Harpalus kaznakovi kaznakovi Kataev & Wrase, 1997

Kataev, Boris M. & Liang, Hongbin, 2007, Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini), Zootaxa 1604, pp. 1-20 : 12-14

Harpalus kaznakovi kaznakovi Kataev & Wrase, 1997

( Figs 25–27 , 30–41 )

Harpalus kaznakovi Kataev & Wrase, 1997: 996 . Type locality: road Luhuo–Sertar, pass 35 km NNE Luhuo, NW Sichuan, China.

Material examined. In addition to the type series the following specimens were examined:

China. SICHUAN: 1 ɗ, same data as holotype: NW Sichuan, road Luhuo–Sertar, pass 35 km NNE Luhuo, alpine region, 3500–4000 m, 27.VII.1994, L. Kalab leg. (cFCCH); 2 ɗ, 2 Ψ, Garzê Co., 3400 m, 29.VI.1983, Chen Yuanging & Zhang Xuezhong leg. ( IZB); 1 ɗ, same but 3300 m, 30.VI.1983, Chen Yuanging leg. ( IZB); 2 ɗ, 2 Ψ, same but 3650 m, 11.VII.1983, Wang Shuyong (2 specimens) & Chen Yuanqing (2 specimens) leg. ( IZB); 2 ɗ, same but 3600 m, 1.VII.1983, Cheng Yuanqing & Wang Shuyong leg. ( IZB); 3 ɗ, 1 Ψ, Wenchuan, 900–1000 m, 1.VIII.1983, Zhang Xuezhong leg. ( IZB); 1 ɗ, W Sichuan, 15 km NE Qianning, 30°35'N 101°41'E, 11.VII.1998, M. Bocak leg. (cWR); 1 ɗ, 1 Ψ, Kangding, 3700 m, 30.VIII. and 7.IX.1982, Wang Shuyong leg. ( IZB); 4 ɗ, 2 Ψ, Barkam Co., 2500 m, 20.VIII.1983, Wang Ruqi leg. ( IZB); 1 ɗ, Dêgê Co., 3900 m, 6.VII.1983, Wang Shuyong leg. ( IZB); 1 Ψ, Dêgê Co., Maniganggo, 3900 m, 8.VII.1983, Chen Yuanqing leg. ( IZB); 2 Ψ, Dêgê Co., 3800 m, 9.VII.1983, Wang Shuyong leg. ( IZB); 2 ɗ, 2 Ψ, same but 3900 m, 10.VII.1983, Zhang Xuezhong leg. ( IZB); 4 ɗ, 1 Ψ, same but 4200 m, 10.VII.1983, Niu Chunlai (3 specimens) and Chai Huaicheng (2 specimens) leg. ( IZB); 1 Ψ, same but 4000 m, 8.VII.1983, Wang Shuyong leg. ( IZB). TIBET: 1 Ψ, Jomda Co., 3400 m, 27.VII.1976, Zhang Xuezhong leg. ( IZB).

Description. Habitus: Figs 25–26 . Metallic tinge on dorsum variable, sometimes recognizable only along margins of pronotum and elytra. Body size very variable: body length 5.9–8.4 mm. Head moderate or relatively large (in male WHmax/WPmax = 0.58–0.72 and WHmin/WPmax = 0.53–0.61), with convex or almost flat eyes (WHmax/WHmin = 1.04–1.25). Pronotum variable in proportions and shape, 1.44–1.59 as wide as long, with sides rounded or straight, converging posteriad in basal half, sometimes sinuate; basal angles obtuse, somewhat sharp or rounded at apices; basal foveae small, variable in shape, usually elongate and not deep; base of pronotum finely punctate or almost smooth, only with few fine punctures in basal foveae. Elytra relatively short or elongate, 1.26–1.42 times as long as wide, 2.14–2.43 times as long and 1.09–1.21 times as wide as pronotum; parascutellar pore present or absent; each third interval with or without one discal pore. Metepisterna ( Fig. 27 ) wider than long, weakly narrowed posteriad. Metacoxae usually without additional setigerous pores, occasionally with a small such pore medially. Median lobe of aedeagus ( Figs 30–31, 34–37, 39–40 ) more or less strongly bent after basal bulb; terminal lamella ( Figs 32–33, 38, 41 ) comparatively long Distribution. Known from north-western Sichuan and eastern Tibet (Jomba County).

Variation. The nominotypical subspecies demonstrates geographical variation in several characters, mainly in body size and shape of eyes and pronotum, and apparently represents a complex of several local forms which however seem to be not sharply separated from each other. At least based on the scarce available material delimitation of these forms involves big difficulties. In Sichuan, the specimens from Garzê County, Wenchuang and Kangding are similar to those of the type series; they are, on average, rather large (6.1–8.2 mm) and robust but eyes are only scarcely convex, almost flat ( Fig. 25 ; WHmax/WHmin = 1.04–1.10). The median lobe of aedeagus of these specimens is usually almost straight from dorsal view or only slightly curved to right ( Figs 30–31, 34–35, 36–37 ); in several specimens from Garzê County the median lobe is narrower and less strongly arcuate ( Figs 34–35 ) than that in most other specimens. The pronotum of the specimens from Garzê County, Wenchuang and Kangding is usually with sides rounded or straight, converging posteriad in basal half, sometimes sinuate (more markedly so in the specimens from Wenchuan), and basal angles obtuse and rounded at apices. In one male examined from the environs of Qianning, eyes are scarcely convex but pronotal sides clearly sinuate basally and basal angles nearly right and rather sharp at apices. By contrast, in the specimens from Barkam and Dêgê counties (Sichuan) as well as Jomba County (Tibet), body is, on average, smaller (5.9–7.0 mm) and more slender and eyes are more convex ( Fig.26 ; WHmax/WHmin = 1.11–1.25) than those in the most specimens from Garzê, Wenchuang and Kangding counties. The median lobe of aedeagus of these specimens is markedly arcuate and with apical portion more strongly curved to right ( Figs 39–40 ). The armature of the internal sac in all forms is very similar.

Remarks. Although the nominotypical subspecies is highly variable, it may be recognized by the short and wide metepisterna, the terminal lamella of the median lobe of aedeagus nearly parallel–sided and the body larger and more robust.

Figures

FIGURES 25 – 29. Harpalus kaznakovi. 25 – 27, H. k. kaznakovi (25, 27, Sichuan, Garzê Co.; 26, Sichuan, Dêgê Co.); 28 – 29, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 25 – 26, 28, habitus, 27 – 29, left metepisternum. Scales = 1.0 mm (A: Figs 25 – 26, 28; B: 27, 29).

FIGURES 30 – 44. Harpalus kaznakovi, median lobe of aedeagus (32 – 33, 38, 41, 44, terminal lamella). 30 – 41, H. k. kaznakovi (30 – 38, Sichuan, Garzê Co.; 39 – 41, Sichuan, Dêgê Co.). 42 – 44, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 30, 32 – 34, 36, 38 – 39, 41 – 42, 44, dorsal aspect; 31, 35, 37, 40, 43, lateral aspect. Scales: A = 0.5 mm (Figs 32 – 33, 38, 41, 44), B = 1.0 mm (Figs 30 – 31, 34 – 37, 39 – 40, 42 – 43).