU); kaiserwasser, 14 May 1908, Watzl s.n. (WU); Marchegg, Hochwasserdamm wenig oh. Schloss, 21 April 1990, R. Hand s.n. (B100327559); Falkenstein, junto a las ruinas del castillo, en prados calizos, 11 June 2001, M. Martínez Ortega MO1337 (SALA124615); Hainburg an der Donau Braunsberg, prados sobre calizas en la parte alta, 350 m, 48º09'16.8''N, 16º57'31.5''E, 29 July 2014, X. Giráldez , M. M. Martínez Ortega, B. Rojas Andrés & N. López González NLG 135 (SALA155858); Hainburg, Braunsberg, prados sobre calizas, 3 June 2001, M. Martínez Ortega MO1302 (SALA124614); Rohrendorf bei Krems, Saubühel, prados secos en un merendero, 286 m, 48º25'40.3''N, 15º39'22.1''E, 22 July 2011, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés BR182 (SALA149040); Styria: Föllig, 25 June 1997, Neumann s.n. (W); Tirol: Sandskügel nach der Weiherburg bei Innsbruck, ¿Kern? s.n. (WU); Oberinntal, Fliess-Gh. Neuer Zoll, Trockenrasen, Moränenschutt, Silikat, 850–1073 m, 6 June 1970, Polatschek s.n. (W); Upper Austria: Alluvialboden der Traunebene (Welserhaick) ¿nachst? hing (illegible word), nicht selten, gessellig, Strobl s.n. (K); Frauenstein bei Mo???ig, 27 April 1887, Engelhardt s.n. (B); ¿Woppl? bei Linz, an Wärmeplätzen hinter dem ¿Waldfriedhof?, April–May 50, Baschant s.n. (B); (illegible group of words) bei Linz, 23 (illegible month) 1885, illegible collector (PR); Welser Heide, Braunstingel s.n. (WU); Vienna: Trockener wiesen in Prater bei Wien, 21 April 1920, Zerny s.n. (W); Wien, auf kalkfelsen bei?odling, 21 May 1899, Krebs s.n. (E); Vindobonae, Rohle s.n. (B); Vienna, Donauinsel, on new roadside, 19 April 1988, Bowen s.n. (RNG); Kalksburg, en la subida desde la calle "In der Klausen", 338 m, 48º08'28.0''N, 16º14'50.2''E, 24 July 2011, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés MO6040 (SALA149316); unknown or dubious state: b. Le???burg, 18 May 84, Karny s.n. (B); Kalten?entgeben, f??fburg, 420 m, 20 May 1924, Berg s.n. (B); Mamersdarf b. Wien, May 1923, Meebold s.n. (K).

BULGARIA. Dobrich: Kavarna, in saxosis ad oppid, 19 May 1965, Weber s.n. ( PR) ; Pazardzhik: In graminosis siccis ad Belovo, 1911, Urumoff s.n. ( WU); Ruse: In graminosis siccis ad Rustzuk, 1901, Urumoff s.n. ( WU) ; Shumen: In graminosis siccis ad Preslav, 1881, Urumoff s.n. ( WU); In graminosis siccis ad Preslav , 1901, Urumoff s.n. ( WU); Madara , junto al pueblo, prados secos sobre calizas, 162 m, 43º16'21.1''N, 27º06'23.8''E, 21 June 2009, X. Giráldez , M. M . Martínez Ortega, B. Rojas Andrés & M . Santos Vicente MS1239 ( SALA149317 ) ; Sofia (province): Entre Dragoman y Golemo Malovo, en las laderas que están frente al humedal, claros abiertos en ladera sobre calizas, 731 m, 42º56'26.2''N, 22º58'04.8''E, 16 June 2009, X. Giráldez , M. M . Martínez Ortega, B. Rojas Andrés & M . Santos Vicente MO4584 ( SALA149314 ) ; Varna: hills north of Varna, 14 May 1924, Gilliat Smith s.n. ( K); Varna, May 1876, unknown collector ( K); Prope Varna, in graminosis ad ¿Francka?, 16 May 1907, Schneider s.n. ( W); Prope Varna, in desertis ad ¿Francka?, 16 May 1907, Schneider s.n. ( BM); Varna district , 24 May 23, Gilliat Smith s.n. ( K); Prope Varna, in desertis ad (illegible word), 16 May 1907, Schneider s.n. ( K); dubious province: Bei Tirnovo, 1896, Urumoff s.n. ( WU) .

CZECH REPUBLIC. Central Bohemia: Srbsko, colina Doutnác , en prados calizos de borde de bosque, 10 June 2001, M . Martínez Ortega MO1331 ( SALA124619 ); Slany, na (two illegible words), 17 May 1886, Bilek s.n. ( W); ¿ Zubist ? bei Prag, 12 May 1884, Lu ??? s.n. ( W); Zavist bei Prag, 12 May 1884, L ?una??? s.n. (K); Liberec: Habstein [=Jestřebí], May 1911, Weder s.n. ( E) ; Olomouc: Olmütz, abh. des Ko ???, May 1932, ¿Laus? s.n .

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Phytotaxa 285 (1) © 2016 Magnolia Press • 51 (BCF41368); Prague: distr. Praha, (reg. Ceský Kras), in nemore mixto ad cotam 377 pr. pag. Solopysky apud opp. Cernosice, 22 May 1943, Dostál s.n. ( PRC); South Moravia: Moravia, montes Pavlovski Kopce, May 1933, F. Weber s.n. (WU); Entre Klentnice y Mikulov, Tabulová, prados secos sobre calizas con vegetación panónica, 376 m, 48º50'08.4''N, 16º38'27.7''E, 23 July 2011, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés BR186 (SALA149311); Entre Mikulov y Klentnice, prados calizos secos, en lugares con vegetación pannónica, 24 June 2001, M. Martínez Ortega MO1445 (SALA124617); Mikulov, junto a la frontera con Austria, Šibeničník, prados secos con vegetación panónica, 236 m, 48º47'25.5''N, 16º37'56.5''E, 23 July 2011, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés BR184 (SALA149310); Mikulov, Svatý kopeček, en la parte de arriba junto a la iglesia, prados secos y rocosos sobre calizas, 350 m, 48º48'24.8''N, 16º38'46.6''E, 23 July 2011, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés BR189 (SALA149257); Vysočina, distr. Třebíč: Mohelno, 7 July 1937, Stearn s.n. (K).

FRANCE. Hautes-Alpes: plateau de Bayard du cote de (illegible word), 23 May 1897, Faure s.n. ( MPU); Briançon, mont Genevre, prairies incultes, 1900 m, 29 May 1868, unknown collector ( K); ¿Charance?, 17 May 1862, Borel s.n. ( BM); Briançon, mont Genevre, prairies, incultes, 1900 m, 29 May 1868, unknown collector ( BM); col Bagard près Gap, June 1902, illegible collector (PRC) ; Mt. Seuse , 29 July 1862, Borel s.n. ( K); La grave, schistes, June 1860, Ozanon s.n. ( BM); les praires de ¿ Sèuse?, 15 June 1872, Reverchon s.n. ( E); La Grave, lieux arides, a 1500 m, June 1869, Reverchon s.n. ( K); Gap, col du Noyer, entre Le Noyer y L'Enclus, prados subalpinos sobre calizas, 1671 m, 44º41'34.7''N, 05º59'13.2''E, 14 June 2012, X. Giráldez , M. M . Martínez Ortega & B . Rojas Andrés BR219 ( SALA149313 ); Gap, col de Bayard, hacia el col de Glezié, prados calizos secos, 1403 m, 44º37'13.5''N, 06º04'06.0''E, 9 July 2011, X. Giráldez , M. M . Martínez Ortega & B . Rojas Andrés MO6037 ( SALA149315 ); Isère : Coteaux de Crémieu , Boullu s.n. ( BM); Savoie : La Côte de l’Ile près Mo ??ne, prairies seches, 24 May 1891, Perrier s.n. ( G); unknown or dubious department: paturages près St. Germain, May 1864, Rostan s.n. ( BM, G, K) .

GERMANY. Berlin: Glienicker Volkspark Wiese, 30 May 1970, Damboldt s.n. ( B) ; Köpenick, auf sonnigen Hügeln , 20 May 1913, Gross s.n. ( B) ; Berlin , unknown collector ( B) ; Collines incultes, sèches et stèriles aux bords des champs sur le diluvium de la plaine autour de Berlin ( Prusse ), fl. 18 May 1847, fr. 14 June 1852, Bolle & F. Koernike s.n. ( K, PRC, W); pr. Berolinum (Fuchsberge), 1841, unknown collector ( P00208676 ) ; Brandenburg: Brandenburg , 15 May 1887, Jachan s.n. ( W) ; Potsdam in locis apricis, sabulosis ad graminosis, ad fin m. May 1905, Garllorph s.n. ( BM) ; Bei Potsdam , (two illegible words), May 1882, Bornmüller s.n. ( B) ; Saxony: Dresden, Grahard s.n. ( WU) ; An Rainen bei Dresden, end April & May, unknown collector ( K, W, B) ; in coll. prope Meissen , 25 May 1892, Hofman s.n. ( PRC, W, WU) ; Saxony-Anhalt: Altmark (illegible locality), 21 May 1931, illegible collector ( W) ; Gutenberg bei Halle , May 1845, unknown collector ( B) ; Kleinwangen bei Nebra , in lückingen Steppenrasen auf den Rogensandsteinbänken der Steinklöbe, 27 May 1956, Hempel s.n. ( B) ; Thuringia: Thüringen, sonnige Hänge , bei Erfurt, 8 June 1889, Rudolph s.n. ( W) ; Thüringen, Gyp ?felsen b. ???hauren, 11 June 1877, ¿ Shafer? s.n. ( WU) ; unknown state: Halle Kerbe b. Brachland flachgründig. Zechstein. -verwitterung, 16 May 1956, Neumann s.n. ( W) ; illegible locality, 4 June 67, Hinneberg s.n. ( B) .

HUNGARY. Borsod-Abaúj-Zemplén: Szendrölád, a la salida del pueblo en dirección a Szendrö, prados secos sobre calizas, junto a la carretera, 165 m, 48º20'59.5''N, 20º44'19.6''E, 26 July 2014, X. Giráldez , M. M . Martínez Ortega, B. Rojas Andrés & N. López González NLG 133 ( SALA155850 ) ; Budapest: Hungaria, Buda, Sadler s.n. ( W) ; Heves: Hungaria borealis, montes Mátra, collis Sár-hegye versus boreo-orientem ab oppido Gyöngyös , 250– 500 m, 10 May 1970, Vasak s.n. ( PR); Hungaria borealis, ad ruinam Sirokivár prope pag. Sirok , versus occidentem ab oppido Eger , 200–309 m, 11 May 1970, Vasak s.n. ( PR); Gyöngyös, Gyöngyösi Sárhegy , prados sobre calizas , 482 m, 47º48'36.1''N, 20º00'02.7''E, 25 July 2014, X. Giráldez , M. M . Martínez Ortega, B. Rojas Andrés & N. López González BR 274 ( SALA155849 ); Sirok , en la subida al castillo, herbazales en la subida al castillo, sobre calizas , 281 m, 47º56'23.0''N, 20º11'48.0''E, 25 July 2014, X. Giráldez , M. M . Martínez Ortega, B. Rojas Andrés & N. López González BR 273 ( SALA155857 ) ; Komárom-Esztergom: Labatlán, haca Bajót, colinas cercanas al pueblo, prados secos sobre calizas, 183 m, 47º44'22.2''N, 18º31'04.4''E, 27 July 2014, X. Giráldez , M. M . Martínez Ortega, B. Rojas Andrés & N. López González MO 6114 ( SALA155848 ) ; Pest: Hungaria, collinis pilisiensis, in locis arenosis ad opp. Kestelcs [= Kesztölc], 6 May 1972, Weber s.n. ( PR); Kiskunsági Nemzeti Park , entre Szabanet y Kunpeszer, prados secos sobre calizas , 96 m, 47º02'57.5''N, 19º18'54.6''E, 28 July 2014, X. Giráldez , M. M . Martínez Ortega, B. Rojas Andrés & N. López González BR 277 ( SALA155860 ); Tatarszentgyörgy, Kikunság, Vitezsos , prados secos sobre arenas , 103 m, 47º03'58.6''N, 19º18'51.1''E, 24 July 2014, X. Giráldez , M. M . Martínez

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Ortega, B. Rojas Andrés & N. López González MO 6109 (SALA155859); unknown or dubious county: Ad margines in valle ¿Zuanoy?, July 90, ¿Holuby? s.n. (K).

ITALY. Alessandria: Colle di Casale Monferrato, July 1877, Negri s.n. (BC661136, WU); Aosta: Piemont, Gran Paradiso, Cognetal SW Aosta, zwischen Vieyes und Epinel, an Silikatböschung nahe der Strasse, ca. 1360 m, 7–20 June 1986, Polatschek s.n. (W); Bergamo: Alpes bergamaques, collines de Trescore, 28 April 1?14, Chenevard s.n. (G); Bolzano: Bozen, Tirol, 30 May 1903, Pringsheim s.n. ( PRC); Sarner Alpen, am Weg zwischen Terlan und Rumsein, Bozner Quarzporphyr, Trockenrasen, ca. 800 m, 29 April to 3 May 1978, Polatschek s.n. (W); Süd tirol, bei Bozen am Weg nach Virgl, alluvium, ca. 300 m, 25 April 1902, Handel-Mazetti s.n. (WU); Firenze: Near Florence, April 1873, Duttrie s.n. (BM); Calenzano, 12 Km nordwest. Florenz, in trockenem Rasen, 80 m, 22 April 1944, Sterneck s.n. (PR); Near Florence, April 73, Duthie s.n. (E); Florence Galuzzo, 2 May 1872, Joad s.n. (K); Luoghi incolti S. Montepescoli (Pelago), April 1884, Beren??? s.n. ( PRC); Grosseto: Entre Seggione y Castiglione d'Orcia, bordes de bosque de Quercus y Pistacia sobre calizas en margen de prado pastoreado, 420 m, 42º57'50.1''N, 11º34'08''E, 15 June 2014, X. Giráldez , M. M. Martínez Ortega, N. Padilla García & N. López González MO6082 (SALA155853); along the road from Seggiano to Poggio Uccello, margin of street, ca. 400 m, 26 April 2000, W. & S. Till Till (WU); L’Aquila: Assergi, entre la estación del funicular Fonte Cerrato y el campo di Imperatore, Gran Sasso, prados herbosos, 1213 m, 42º25'15.3''N, 13º32'48.8''E, 13 June 2014, X. Giráldez , M. M. Martínez Ortega, N. Padilla García & N. López González NPG8 (SALA155855); Gioia dei Marsi, pista desde Sperone al Monte Serrato, prados sobre calizas, 1333 m, 41º56'48.8''N, 13º43'36.3''E, 13 June 2014, X. Giráldez , M. M. Martínez Ortega, N. Padilla García & N. López González NPG6 (SALA155856); Parma: In agro parmensi, N. lungo le strade sui colli, F. in April–May, unknown collector (K); Perugia: Ex collibus Spoletanis, Umbria, April 1842, J. Ball s.n. (K); Sondrio: Bormio, limite del campi al Forte, solo calcareo, 1230 m, 29 May & 3 June 1909, Longa s.n. (RNG); Teramo: Aprutii, Monti dei Fiori, Orsini s.n. (BM); Trento: Tirolia austral., Val di Ledro, in pratis apricis, sol. calcar., 800–1000 m, 10 May 1884, Porta s.n. (K); Dofs, Trento, 6 May 1880, Sardagna s.n. (WU); Verona: Veron. Prov. in prat. (illegible word), spl. calcar., torri, 240–800 m, 25 April 69, Rigo s.n. (K); Venetia, distr. Veron. mte. Baldo S?nago, sol. calcar., 2–3000 m, 22 May 1870, Rigo s.n. (K); subida al Monte Baldo, desde Spiazzi a la Malga de Ime, prados secos sobre calizas, 1047 m, 45º39'42.6''N, 10º50'36.2''E, 20 June 2014, X. Giráldez , M. M. Martínez Ortega, N. Padilla García & N. López González NLG118 (SALA155854); Vicenza: Colli Berici, in herbosis aridis, in summitate montis Bellaguarda, solo basaltico, 140 m, May 1911 –13, Béguinot s.n. (K); unknown or dubious province: Monte di prato, Toscana, a Monte lapi presso la Caprain, solo arenaceo, 6 May 1906, Lacaita s.n. (BM); Casale, coteaux des environs de la ville, May 1877, Negri s.n. (G).

POLAND. Lublin: Ciechanki prope Leczna, in declivio loessaceo aprico ad fluvium Wieprz prope eius oram, 15 May 1959, Fijalkowski s.n. ( K) ;

REPUBLIC OF MACEDONIA ( FYROM). Tetovo: In ca. Mts. Kobelica (alp. Scard. Occ.); cult. in horto alp. bot. Sundermann-Lindau , July 1919, J . Bornmüller s.n. ( B100325910 ) ;

ROMANIA. Brașov: Gesprengberg bei Kronstadt [= Brașov], Jurakalk, May 1887, RÖmer s.n. (PR); Wiesen bei Honigberg [= Hărman], 540 m, 3 July 1912, Bornmüller s.n. (B); Cluj: Transsilvania, distr. Cluj, in declivibus meridionalibus collinae Ciresii prope pagum Pata, ca. 390 m, 6 June 1923, Bujoreau & al. s.n. (W); Cluj Napoca. Cojocna, en las afueras del pueblo hacia Ceanu Mare, prados, 323 m, 46º44'32.8''N, 23º50'18.5''E, 19 July 2014, X. Giráldez , M. M. Martínez Ortega, B. Rojas Andrés & N. López González MO 6099 (SALA155845); Cluj Napoca, Fanatele Clujului, prados sobre calizas, 503 m, 46º50'15.7''N, 23º37'14.3''E, 20 July 2014, X. Giráldez , M. M. Martínez Ortega, B. Rojas Andrés & N. López González MO 6101 (SALA155846); Fanatele Clujului, a 8 Km de Cluj-Napoca, prados sobre calizas, ca. 550 m, 30 June 2000, X. Giráldez , M. Martínez Ortega & J. A. Sánchez Agudo MO904 (SALA124620); Kojocna, prados sobre calizas, 1 July 2000, X. Giráldez , M. Martínez Ortega & J. A. Sánchez Agudo MO906 (SALA124618); Hunedoara: Hateg, entre Hätägel y Civla Mare, prados herbosos secos, 386m, 45.58382ºN, 22.82140ºE, 21 July 2014, X. Giráldez , M. M. Martínez Ortega, B. Rojas Andrés & N. López González BR 272 (SALA155847); Dolj: Oltenia, distr. Dolj, in pratis siccis, loco “Valea rea” dicto inter Segarcea et Radovan, ca. 140 m, 28 April 1967, Cirtu & Cirtu s.n. ( PRC); Oltenia, distr. Dolj, in pratis siccis, loco “Valea rea” dicto inter Segarcea et Radovan, ca. 140 m, 28 April 1967, Cirtu & Cirtu s.n. (WU); Iași: Iasi, Brezau, pasune, 10 May 1962, Toma s.n. (BM); Sibiu: Sura Mare, alrededores de Hamba, prados herbosos, 494 m, 45º52'16.8''N, 24º12'14.8''E, 21 July 2014, X. Giráldez , M. M. Martínez Ortega, B. Rojas Andrés & N. López González BR 269 (SALA155852).

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Phytotaxa 285 (1) © 2016 Magnolia Press • 53

RUSSIA. Omsk Oblast: On dry banks of the river Irtysch near Omsk, September 1900, N . Sidelnikow s.n. ( TU265983 ); dubious region: Daurica, 1829, Trinius 267 ( W0049262 ) .

SERBIA. South Bačka: Sirmia Petrovaradinum, 21 April 1910, Kupcok s.n. (PR); South Banat: Jugoslavia, Serbia, Banat, Dliblatska pescara-Devojacki bunari, in arenosis, 140 m, 26 April 1967, Mayer s.n. (BM); A. S. P. Vojvodina, südl. Banat, im Sande bei Deliblát, 31 May 1909, Watzl s.n. (WU); Devojački Bunar, Vladimirovac, prados secos sobre arenas, 163 m, 45º00'57.8''N, 20º57'17.3''E, 27 July 2010, S. Andrés , X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés SA428 (SALA149319); Devojački Bunar, Vladimirovac, prados sobre arenas, 155 m, 45º00'10.8''N, 20º56'21.1''E, 27 July 2010, S. Andrés , X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés SA430 (SALA149320); Pirot: Jugoslavien, Serbien, 10 km N Pirot, Strassenrand, trockenrasenfragmente (Terra fusca), c. 500 m, 17 May 1962, Ehrendorfer & Sorger s.n. (WU); Entre Pirot y Koprivštica, prados secos sobre calizas, 806 m, 43º13'35.5''N, 22º36'24.3''E, 21 July 2010, S. Andrés , X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés SA399 (SALA149318); Prizren: Entre Stirilje y Mališevo, cerca de la rotonda para coger la autovía hacia Pizren, 670 m, 42º23'36.4''N, 20º51'59.7''E, 7 June 2015, M. Martínez Ortega, X. Giráldez, N. Padilla & N. López NPG 30 (SALA157014).

SLOVAKIA. Banská Bystrica: Hradisko ( Hrusov ), 18 May 1985, V . Mikolás s.n. ( B100327614 ) ; Košice: in pratis ad pagum Szepessümeg [= Smizany] prope opp. Igló [= Spišská Nová Ves ], solo calcareo, ca. 500 m, 9 May 1916, Filarszky & Kümmerle s.n. ( K, PR, W) ; Nitra: Kamenica nad Hronomm, en la salida del pueblo hacia Chilaba , prados secos sobre caliza en borde de bosque, 118 m, 47º49'30.7''N, 18º44'46.4''E, 27 July 2014, X. Giráldez , M. M . Martínez Ortega, B. Rojas Andrés & N. López González MO 6113 ( SALA155851 ) .

SLOVENIA. Central Slovenia: Auf trockenen Wiesen b. Laibach, D. Graf s.n. (BM).

SWITZERLAND. Valais: Ste. Maurice, La Batia près Martigny, Charat s.n. (G); Fully, May & June 64 & 71, unknown collector - communicated to J. D. Hooker by Favrat and Barbey s.n. (K); Kt. Wallis, Gemeinde Orsières, Westhaang, chez les Girouds, Verunkrauteter Acker, 1100 m, 24 May 1950, H. Bührer s.n. (SEV16475); Charat, June 1865, Chevalier s.n. (G); Saillon, May, unknown collector (G); environs de Martigny, unknown collector (G); Sembraucher Valais, 800 m, 1903, DÖrfler s.n. (W); Saxon, lieux secs, bords des chemins, 540 m, 8 June 1876, Masson s.n. (WU); Vallée de Bagnes, 700 m, 20 May 1900, Wilczek s.n. (E); Saxon, 19 May 1806, Hamilton s.n. (E); unknown locality, unknown date, Thomas s.n. (E); Charrat cerca de Martigny, cerca de la "Ruta de los Adonis", terraplenes herbosos cerca de viñedos, 532 m, 46º07'38.6''N, 07º08'44.3''E, 12 June 2012, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés BR215 (SALA149312).

UKRAINE: Ternopil: Na stromym zboczu lewego brzegu jaru Dniestru powyzej mostu kolejowego, in ripa abrupta sinistra flum. Tyrae , 18 May 1938, J . Madalski & K . Matusiak s.n. ( B100327624 ) .

UNKNOWN OR DUBIOUS LOCATION: Transilvania, Czetz s.n. (K); Spont. prope Francofuntum, unknown collector (K).

Material provisionally assigned to V. prostrata : — GERMANY. Dubious state: in apricis calcareis prope Kal ?satdt (Palatinatum), 18 May 1850, illegible collector ( PR) .

Figures

FIGURE 15. Veronica prostrata (a–h. Ukraine, Zalishchyky Raion, Zalischyky, B 100327624; j–k. Czech Republic, Beroun, Srbsko, Doutnáč hill, M. Martínez-Ortega MO1331, SALA124619; i. Austria, Burgenland, Leithagebirge, Donnerskirchen, M. Martínez- Ortega MO1055, SALA124616). a. Habit; b. Stem indument; c. Medium leaf (adaxial leaf); d. Detail of a medium leaf (adaxial face); e. Medium leaf (abaxial leaf); f. Detail of a medium leaf (abaxial face); g. Apical shoot leaf (adaxial face); h. Apical shoot leaf (abaxial face); i. Calyx; j. Fruiting raceme; k. Part of a fruiting raceme, with a capsule. Illustration by Juan Castillo.

FIGURE 16. Distribution map of V. prostrata based on herbarium specimens (black circles) and collected material (open circles). Localities corresponding to the specimens examined from Russia are not represented.