Harpalus giacomazzoi giacomazzoi Kataev & Wrase, 1996

Kataev, Boris M. & Liang, Hongbin, 2007, Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini), Zootaxa 1604, pp. 1-20 : 16-17

Harpalus giacomazzoi giacomazzoi Kataev & Wrase, 1996

( Figs 45–53 )

Harpalus giacomazzoi Kataev & Wrase, 1996: 519 . Type locality: 50 km E Batang, road Litang–Batang, Sichuan, China.

Type material. Holotype: ɗ, China, Sichuan, road Litang–Batang, 50 km E Batang, 14–16.VII.1992, E. Giacomazzo leg. (cWR). Paratypes: 1 ɗ, same data as holotype (cSCI); 1 ɗ, 1 Ψ, Batang, 2500 m, 15.VII.1992, E. Giacomazzo leg. (cSCI).

Additional material examined. China. SICHUAN: 5 ɗ, 2 Ψ, Dêgê Co., 3100–3300 m, 4.VII.1983, Niu Chunlai (6 specimens) and Wang Shuyong (1 specimen) leg. ( IZB); 1 Ψ, Dêgê Co., 3200 m, 4.VII.1983, Chen Yuanqing leg. ( IZB); 7 Ψ, Batang, 2500 m, 13.VIII.1982, Wang Shuyong leg. ( IZB); 23 ɗ, 6 Ψ, "Yun-nan: Tsekou et Se-Tchouen, Principante de Batang: Yargong, J.A. Soulie, 1907" ( MNHN, ZISP); 5 ɗ, 5 Ψ, Xiangcheng Co., 2900 m, 18.VI.1982, Wang Shuyong (8 specimens) and Zhang Xuezhong (2 specimens) leg. ( IZB).

Questionable locality (probably mislabelled and actually Sichuan): 1 ɗ, 1 Ψ, "Peking, Westberge, Exp. Stötzner", " type " [male] or "cotype" [female], " pekinensis Schaub." [according to our data, this taxon has never been described by E. Schauberger] ( OÖLL).

Description. Size variable: body length 7.8–10.8 mm. Head moderate (WHmax/WPmax = 0.55–0.63 and WHmin/WPmax = 0.46–0.52). Pronotum variable in form, from nearly oval to more or less trapezium– shaped, but in all cases more strongly narrowed apicad than basad and with base notably wider than apex; basal angles narrowly or widely rounded. Elytra 1.33–1.45 times as long as wide, 2.28–2.57 times as long and 1.07–1.16 times as wide as pronotum; preapical sinuation weak or moderately deep. Wings reduced to scale extending to about 1/3 –3/4 of elytra. Three last abdominal sternites glabrous or with scattered additional short setae. Metacoxae with or without additional setae medially. Metafemur with 4–10 setigerous pores along posterior margin. Median lobe of aedeagus with terminal lamella notably widened apically ( Fig. 47, 50 ); internal sac with moderately wide longitudinal spiny patch and with or without two extremely small separate teeth ( Figs 45–46, 48–49 ).

Distribution. The nominotypical subspecies is distributed in western part of Sichuan.

Variation. In the specimens from Dêgê County, the pronotum is only slightly narrower than elytra (WPmax/WE = 1.07–1.08), the metacoxae are with many additional setae medially, the abdominal sternites are covered with scattered additional short setae, the metafemur is with 8–10 setigerous pores along posterior margin and the aedeagus has separate teeth in the internal sac larger than in the specimens from other known localities ( Figs 45–46 ). In the specimens from Batang County including the type series, the pronotum is variable but usually notably narrower than elytra (WPmax/WE = 1.09–1.16), the metacoxae are with or without additional setae medially, the abdominal sternites are usually glabrous, at most with few very short setae on three penultimate sternites, the metafemur possesses 4–7 setigerous pores along posterior margin and the separate teeth in the internal sac of the aedeagus are small or absent ( Figs 48–49 ). Among the specimens from Batang County, the specimens from “Yargong” are characterized by deeper preapical sinuation of elytra. The specimens examined from Xiangcheng County and “Peking” are similar to those from Batang County (WPmax/WE = 1.10–1.16) but the metacoxae seem to be constantly without additional setae.

Remarks. The most important distinctive characters of the nominotypical subspecies are the pronotum with comparatively wide base and the terminal lamella of the aedeagus notably widened apically. This taxon is highly variable in many of its morphological characteristics and it is possible that it will be divided into two or several subspecies when more populations will be examined.

Figures

FIGURES 45 – 53. Harpalus giacomazzoi, median lobe of aedeagus (47, 50, 53, terminal lamella). 45 – 50, H. g. giacomazzoi (45 – 47, Sichuan, Dêgê Co.; 48 – 50, Sichuan, Yargong); 51 – 53, H. g. gracilis ssp. n. (Tibet, Markam Co., holotype). 45, 47 – 48, 50 – 51, 53, dorsal aspect); 46, 49, 52, lateral aspect). Scales: A = 0.5 mm (Figs 47, 50, 53), B = 1.0 mm (Figs 45 – 46, 48 – 49, 51 – 52).