Anthobium nigrum (Cameron, 1924)

Shavrin, Alexey V. & Smetana, Aleš, 2018, A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group, Zootaxa 4508 (4), pp. 451-506 : 500

Key to species of Anthobium nigrum group (modified Shavrin & Smetana 2017)

1 Lateral edges of pronotum with very indistinct smooth crenulation, latero-apical margins of elytra with moderately large teeth. Head reddish-brown, pronotum, elytra and abdomen yellow-brown. Aedeagus as in Fig. 98 . Habitus as in Fig. 95 . Body length: 3.15–3.64 mm. Nepal................................................... A. flaveolum Shavrin & Smetana , sp.n.

- Lateral edges of pronotum with distinct crenulation, latero-apical margins of elytra with small acute teeth. Body darker..... 2

2 Anterior angles of pronotum strongly protruded anteriad. Elytra markedly wider than long. Aedeagus as in Fig. 77 ( Shavrin & Smetana 2017). Habitus as in Fig. 67 ( Shavrin & Smetana 2017). Body length: 3.90–4.20 mm. China: Yunnan ................................................................................. A. conjunctum Shavrin & Smetana, 2017

- Anterior angles of pronotum slightly protruded anteriad. Elytra as wide as long or slightly longer than wide.............. 3

3 Punctation of pronotum very dense. Aedeagus as in Fig. 80 ( Shavrin & Smetana 2017). Habitus as in Fig. 68 ( Shavrin & Smetana 2017). Body length: 3.50–4.27 mm. China: Yunnan ............... A. densepunctatum Shavrin & Smetana, 2017

- Punctation of pronotum sparse.......................................................................... 4

4 Pronotum slightly flattened in cross section, with indistinct mediobasal impression. Aedeagus as in Fig. 72 ( Shavrin & Smetana 2017). Habitus as in Fig. 66 ( Shavrin & Smetana 2017). Body length: 4.15–4.37 mm. China: Gansu .................................................................................A. anishchenkoi Shavrin & Smetana, 2017

- Pronotum very convex, with deep mediobasal impression...................................................... 5

5 Middle portion of head distinctly convex, with indistinct transverse impression. Aedeagus as in Fig. 70 ( Shavrin & Smetana 2017). Habitus as in Fig. 65 ( Shavrin & Smetana 2017). Body length: 3.44–4.75 mm. India..... A. nigrum ( Cameron, 1924)

- Middle portion of head flattened, without or with very indistinct transverse impression. Aedeagus different.............. 6

6 Elytra as wide as long. Body reddish-brown to black. Median lobe of aedeagus narrow, apical portions of parameres widened ( Shavrin & Smetana 2017: Fig. 82 ). Habitus as in Fig. 69 ( Shavrin & Smetana 2017). Body larger: 4.00– 4.71 mm. Nepal, Bhutan...................................................................... A. ivani Shavrin & Smetana, 2017

- Elytra slightly longer than wide. Body brown. Median lobe of aedeagus moderately wide, apical portions of parameres narrow ( Fig. 96 ). Habitus as in Fig. 94 . Body smaller: 3.50 mm. Nepal............................ A. alticola ( Coiffait, 1977)

Figures

FIGURE 65. Distribution of Anthobium confucii (black circles) and A. liliputense (black square) in the south-western China.

FIGURES 96–99. Aedeagus of Anthobium alticola (96–97: NEPAL: Annapurna Mts.) and A. flaveolum (98–99: holotype): 96, 98—parameral view, 97, 99—lateral view. Scale bar: 0.1 mm.

FIGURES 94–95. Habitus of Anthobium nigrum group: 94—A. alticola (NEPAL: Annapurna Mts.) and 95—A. flaveolum (paratype). Scale bar: 1.0 mm.

FIGURES 74–78. Habitus of Anthobium fusculum group: 74—A. liliputense (paratype), 75—A. monticola (NEPAL: Lamjung), 76—A. rugosum (NEPAL: Dhaulagiri), 77—A. sociale (INDIA: Uttarakhand), 78—A. splendidulum (CHINA: Yunnan, Gaoligong Shan). Scale bar: 1.0 mm.

FIGURES 66–67. Aedeagus of Anthobium cuccodoroi (holotype): 66—parameral view, 67—lateral view. Scale bar: 0.1 mm.

FIGURES 79–82. Aedeagus of Anthobium liliputense (79–80: holotype) and A. monticola (81–82: NEPAL: Lamjung): 79, 81—parameral view, 80, 82—lateral view. Scale bar: 0.1 mm.

FIGURES 68–72. Aedeagus of Anthobium laozii (68–69: paratype; CHINA: Yunnan, Diancang Shan, 70: paratype; CHINA: Cangshan) and A. latissimum (71–72: paratype): 68, 71—parameral view, 69, 72—lateral view, 70—apical part of aedeagus, parameral view. Scale bar: 0.1 mm.