Scathophaga obscura ( Fallén, 1819 )

Ozerov, A. L. & Krivosheina, M. G., 2021, A review of the genus Scathophaga Meigen, 1803 (Diptera: Scathophagidae) of Russia, Russian Entomological Journal 30 (2), pp. 201-246 : 232-233

Scathophaga obscura ( Fallén, 1819)

Figs 1 , 34, 103–105, 136.

obscura Fallén, 1819: 9 ( Cordylura ). Type-locality: “ Scania ... Lårketorp Ostrogothiae” ( Sweden).

REMARK. The species was registered in Russia from Kola Peninsula [ Gorodkov, 1970: 453], Caucasus and Kuril Islands [ Gorodkov, 1986: 33], Siberia (Taimyr Peninsula) [ Ozerov, Barkalov, 2014: 564] and Far East [ Ozerov, Krivosheina, 2014: 218].

MATERIAL EXAMINED. Altai: Kosh-Agach env., (50.01ºN 88.66ºE), 1750 m, 2–4.VII.2016, N. Vikhrev (1 ♀, ZMUM); Kosh-Agach, plateau Ukok, 2400 m, surrounding area of Lake Muzdy-Bulak (49.26ºN, 87.65ºE), 10.VII.2008, A. Barkalov (1 ♂, ISEA); Onguday env., (50.77ºN 86.09ºE), 8–13.VII.2016, N. Vikhrev (3 ♂♂, ZMUM); Seminsky pass, River Turala (50.99ºN 85.68ºE), 1350 m, 8–12.VII.2016, N. Vikhrev (1 ♂, ZMUM); Tigirek Vill. (51.13ºN 83.05ºE), 550 m, 9–12.VII.2014, I. Shamshev (1 ♂, 3 ♀♀, ZISP); Arkhangelsk Oblast: 73 km NW of Nar’yan-Mar (ca. 67.981ºN 51.608ºE), 3.VIII.1978, Gorodkov (1 ♂, 1 ♀, ZISP); Nar’yan-Mar (67.631ºN, 52.985ºE), 12.VII.2008, A.L. Ozerov (1 ♂, ZMUM); Solvychegovsk (61.342ºN 46.913ºE), 12– 13.VIII.2010, D. Gavryushin (4 ♂♂, 12 ♀♀, ZMUM); Bashkiria: between Abzakovo and Murakaevo (ca. 53.797ºN 58.651ºE), 2–8.VIII.2008, K. Tomkovich (1 ♀, ZMUM); Buryatia: the upper reaches of the River Irkut near Lake Il’chir (51.961ºN 100.952ºE), 7.VIII.2012, A. Medvedev (1 ♂, 1 ♀, ZMUM); Chelyabinsk Oblast: Taganay (55.277ºN 59.795ºE), 18–24.VII.2008, K. Tomkovich (2 ♂♂, 1 ♀, ZMUM); Chukotka: Meynypil’gyno env. (62.628ºN 176.976ºE), 14.VII.2015, P.S. Tomkovich (1 ♂, ZMUM); Kamchatka Krai: Kozyrevsk (56.048ºN 159.870ºE), 23.VII.1985, V. Zlobin (1 ♀, ZISP); Karelia: Primorsky (66.555ºN 33.006ºE), 5.VII.2010, A.L. Ozerov (7 ♂♂, 4 ♀♀, ZMUM); Khabarovsk Krai: Bulgino env. (59.371ºN 143.147ºE), 29.VIII.1987, Gorodkov (1 ♂, ZISP); Khakassia: W Sayan pass (51.717ºN 89.882ºE), 2100 m, 9–12.VII.2017, N. Vikhrev (1 ♂, ZMUM); Krasnodar Krai: Teberdinskiy Nature Reserve, (43.443ºN 41.738ºE), 2.IX.1965, Gorodkov (1 ♂, ZISP); Krasnoyarsk Krai: Igarka (67.457ºN 86.598ºE), River Yenisei, 30. VI.1967, Gorodkov (4 ♂♂, 2 ♀♀, ZISP); Igarka (67.457ºN 86.598ºE), River Yenisei, 1.VII.1967, Gorodkov (1 ♀, ZISP); Kryuchkovo Station (56.096ºN 92.109ºE), 14–23.VII.2009, K. Tomkovich (1 ♂, ZMUM); Leningrad Oblast: Bol’shoy Berezovyy I. (60.3ºN 28.62ºE), 7.VIII.1981, Kandybina (1 ♂, ZISP); Gor’kovskoe, Lake Lebyazh’e (60.312ºN 29.532ºE), 24.VIII.2003, A. Ovchinnikov (1 ♀, ZISP); Luga (ca. 58.73ºN 29.84ºE), 10.VII.1952, 27.VIII.1953, A. Stackelberg (1 ♂, 1 ♀, ZISP); Rozhdestveno (59.322ºN 29.946ºE), 28.VIII.1956, A. Stackelberg (1 ♂, ZISP); Yashchera (58.894ºN 29.820ºE), 28.VIII.1957 and 11.VII.1959, A. Stackelberg (4 ♂♂, 2 ♀♀, ZISP); Moscow and Moscow Oblast: Dmitrov env. (56.316ºN 37.725ºE), 7. V.2006, N.Vikhrev (1 ♂, ZMUM); Ivanovskoe (55.925ºN 35.605ºE), 17.VII.2007, A.L. Ozerov (1 ♂, ZMUM); Izmaylovo (55.800ºN 37.832ºE), 5.VII.1983, A.L. Ozerov (3 ♂♂, 3 ♀♀, ZMUM); Naro-Fominsk (55.375ºN 36.756ºE), 8–13. V.2006, 14. V., 22. VI.2010, D. Gavryushin (1 ♂, 2 ♀♀, ZMUM); Pavlovskoe env. (55.921ºN 35.588ºE), 14.VIII.2006, A.L. Ozerov (2 ♂♂, 2 ♀♀, ZMUM); Rozhdestveno (56.932ºN 35.625ºE), 17.VII.2007, A.L. Ozerov (1 ♂, 1 ♀, ZMUM); Yakshino (55.922ºN 35.577ºE), 1.VIII.2005, A.L. Ozerov (4 ♂♂, 4 ♀♀, ZMUM); Zelenograd (55.986ºN 37.202ºE), 23.IV.1999, A. Gusakov (1 ♂, ZMUM); Murmansk Oblast: Molochnoe (68.853ºN 33.016ºE), 20.VII.2011, A. Ozerov (4 ♂♂, 1 ♀, ZMUM); Murmansk env. (68.979ºN 33.114ºE), 21.VII.2011, A. Ozerov (1 ♀, ZMUM); Revda (67.942ºN 34.552ºE), 15.VIII.1981, Gorodkov (8 ♂♂, 6 ♀♀, ZISP); Novgorod Oblast: Terekhovo (58.072ºN 33.333ºE), 29.VIII.1948 (1 ♀, ZMUM); Novosibirsk Oblast: River Inya (55.0ºN 83.2ºE), 22.VII.2012, O. Kosterin (2 ♂♂, ZMUM); Primorsky Krai: Gornotaezhnoe (43.697ºN 132.156ºE), 5.IX.1980, Zlobin (1 ♀, ZISP); Lazovsky Nature Reserve, Proselochnaya Bay (43.015ºN 134.124ºE), 3– 5.VII.2007, 6.VIII.2009, A. Ovchinnikov (3 ♂♂, 2 ♀♀, ZISP); Suputinsky [=Ussuriysky] Nature Reserve (ca. 46.591ºN 137.021ºE), 18.VIII.1980, Zlobin (1 ♂, 1 ♀, ZISP); Lazovsky Nature Reserve, cordon Korpad’ (43.263ºN 134.131ºE), 8–10.VII.2007, 6.VIII.2009, A. Ovchinnikov (1 ♂, 1 ♀, ZISP); Preobrazhenie (42.909ºN 133.890ºE), 16–17.VII.2007, A. Ovchinnikov (1 ♂, ZISP); Tigrovyy (=Tigrovoe) (43.189ºN 132.894ºE), 22.VIII.1979, Zinov’ev (1 ♂, ZISP); Vladivostok env., Sedanka (43.213ºN 131.960ºE), 20. VI.19278, A. Stackelberg (1 ♂, ZISP). Ryazan’ Oblast: Zalesnoe (54.969ºN 41.327ºE), 21–26.VII.2013, N. Vikhrev (1 ♂, ZMUM); Sakhalin Oblast: Kunashir I., Golovnin Volcano (43.841ºN 145.504ºE), 24.VII.1973, Kasparyan (1 ♂, ZISP); Kunashir I., Tret’yakovo (43.989ºN 145.644ºE), 7.VIII.1971, Nartshuk (1 ♂, ZISP); same place, 3.VIII.1973, Kasparyan (1 ♂, ZISP); Sakhalin I., Yuzhno-Sakhalinsk (46.954ºN 142.736ºE), 22.VII.1956, Violovich (1 ♂, 1 ♀, ZISP); Tuva: W Sayan pass (51.69ºN 89.96ºE), 1840 m, 9.VII.2017, N. Vikhrev (1 ♂, ZMUM); Tyumen’ Oblast: 75 km WSW of Samburg (ca. 67.035ºN 76.541ºE), 7–9.VIII.1976, Gorodkov (6 ♂♂, 5 ♀♀, ZISP); Neroyka (ca. 64.57ºN 59.67ºE), 600 m, 5.VIII.1990, Malozemov (2 ♂♂, ZISP); Voronezh Oblast: Ramon’ (51.908ºN 39.333ºE), 15.IX.1978, A. Shatalkin (1 ♀, ZMUM); Yakutia: Deputatskiy (69.319ºN 139.966ºE), 6.VIII.1974, Gorodkov (11 ♂♂, 10 ♀♀, ZISP). See also Ozerov & Krivosheina [2014: 218] and Ozerov & Barkalov [2014: 564].

DESCRIPTION. Male. Female ( Fig. 1 ). Body-length 4.5–6.1 mm.

Head. Frontal vitta yellow or reddish-yellow, matt; fronto-orbital plate blackish, greyish dusted. Face, parafacial and gena yellow, with delicate whitish or golden reflection. Postcranium blackish. 3 orbital and 3–4 frontal setae present. Antenna black; postpedicel about twice as long as wide. Arista bare. Palpus yellow.

Thorax black, densely pale grey dusted. Acrostichals absent, dorsocentrals 2+3, intra-alars absent, supra-alars 1+2, postpronotals 1–2. Scutellum greyish dust- ed, with a pair of strong basal scutellar and a pair of strong apical scutellar setae.

Legs. All coxae black, greyish dusted. All femora greyish dusted, black, except yellow apex. All tibiae and tarsi yellow. Fore femur covered with whitish hairs, with 4–5 dorsal setulae in apical half. Fore tibia with 1

dorsal, 0–1 posterior, 1 preapical dorsal, and 1 posteri- or apical setae. Mid femur with 1 preapical posterior seta. Mid tibia with 1 anterodorsal, 1 posterodorsal, 0– 1 anteroventral setae, also with a ring of apicals. Hind femur with a row of anterodorsal setulae. Hind tibia with 2 posterodorsal, 2 anterodorsal, 1 preapical dorsal, 1 preapical anterodorsal and 1 apical anteroventral setae.

Wing clear; crossveins r-m and d-m not darkened.

Abdomen black, densely pale grey dusted, covered with short whitish hairs, without marginal setae. Male sternite 4 about twice as long as wide (Fig. 34). Male sternite 5 with moderately long and narrow lateral lobes ( Fig. 103 ). Cercal plate longer than surstyli and narrowing to a slender incurved tip ( Figs 104, 105 ). Aedeagus as in Fig. 136 .

DISTRIBUTION. Russia: throughout midland, north to southern tundra. — Mongolia, North America.

Figures

Figs 1–2. Males of Scathophaga obscura (Fallén) (1) and Scathophaga stercoraria (Linnaeus) (2). Photographs by D.I. Gavryushin. 2 — after Ozerov & Freidberg, 2011, fig. 24. Рис. 1–2. Самцы Scathophaga obscura (Fallén) (1) и Scathophaga stercoraria (Linnaeus) (2). Фото Д.И. ГаврюШина. 2 — по Ozerov & Freidberg, 2011, fig. 24.

Figs 97–105. Scathophaga multisetosa (Holmgren) (97–99), S. nigripalpis (Becker) (100–102) and S. obscura (Fallén) (103–105): 97, 100, 103 — male sternite 5; 98, 101, 104 — epandrium, cercal plate and surstyli, lateral view; 99, 102, 105 — same, dorsal view. Рис. 97–105. Scathophaga multisetosa (Holmgren) (97–99), S. nigripalpis (Becker) (100–102) и S. obscura (Fallén) (103–105): 97, 100, 103 — стернит 5 самца; 98, 101, 104 — Эпандрий, церки и сурстили, сбоку; 99, 102, 105 — то же, сверху.

Figs 130–137. Aedeagus of Scathophaga spp., lateral view: 130 — S. buryatica Ozerov et Krivosheina; 131 — S. decipiens (Haliday); 132 — S. incompleta Ozerov et Krivosheina; 133 — S. lapponica (Ringdahl); 134 — S. mollis (Becker); 135 — S. nigripalpis (Becker); 136 — S. obscura (Fallén); 137 — S. tinctinervis (Becker). 130 — after Ozerov & Krivosheina, 2019, fig. 2; 131, 132 — after Ozerov & Krivosheina, 2020, figs 5, 11. Рис. 130–137. Эдеагус Scathophaga spp., сбоку: 130 — S. buryatica Ozerov et Krivosheina; 131 — S. decipiens (Haliday); 132 — S. incompleta Ozerov et Krivosheina; 133 — S. lapponica (Ringdahl); 134 — S. mollis (Becker); 135 — S. nigripalpis (Becker); 136 — S. obscura (Fallén); 137 — S. tinctinervis (Becker). 130 — по Ozerov & Krivosheina, 2019, fig. 2; 131, 132 — по Ozerov & Krivosheina, 2020, figs 5, 11.