Anteon madagascolum ( Benoit, 1954 )

Olmi, Massimo, Copeland, Robert S. & Noort, Simon Van, 2019, Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea), Zootaxa 4630 (1), pp. 1-619 : 161-165

57. Anteon madagascolum ( Benoit, 1954)

( Figs 56E, F , 57D )

Xenanteon madagascola Benoit 1954: 418 .

Anteon madagascolum (Benoit) : Olmi 1984: 393; 1994c: 4; Azevedo et al. 2010: 878.

Anteon brooksi Olmi 2003: 26 (syn. nov.); type locality: Antananarivo ( Madagascar); Azevedo et al. 2010: 876.

Description. ♀. Fully winged; body length 2.5–3.0 mm. Head black, except mandible testaceous (with teeth brown); antenna brown, except antennomeres 1–6 or 1–4 testaceous (occasionally antenna completely brown or testaceous, except antennomere 10 darkened); mesosoma and metasoma black; tegula testaceous; legs brown, except tarsi testaceous, or testaceous, except metacoxa brown. Antenna clavate; antennomeres in following proportions: 13:5:4:4:4:4:5:5:5:7. Head shiny, granulate and reticulate rugose; frontal line present; frons with or without lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 7; OL = 4; OOL = 5; OPL = 6; TL = 4. Pronotum shiny, granulate; posterior surface very long, more than 0.5 × as long as mesoscutum (10:13); pronotal tubercle reaching tegula. Mesoscutum shiny, granulate. Notauli absent; occasionally notauli present and incomplete, reaching about 0.2–0.3 × length of mesoscutum. Mesoscutellum and metanotum shiny, slightly granulate. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity granulate and with numerous irregular keels or sculptured by small areolae, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2r-rs&Rs vein much shorter than proximal part (3:9). Protarsomeres in following proportions: 7:2:3:3:12. Protarsomere 4 produced into hook, about 0.5 × as long as basal part of 5. Enlarged claw ( Fig. 57D ) with proximal prominence bearing one long bristle. Protarsomere 5 ( Fig. 57D ) with two rows of 10–14 lamellae, among which one very long distal lamella. Tibial spurs 1/1/2.

♂ (not reared; tentatively associated to the above ♀). Fully winged; body length 1.0– 1.9 mm. Head black, except mandible testaceous; antenna testaceous, except antennomeres 3–10 darkened (in holotype of A. brooksi , antenna totally testaceous; in other specimens, antenna brown, except scape and pedicel testaceous); mesosoma and metasoma black; occasionally metasoma brown; legs testaceous, except coxae, clubs of femora and tibiae partly brown (in holotype of A. brooksi , pro- and mesoleg completely testaceous). Antenna filiform; antennomeres in following proportions: 6:3.5:3:3:3:3:3:3:3:4.5. Head dull, reticulate rugose; frontal line complete; occipital carina complete; POL = 6; OL = 3; OOL = 3; OPL = 2; TL = 3. Mesoscutum shiny, slightly or strongly granulate. Notauli incomplete, reaching about 0.3–0.4 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured.

Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity without longitudinal keels, completely reticulate rugose (in specimen from Madagascar, 21°13.67’S 47°22.19’E, propodeal declivity with tracks of two complete longitudinal keels). Forewing hyaline, without dark transverse bands; distal part of 2r-rs&Rs vein much shorter than proximal part (1.5:5.5). Paramere ( Figs 56E, F ) candelabrum shaped, with distal part very slender, with an inner lobe large or small, covered or not with papillae. Tibial spurs 1/1/2.

Material examined. Types: ♀ holotype of A. madagascolum : MADAGASCAR: Toliara, Bekily , XI.1938 , A. Seyrig leg. ( MNHN); ♂ holotype of A. brooksi : MADAGASCAR: Antananarivo, Antananarivo, 25–27.X.1984 , YPT, Robert W. Brooks leg. ( NAMUK). Paratype of A. madagascolum : MADAGASCAR: same locality label as holotype, IX.1938 , A. Seyrig leg., 1♀ ( MRAC); Antsiranana, Antsiranana, 1893, Ch. Alluaud leg. , 1♀ (not exam- ined) ( Benoit 1954); Fianarantsoa, Ranomafana , X.1938 , A. Seyrig leg., 1♀ (not examined) ( Benoit 1954). Other material: COMOROS ISLANDS: Mohéli Island, Ouallah , 12°18’13’’S 43°40’06’’E, 500 m, 18–22.I.2009 , MT, rainforest, B.L. Fisher et al. leg., BLF20683 , 1♂ ( CAS). MADAGASCAR: Antananarivo, 11.5 km 147° SSE of Anjozorobe, 3 km 41°NE of Andranomay, 18°28.24’S 47°57.36’E, 1300 m, 5–13.XII.2000 , MT, montane rainforest, Fisher , Griswold et al. leg., code BLF 2375 , 2♂♂ ( CAS); Antananarivo, 46 km NE of Ankazobe , Ambohitan- tely, 18°11.88’S 47°16.89’E, 700 m, 17–24.X.2003 , MT, sclerophyl forest, M. Irwin & R. Harin’Hala leg., MA-27- 02, 1♂ ( CAS); Antsiranana, Ankarana, IX.1986 , MT, Jane Wilson leg., 1♀ ( NHMUK); Antsiranana, 7 km N of Joffreville , 12°20’S 49°15’E, 300 m, 7–27.IV.2001 , dry forest, MT, R. Harin’Hala leg., MA- 01-07-10, 1♂ ( CAS); Antsiranana, Diana Region, Orangea dry forest, Ramena , Baie de dune, 900 m E of Camp Minier , 23.II–2.III.2011 , 12°13.97’ S 49°21.99’E, M. Irwin, R.Harin’Hala leg., MT, dry forest, elev. 152 m, MG-62-11, 1♂ ( CAS); same locality label, 15–22.VI.2011 , MG-62-27, 1♂ ( CAS); same locality label, 18–26.V.2011 , MG-62-23, 1♂ ( CAS); Antsiranana, Orangea , 3 km E of Ramena near fort, 12°14.49’S 49°22.17’E, 65 m, 23–27.I.2001 , MT, littoral forest on sand, MA- 01-05-02, M.E. Irwin, E.I. Schlinger & R. Harin’Hala leg., 1♂ ( CAS); Antsiranana, Montagne d’Ambre Nat. Park , 12°30.52’S 49°10.53’E, Antsiranana, 960 m, 26–29.I.2001 , MT, M.E. Irwin, E.I. Schlinger & R. Harin’Hala leg., MA-01-01A-02, 3♂♂ (2 in CAS, 1 in MOLC); same locality label, 7–27.I.2007 , M. Irwin, F. Parker & R. Harin’Hala leg., MA-01-01A-16, 1♂ ( CAS); same locality label, 7–27.I.2007 , MA-01-01A-16, 1♂ ( CAS); same locality label, 21–28.I.2001 , M.E. Irwin, E.I. Schlinger & R. Harin’Hala leg., MA-01-01A-01, 1♂ ( CAS); same locality label, 27.I–11.II.2007 , MA-01-01A-17, 1♀ ( MOLC); same locality label, 21–26.I.2001 , MA- 01-01A-01, 4♂♂ (3 in CAS, 1 in MOLC); Antsiranana, Montagne d’Ambre National Park , 12°31’S 49°11’E, 975 m, 19.III–5.IV.2001 , MT, M.E. Irwin, E.I. Schlinger & R. Harin’Hala leg., MA-01-01B-07, 1♂ ( CAS); Antsirinana, Montagne d’Ambre National Park , 12°31.13’S 49°10.45’E, 1125 m, 29.I–11.II.2001 , MT, R. Harin’Hala leg., MA- 01-01D-03, 2♂♂ ( CAS); same locality label, 14–30.V.2001 , MA-01-01D-11, 1♂ ( CAS); Antsiranana, Montagne d’Ambre National Park , 12°31’S 49°11’E, 975 m, MT, M.E. Irwin, E.I. Schlinger & R. Harin’Hala leg., MA-01- 01B-01 , 1♂ ( CAS); Antsiranana, Marojejy National Park, Manantenina river , 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 14°26.06’S 49°45.36’E, 775 m, 11–14.XII.2005 , MT, rainforest, B.L. Fisher et al. leg., BLF13383 , 1♂ ( CAS); Antsiranana, SAVA Region, Vohemar District, Andranotsimaty dry forest, 9 km NE Daraina, 13°10.17’S 49°42.04’E, 90 m, 19–26.XII.2010 , MT, dry dense forest, M. Irwin, R. Harin’Hala leg., MG-57-12, 1♂ ( CAS); same locality label, 4–10.X.2010 , MG-57-01, 1♂ ( CAS); Fianarantsoa, 7 km W of Ranomafana , 900 m, 1–7.III.1990 , W.E. Steiner leg., 1♀, 1♂ ( USNM); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21°15.99’S 47°25.21’E, 1020 m, 10–14.I.2002 , secondary tropical forest, MT, R. Harin’Hala leg., 1♀ ( MOLC); same locality label, 4–12.II.2002 , MA-02-09C-15, 2♂♂ ( CAS); same locality label, 10–14.I.2002 , MA-02-09C-11, 2♂♂ ( CAS);same locality label, 26.II–4.III.2002 , MA-02-09C-18, 1♂ ( CAS); same locality label, 28.XI–6.XII.2001 , MA-02-09C-05, 3♂♂ ( CAS); same locality label, 4–14.VII.2002 , MA-02-09C-34, 1♂ ( CAS); same locality label, 24.VII–4.VIII.2002 , MA-02-09C-36, 5♂♂ ( CAS); same locality label, 8–15.XI.2001 , MA-02-09C-02, 1♂ ( MOLC); same locality label, 4–16.V.2003 , MA-02-09C-60, 2♂♂ ( CAS); same locality label, 12–23.IV.2003 , MA-02-09C- 58, 1♂ ( CAS); same locality label, 15–22.XI.2001 , MT, secondary tropical forest, R. Harin’Hala leg., MA-02-09C- 03, 1♂ ( CAS); same locality label, 19–26.II.2002 , MA-02-09C-17, 13♂♂ (8 in CAS, 5 in MOLC); same locality label, 15–22.XI.2001 , MA-02-09C-03, 1♂ ( CAS); same locality label, 23.VI–4.VII.2002 , MA-02-09C-33, 2♂♂ ( CAS); same locality label, 10–21.III.2003 , MA-02-09C-56, 3♂♂ ( CAS); same locality label, 14–21.I.2002 , MA- 02-09C-12, 1♂ ( CAS); same locality label, 8–15.XI.2001 , MA-02-09C-02, 2♂♂ ( CAS); same locality label, 13– 23.VI.2002 , MA-02-09C-32, 3♂♂ ( MOLC); same locality label, 26–31.III.2002 , MA-02-09C-22, 3♂♂ ( CAS); same locality label, 12–19.II.2002 , MA-02-09C-16, 4♂♂ ( CAS); same locality label 22–28.XI.2001 , MA-02-09C- 04, 9♂♂ (7 in CAS, 2 in MOLC); same locality label, 16.X–8.XI.2001 , MA-02-09C-01, 6♂♂ ( CAS); Fianarantsoa, Ranomafana , JIRAMA water works, 21°14.91’S 47°27.13’E, 690 m, 28.XI–6.XII.2001 , MT, near river, R. Harin’Hala leg., MA-02-09D-05, 4♂♂ (1 in CAS, 3 in MOLC); same locality label, 10–14.I.2002 , MA-02-09D-11, 1♂ ( CAS); same locality label, 21–24.XII.2001 , MA-02-09D-08, 1♂ ( CAS); same locality label, 2–10.I.2002 , MA- 02-09D-10, 3♂♂ ( CAS); Fianarantsoa, Ranomafana National Park, Vohiparara , 21°13.57’S 47°22.19’E, at broken bridge, 1110 m, 21–28.I.2002 , MT, high altitude rainforest, R. Harin’Hala leg., MA-02-09A-13, 2♂♂ ( CAS); same locality label, 19–26.II.2002 , MA-02-09A-17, 2♂♂ ( CAS, MOLC); same locality label, 15–25.VII.2002 , MA-02- 09A-35, 1♂ ( MOLC); same locality label, 14–21.I.2002 , MA-02-09A-12, 7♂♂ (5 in CAS, 2 in MOLC); same lo- cality label, 21–28.I.2002 , MA-02-09A-13, 2♂♂ ( CAS); same locality label, 22–28.XI.2001 , MA-02-09A-04, 3♂♂ ( CAS); same locality label, 15–21.XII.2001 , MA-02-09A-07, 2♂♂ ( CAS); same locality label, 6–15.XII.2001 , MA-02-09A-06, 2♂♂ ( CAS); Fianarantsoa, Ranomafana National Park, Vohiparara , 21°13.67’S 47°22.19’E, at broken bridge, 1110 m, 25.VII–3.VIII.2002 , MT, high altitude rainforest, R. Harin’Hala leg., MA-02-08A-36, 5♂♂ (3 in CAS, 2 in MOLC); Fianarantsoa, Ranomafana National Park , radio tower at forest edge, 21°15.05’S 47°24.43’E, 1130 m, 4–12.III.2002 , MT, mixed tropical forest, R. Harin’Hala leg., MA-02-09B-19, 4♂♂ (3 in CAS, 1 in MOLC); same locality label, 16.X–8.XI.2001 , MA-02-09B-01, 9♂♂ (8 in CAS, 1 in MOLC); same locality label, 7– 18.VI.2003 , 1♂ ( CAS); same locality label, 12–19.II.2002 , MA-02-09B-16, 3♂♂ ( CAS); same locality label, 14– 24.VI.2002 , MA-02-09B-32, 3♂♂ ( CAS); same locality label, 15–27.IV.2003 , MA-02-09B-58, 1♂ ( CAS); same locality label, 12–19.III.2002 , MA-02-09B-20, 4♂♂ ( CAS); same locality label, 9–20.III.2003 , MA-02-09B-55, 1♀ ( MOLC) , 2♂♂ ( CAS); same locality label, 27.II–9.III.2003 , MA-02-09B-54, 5♂♂ ( CAS); same locality label, 18–29.VI.2003 , MA-02-09B-64, 3♂♂ ( CAS); same locality label, 15–21.XII.2001 , MA-02-09B-07, 2♂♂ ( CAS); same locality label, 22–28.XI.2001 , MA-02-09B-04, 2♂♂ ( CAS); same locality label, 20.III–3.IV.2003 , MA-02- 09B-56, 1♂ ( CAS); same locality label, 27.IV–7.V.2003 , MA-02-09B-59, 2♂♂ ( CAS); same locality label, 24.V– 4.VI.2002 , MA-02-09B-30, 1♂ ( CAS); same locality label, 7–17.V.2003 , MA-02-09B-60, 2♂♂ ( CAS); same local- ity label, 14–21.I.2002 , MA-02-09B-12, 10♂♂ (8 in CAS, 2 in MOLC); same locality label, 27.VI–7.VII.2003 , MA-02-09B-65, 2♂♂ ( CAS); same locality label, 18–27.II.2003 , MA-02-09B-53, 2♂♂ ( CAS, MOLC); same local- ity label, 7–17.VII.2003 , MA-02-09B-66, 1♂ ( CAS); same locality label, 17–30.V.2003 , MA-02-09B-61, 3♂♂ ( CAS); Fianarantsoa, Manombo Special Reserve camp site, 32 km SSE of Farafangana, 23°01.31’S 47°43.20’E, 36 m, 22.V–6.VI.2005 , MT, lowland rainforest, MA-28-20, M. Irwin, R. Harin’Hala leg., 1♂ ( CAS); same locality label, 30.X–12.XI.2004 , MA-28-03, 1♂ ( CAS); Fianarantsoa, near Isalo National Park , in dry wash east of Interpre- tive Center, 22°37.60’S 45°21.49’E, 885 m, 12–22.VI.2002 , MT, in open area, MA-02-11B-32, R. Harin’Hala leg., 1♂ ( MOLC); same locality label, 26.I–3.II.2002 , MA-02-11B-13, 1♂ ( CAS); same locality label, 7–15.IV.2002 , MA-02-11B-23, 1♀ ( CAS); Fianarantsoa, Ananalava Forest , 29.6 km 280° W Ranohira, 22°35.30’S 45°07.42’E, 700 m, 1–5.II.2003 , MT, tropical dry forest, Fisher, Griswold et al. leg., BLF7383, 1♂ ( CAS); Mahajanga, Marotan- drano Special Reserve, Marotandrano , 48.3 km S Mandritsara, 16°17’00’’S 8°48’52’’E, 865 m, 7–8.XII.2007 , transition humid forest, MT, B.L. Fisher et al. leg., BLF19157 , 1♂ ( MOLC); Mahajanga, Sofia District , 45 km S Antso- hihy, 5 km W of Anjiamangirana, elev. 97 m, 27I–2. II.2011, 15°09.42’S 47°47.05’E, M.Irwin, R.Harin’Hala, MT, Analagnambe dry degraded forest, MG-61-08, 1♂ ( MOLC); same locality label, 2–6.I.2011 , MG-61-04, 1♂ ( CAS); Mahajanga, Besalampy District, Analangidro dry forest, 7 km NE Besalampy, 16°41.49’S 44°31.41’E, 200 ft., 9– 16.X.2007 , MT, dry forest on sand, M. Irwin & R. Harin’Hala leg., MG-41-03, 2♂♂ ( CAS); same locality label, 28.X–4.XI.2007 , MG-41-06, 4♂♂ ( CAS); same locality label, 4–11.XI.2007 , 1♂ ( CAS); same locality label, 11– 18.XI.2007 , 1♂ ( CAS); same locality label, 18–25.XI.2007 , 4♂♂ ( CAS); same locality label, 24.IX–2.X.2007 , 5♂♂ ( CAS); Mahajanga, Maintirano District, Asondrodava dry forest, 15 km N of Maintirano, 17°57.92’S 44°2.13’E, 200 ft., 21–28.I.2008 , MT, dry forest, M. Irwin, R. Harin’Hala leg., MG-43A-16, 1♂ ( CAS); Mahajanga, Beanka Forest Reserve , 50.2 km E Maintirano, 18°01’35’’S 44°03’02’’E, 250 m, 19–26.X.2009 , tropical dry forest on tsingy, MT, B.L. Fisher et al. leg., BLF22787 , 1♂ ( CAS); Mahajanga, Analamanitra Forest , 14 km NE Misinjo, 15°55’S 45°57’E, 65 ft., 15–22.I.2008 , MT, dense dry forest, M. Irwin, R. Harin’Hala leg., MG-38-14, 1♂ ( CAS); Mahajanga, Ampijoroa National Park , 160 km N of Maevatanana on RN 04, 16°19’16’’S 46°48’80’’E, 43 m, 24– 31.VIII.2003 , MT, deciduous forest, R. Harin’Hala leg., MA-25-12, 1♂ ( CAS); same locality label, 9–20.XI.2003 , MA-25-23, 1♂ ( CAS);same locality label, 14–22.IX.2003 , MA-25-15, 1♂ ( CAS); same locality label, 7–14.IX.2003 , MA-25-14, 2♂♂ ( CAS); same locality label, 31.VIII–7.IX.2003 , MA-25-13, 1♂ ( CAS); Mahajanga, Namoroka National Park , 9.8 km 300° WNW Vilanandro, 16°28.00’S 45°21.00’E, 140 m, 4–8.XI.2002 , MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6445, 1♂ ( CAS); Toamasina, Ivondro , 1♀ ( AMNH); Mahajanga, Sofia Dis- trict, 45 km S Antsohihy, 5 km W Anjiamangirana, Analagnambe dry degraded forest, 15°09.42’S 47°47.05’E, 97 m, 26.V–2.VI.2011 , MT, MG-61-15, 2♂♂ ( CAS); Toliara, 12 km NW of Amboasary, Berenty , 5–15.V.1983 , J.S. Noyes & M.C. Day leg., 1♂ ( NHMUK); same locality label, 18–23.V.1983 , 1♂ ( AMNH); Toamasina, Mobot site, Analalava , 7 km SW of Foulpointe, 17°41’36’’S 49°27’37’’E, 60 ft., 16–23.XI.2007 , MT, sand in low altitude dense humid forest, MG-37B-10, 1♂ ( MOLC); Toliara, 60 km NE of Morondava, Bereboka , 18–23.V.1983 , J.S. Noyes & M.C. Day leg., 4♂♂ ( AMNH, NHMUK); Toliara, Tsimanampetsotsa Nat. Park, Bemanateza Forest, 20.7 km 81°E Efoetse , 23 km 131° SE Beheloka, 23°59.32’S 43°52.50’E, 90 m, 22–26.III.2002 , beating low vegetation, in spiny forest thicket, Fisher, Griswold et al. leg., BLF 6256, 1♀ ( CAS); same locality label, 22–26.III.2002 , BLF 6257, 1♂ ( CAS); same locality label, 22–26.III.2002 , BLF 6257, 1♂ ( CAS); Toliara, Andohahela National Park, Ihazofotsy Parcelle III, 24°50.09’S 46°29.21’E, 80 m, 26.XII.2002 – 5.I.2003 , MT, dry spiny forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02-21-05, 1♀ ( CAS); same locality label, 7–17.XII.2003 , MA- 02-21-41, 1♂ ( CAS); same locality label, 28.IV–6.V.2003 , MA- 02-21-17, 1♂ ( CAS); Toliara, Andohahela National Park, Tsimelahy , parcelle II, 24°56.21’S 46°37.60’E, 180 m, 16–17.XII.2002 , MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02-20-09, 2♂♂ ( CAS, MOLC); same locality label, 28.III–8.IV.2003 , 3♂♂ (2 in CAS, 1 in MOLC); same locality label, 10–20.VII.2003 , 1♂ ( CAS); same locality label, 17–24.VIII.2003 , MA- 02-20-36, 1♂ ( CAS); same locality label, 22–29.VI.2003 , MA- 02-20-30, 3♂♂ ( CAS); same locality label, 30.XI–11.XII.2003 , MA- 02-20-47, 2♂♂ ( CAS); Toliara, Fiherenana , 23°14.07’S 43°52.15’E, 50 m, 16.X–16.XI.2002 , MT. degraded gallery forest, MGF047, Frontier Wilderness Project, 1♀ ( CAS); Toliara, Fiherenana , 23°10.37’S 43°57.39’E, 100 m, 22– 28.X.2002 , MT. gallery forest, MGF041, Frontier Wilderness Project, 1♂ ( MOLC); Toliara, Ranobe , 23°02.22’S 43°36.377’E, 30 m, 5–9.II.2003 , MT, spiny forest thicket, MGF058, Frontier Wilderness Project, 1♂ ( CAS); Toli- ara, Ranobe , 23°02.367’S 43°36.617’E, 20 m, 8–15.XI.2003 , MT, gallery forest, MGF084, Frontier Wilderness Project, 1♀ ( MOLC); Toliara, Mikea Forest , NW of Manombo, 22°54.22’S 43°28.53’E, 30 m, 6–16.I.2002 , MT, deciduous dry forest, R. Harin’Hala leg., MA-02-18A-09, 1♀ ( CAS); same locality label, 6–16.I.2002 , MA-02- 18A-09, 1♂ ( CAS); Toliara, Mikea Forest , NW of Manombo, 22°54.80’S 43°28.93’E, 37 m, 18–29.IV.2002 , MT, spiny forest, R. Harin’Hala leg., MA-02-18B-20, 1♂ ( CAS); same locality label, 11–24.II.2003 , MA-02-18A-46, 1♂ ( CAS); Toliara, Zombitse National Park , near national road, 22°50.43’S 44°43.87’E, 825 m, 15.X–9.XI.2001 , MT, deciduous spiny forest, R. Harin’Hala leg., MA-02-13B-01, 1♂ ( CAS); same locality label, 7–13.VII.2002 , 1♂ ( CAS); same locality label, 13–20.III.2002 , 1♂ ( MOLC); Toliara, Zombitse National Park , near ANGAP office, 22°53.19’S 44°41.53’E, 840 m, 15.X–9.XI.2001 , MT, deciduous spiny forest, M.E. Irwin, F.D. Parker & R. Harin’Hala leg., MA-02-13A-01, 2♂♂ ( CAS, MOLC); Toliara, Beza Mahafaly Reserve , parcelle I near research station, 23°41.19’S 44°35.46’E, 165 m, 20–24.XII.2002 , MT, dry deciduous forest, R. Harin’Hala leg., MA-02- 14A-45, 1♂ ( CAS); Toliara, Beza Mahafaly Reserve, parcelle II near Bellevue , 23°41.39’S 44°34.53’E, 180 m, 15.X–10.XI.2001 , MT, spiny forest, M.E. Irwin, F.D. Parker & R. Harin’Hala leg., MA-02-14B-01, 1♂ ( CAS); Toliara, Antafoky , 23°28.43’S 44°03.51’E, 55 m, 25–28.I.2002 , MT, gallery forest, Frontier Project leg., MGF003, 1♂ ( CAS); Toliara, Berenty Special Reserve , 8 km NW Amboasary, 25°00.40’S 46°18.20’E, 85 m, 24.III–3.IV.2003 , MT, gallery forest, M. Irwin, F. Parker, R. Harin’Hala leg., MA- 02-22-20, 1♂ ( CAS); same locality label, 16– 27.XII.2002 , 2♂♂ ( CAS); same locality label, 25.II–7.III.2003 , MA- 02-22-17, 2♂♂ ( CAS, MOLC); same locality label, 17–24.III.2003 , MA- 02-22-19, 3♂♂ (2 in CAS, 1 in MOLC); same locality label, 5–15.II.2003 , MA- 02-22- 15, 4♂♂ ( CAS); same locality label, 7–17.III.2003 , MA- 02-22-18, 5♂♂ ( CAS); same locality label, 15–25.II.2003 , MA- 02-22-16, 3♂♂ ( CAS); same locality label, 17–26.I.2003 , MA- 02-22-13, 4♂♂ ( CAS); same locality label, 16–27.XII.2002 , MA- 02-22-10, 2♂♂ ( CAS); same locality label, 26.I–5.II.2003 , MA- 02-22-14, 3♂ # ♂ ( CAS); To- liara, Berenty Special Reserve , 8 km NW Amboasary, elev 35 m, 25°01.26’S 46°18.33’E, 22–30.XI.2003 , M.Irwin, F. Parker & R.Harin’Hala, MT, spiny forest, MA-02-22A-04, 3♂♂ (2 in CAS, 1 in MOLC); same locality label, 7–17.X.2004 , 1♂ ( CAS); Toliara, Berenty Reserve , 80 km W Fort Dauphin, 25°00’S 46°17’E, 7.IV.1994 , MT, M. Wasbauer leg., 1♂ ( DEUCD); Toliara, Beroboka village , 45 km NE Morondava, 19°58.65’S 44°39.92’E, 420 ft., 8–16.VI.2008 , MT, Antsarongaza gallery forest, MG-45B-26, M. Irwin, R. Harin’Hala leg. , 3♂♂ ( CAS); same lo- cality label, 430 ft., 22–30.V.2008 , MG-45A-24, 1♂ ( MOLC); same locality label, 1–9.VII.2008 , MG-45A-29, 2♂♂ ( CAS); Toliara, Mite Forest , 20.7 km 29° WNW Tongobory, 23°31.27’S 44°07.17’E, 75 m, 27.II–3.III.2002 , MT, in gallery forest, Fisher, Griswold et al. leg., BLF 5849, 1♂ ( CAS); Toliara, Forêt de Kirindy , 15.5 km 64° ENE Marofandilia, 20°04.09’S 044°39.38’E, 6–13.VIII.2008 , B.L. Fisher , MT, tropical dry forest, el 30 m, BLF18285_ 40 , 1♂ ( MOLC); same locality label, 15–22.X.2008 , BLF18285_50 , 1♂ ( MOLC); Toliara, Androimpano Forest , 5 km E of Itampolo, 24°39.02’S 43°57.79’E, 130 m, 3–16.VII.2009 , MT, path in dry forest, M. Irwin, R. Harin’Hala leg., MG-54C-33, 1♂ ( CAS); Toliara, Tsimanampetsotsa National Park, Mitoho Forest , 24°02.91’S 43°45.14’E, 120 m, 9–15.I.2009 , MT, transitional forest, M. Irwin, R. Harin’Hala leg., MG-53C-12, 1♂ ( CAS); same locality label, 24–29.X.2008 , MG-53C-01, 1♂ ( CAS); same locality label, 4–12.XII.2008 , MG-53C-07, 1♂ ( CAS); same locality label, 24–29.X.2008 , across trail at base of escarpment, dense dry forest, MG-53B-01, 1♂ ( CAS) .

Hosts. Unknown.

Distribution. Comoros Islands, Madagascar.

Remarks. The record from Diego Suarez Island reported by Olmi (1984) is erroneous, because it was based on a misinterpretation of a paper of Benoit (1954), and instead refers to the town of Diégo-Suarez situated in Madagascar. The new synonymy of Anteon brooksi is justified because the small difference in the ♂ genitalia of A. madagascolum (the only difference between these two species) falls within the range of variability of the species, as shown by the study of hundreds of specimens.

Figures

FIGURE 56. ♂ genitalia of Anteon species.A: A. kivuanum (Benoit), from South Africa, Graskop; B–D: A. kwazuluense Olmi, holotype (B); ♂ from Madagascar, 21°15.99’S 47°25.21’E (C); ♂ from South Africa, Royal Natal National Park (D); E–F: A. madagascolum (Benoit), ♂ from Madagascar, Antananarivo (holotype of a junior synonym, Anteon brooksi Olmi) (E); ♂ from Madagascar, 12°30.52’S 49°10.53’E (F). Scale bar: A: 0.12 mm; B, C: 0.08 mm; D, F: 0.05 mm; E: 0.07 mm.

FIGURE 57. Chelae of Anteon species.A: A. kivuanum (Benoit), ♀ from Democratic Republic of the Congo, Upemba National Park; B: A. longiscapum (Ceballos), holotype; C: A. mabibiense Olmi, Copeland & van Noort sp. nov., holotype; D: A. madagascolum (Benoit), holotype; E: A. mcguirkae Olmi, Copeland & Guglielmino, holotype; F: A. maritimum (Turner), ♀ from South Africa, Milnerton. Scale bar: A, D: 0.09 mm; B: 0.07 mm; C, E: 0.04 mm; F: 0.06 mm.