Exiligada punctata, Criscione & Law & Köhler, 2012

Criscione, Francesco, Law, Margot Louisa & Köhler, Frank, 2012, Land snail diversity in the monsoon tropics of Northern Australia: revision of the genus Exiligada Iredale, 1939 (Mollusca: Pulmonata: Camaenidae), with description of 13 new species, Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 166 (4), pp. 689-722 : 709-710

publication ID	https://doi.org/ 10.1111/j.1096-3642.2012.00863.x
publication LSID	lsid:zoobank.org:pub:AC086999-76A8-4F93-9A51-5B5B213A50E6
DOI	https://doi.org/10.5281/zenodo.5480110
persistent identifier	https://treatment.plazi.org/id/03E9878D-FFCE-FF86-7000-A65D5B48F947
treatment provided by	Marcus
scientific name	Exiligada punctata
status	sp. nov.

Treatment

EXILIGADA PUNCTATA View in CoL SP. NOV.

( FIGS 4D View Figure 4 , 6D View Figure 6 , 20D–F View Figure 20 , 22 View Figure 22 )

Holotype: AUSTRALIA, WA, EK, Halls Creek District , south of Panton River, low and open limestone area, numerous gullies, under well-developed spinifex, in talus, 17°49 ′ 23 ′′ S, 128°14 ′ 58 ′′ E (coll. V. Kessner, R. Hokkanen, 1.ix.2009; WA-76 /09) ( WAM S 83175 View Materials ). GoogleMaps

Paratypes: AUSTRALIA, WA, EK , same data as for holotype, ten wet ( WAM S49221 View Materials ); Halls Creek District , 57 km east of Nicholson Station H /S, very open limestone area, well-developed spinifex, talus, under spinifex and slabs, 17°58 ′ 43 ′′ S, 128°21 ′ 34 ′′ E (coll. V. Kessner, R. Hokkanen, 1.ix.2009; WA-75 /09), 15 wet,> 24 dry ( WAM S49220 View Materials ); Halls Creek District, large rather open limestone area, 4.5 km north of Panton River, numerous gullies, under spinifex, in talus, 17°46 ′ 47 ′′ S, 128°13 ′ 59 ′′ E (coll. V. Kessner, R. Hokkanen, 1.ix.2009; WA-77 /09), seven wet ( WAM S49222 View Materials ), 22 dry ( WAM S49139 View Materials ) GoogleMaps .

Other material: AUSTRALIA, WA, EK , Halls Creek District , 57 km east of Nicholson Station H /S, very open limestone area, well-developed spinifex, talus, under spinifex and slabs, 17°58 ′ 43 ′′ S, 128°21 ′ 34 ′′ E (coll. V. Kessner, R. Hokkanen, 1.ix.2009; WA-75 /09), 30 dry ( WAM S49137 View Materials ); Halls Creek District , south of Panton River , low and open limestone area, numerous gullies, under well-developed spinifex, in talus, 17°49 ′ 23 ′′ S, 128°14 ′ 58 ′′ E (coll. V. Kessner, R. Hokkanen, 1.ix.2009; WA-76 /09), 17 dry ( WAM S49138 View Materials ); Halls Creek District, south of Panton River, low and open limestone area, numerous gullies, under welldeveloped spinifex, in talus, 17°49 ′ 23 ′′ S, 128°14 ′ 58 ′′ E (coll. V. Kessner, R. Hokkanen, 1.ix.2009; WA-76 /09), four dry ( WAM S49144) GoogleMaps .

Etymology: In reference to the peculiar colour pattern, formed by fine maculations, derived from ‘punctatus’ (Latin = dotted), adjective of feminine gender.

Description: Shell ( Figs 4D View Figure 4 , 6D View Figure 6 , 20D–F View Figure 20 ). Large, moderately elevated, moderately thick; micropustulations absent. Shell background colour yellowish horn fading to whitish towards shell base, with several brown-reddish spiral bands, most conspicuous on body whorl, never present on shell base, often interrupted radially; sparse brown maculations in radial clusters present on whole spire and increasing in number on last whorls.

Genitalia ( Fig. 22 View Figure 22 ). Epiphallus one third of penis. Vas deferens entering penial sheath from one third up. Penis five times longer than wide, not coiled inside sheath, with inner wall supporting a large apical corrugated pad and six narrow, regularly shaped longitudinal pilasters, five of which interrupted posteriorly and one running along entire wall length. Vagina one third shorter than penis and as long as free oviduct, bursa copulatrix shape variable, without well-differentiated end or with a short to long rounded end.

Remarks: Shell colour pattern similar to E. pallida , E. longicauda , E. nodulicauda , and E. negriensis , but easily distinguishable by presence of maculations. Penis length relative to width (pl/pw) and epiphallus length relative to penis (el pl-1) smaller than average. Inner penial wall sculpture similar to E. negriensis and E. pallida , to be distinguished from these species by narrower posterior pad and longer longitudinal pilasters, always reaching the posterior part of the wall, and by latter by posterior pad being corrugated. Vagina length relative to penis (vl/pl) and relative to free oviduct (vl/ol) average.