Eucerini, Latreille, 1802

Eardley, Connal & Urban, Rosalind, 2010, Catalogue of Afrotropical bees (Hymenoptera: Apoidea: Apiformes) 2455, Zootaxa 2455, pp. 1-548 : 425-428

Eucerini :

Tetralonia Spinola

Macrocera Latreille, 1810: 339 , 439, not Macrocera Meigen, 1803 . Type species: Eucera antennata Fabrciius, 1793 , by original designation.

Tetralonia Spinola, 1839: 538 [replacement name for Macrocera Latreille ]. Type species: Eucera antennata Fabricius , designated by Latreille, 1810; Friese 1909a: 256–260; Strand 1911b: 144; Friese 1915: 267; Brauns 1926a: 193; Alfken 1932: 52–53; Cockerell 1933d: 456; Cockerell 1933n: 362–363; Cockerell 1933zd: 442; Cockerell 1935a: 86–88; Cockerell 1936i: 3–5; Arnold 1947: 198, 201, 202, 212, 218; Michener 1968c: 484; Rozen 1969a: 102; Medler 1980: 483; Eardley 1989: 15–21; Alexander 1990: 124; Pauly 1990: 153; Baker 1996b: 544; Michener 1997a: 18–19.

Eucera (Tetralonia) Spinola : Gerstaecker 1870: 349–350.

Anthophora (Eucara) Friese, 1905f: 241 . Type species: Anthophora (Eucara) laticeps Friese , designated by Cockerell, 1933d; Cockerell 1933d: 456.

Eucera [!] Friese: Friese 1909a: 86, 291; Pesenko and Sitdikov, 1990: 88.

Eucara Friese : Friese, 1909a: 264; Friese 1911a: 667; Friese 1921: 1093–4; Eardley 1989: 1–15; Michener 2000: 69, 715, 716.

Tetralonia (Eucara) Friese : Alfken 1932: 52–53; Michener 2000: 715, 716.

Tetralonia (Eucera) [!] Friese: Arnold 1947: 198, 204, 217, 218.

Thygatina Cockerell, 1911b: 237 . Type species: Thygatina fumida Cockerell, 1911 , by monotypy.

Tetralonia (Thygatina) Cockerell : Michener 2000: 69, 100, 715, 716.

Tetralonia boharti (Eardley)

Kenya, Tanzania.

Eucara boharti Eardley, 1989: 2–8 , 10, male holotype (BLCU) Tanzania.

Tetralonia boharti (Eardley) : Eardley and Urban 2006: 162.

Tetralonia caudata (Friese)

Kenya, Malawi, Mozambique, Nigeria, Tanzania, Zimbabwe.

Tetralonia caudata Friese, 1905d: 20–23 , female lectotype [designated by Eardley, 1989] (ZMHB) Tanzania; Friese 1909a: 256–257; Friese 1909b: 164; Meade-Waldo 1914b: 400; Rasmussen and Ascher 2008: 36.

Eucara caudata (Friese) : Eardley 1989: 2–4, 8–11.

Tetralonia (Eucara) caudata (Friese) : Michener 2000: 715 [by implication].

Anthophora (Eucara) piligera Friese, 1905f: 241 , female holotype (ZMHB) Kenya; Friese 1909a: 262, 265, 282; Friese 1909b: 140, 145, 165; Eardley 1989: 8, syn.; Rasmussen and Ascher 2008: 86.

Eucara piligera Friese : Cockerell 1933n: 363.

Tetralonia cinctula Cockerell

Botswana, Kenya, Namibia.

Tetralonia cinctula Cockerell, 1936o: 558–559 (part), female holotype (BMNH) Botswana; Eardley 1989: 15–16, 18– 20; Danforth et al. 2006b: 366, 367, 368.

Tetralonia (Thygatina) cinctula Cockerell : Michener 2000: 716 [by implication].

Tetralonia fraterna Friese

Botswana, Mozambique, Namibia, Niger, Senegal, South Africa.

Tetralonia fraterna Friese, 1911a: 655 , male lectotype [designated by Eardley, 1989] (ZMHB) Mozambique; Friese 1941: 100; Eardley 1989: 15–19, 21; Rasmussen and Ascher 2008: 51.

Tetralonia (Thygatina) fraterna Friese : Michener 2000: 716 [by implication].

Tetralonia rikatlaensis Friese, 1916a: 443–444 , male holotype (ZMHB) Mozambique; Friese 1941: 101; Eardley 1989: 18, syn.; Rasmussen and Ascher 2008: 90.

Tetralonia labrosa Friese

Kenya.

Tetralonia labrosa Friese, 1911a: 657 , male holotype (ZMHB) Kenya; Meade-Waldo 1914b: 400; Eardley 1989: 15, 17– 18; Rasmussen and Ascher 2008: 63.

Tetralonia (Thygatina) labrosa Friese : Michener 2000: 716 [by implication].

Tetralonia macrognatha (Gerstaecker)

Angola, Botswana, Burkina Faso, Congo, Democratic Republic of the Congo, Ethiopia, Ghana, Kenya, Malawi, Mozambique, Namibia, South Africa, Tanzania, Uganda, Zambia, Zimbabwe. On flowers of: Gossypium sp. , Hibiscus esculentus , Hibiscus sp. , Pavonia sp. , Taraxacum sp.

Eucera (Tetralonia) macrognatha Gerstaecker, 1870: 349–350 , male holotype (ZMHB) Kenya; Stadelmann 1898: 24.

Eucera macrognatha Gerstaecker : Dalla Torre 1896: 239.

Anthophora (Eucara) macrognatha (Gerstaecker) : Friese 1909a: 283.

Tetralonia macrognatha (Gerstaecker) : Friese 1909b: 164; Arnold 1947: 212.

Tetralonia (Eucara) macrognatha (Gerstaecker) : Alfken 1932: 53; Michener 2000: 715 [by implication].

Eucara (Tetralonia) macrognatha (Gerstaecker) : Cockerell 1933n: 362, 364.

Eucara macrognatha (Gerstaecker) : Cockerell 1933d: 456; Cockerell 1936l: 483; Cockerell 1938b: 370; Eardley 1989: 4–7, 10.

Anthophora (Eucara) laticeps Friese, 1905f: 241 , male holotype (ZMHB) 'Massailand' [ Kenya or Tanzania]; Friese 1909a: 262, 265, 282; Friese 1909b: 140, 144, 165; Friese 1915: 288; Cockerell 1933d: 456; Cockerell 1933g: 136; Cockerell 1933n: 363–364; Cockerell 1937i: 11; Sandhouse 1943: 549; Eardley 1989: 4, 7, syn.; Rasmussen and Ascher 2008: 63.

Anthophora (Eucara) haefligeri Friese, 1905f: 242 , female holotype (ZMHB) Tanzania; Friese 1909a: 262, 283; Friese

1909b: 140–164; Rasmussen and Ascher 2008: 56. Eucara haefligeri (Friese) : Friese 1911a: 666–667; Cockerell 1937f: 280, syn. laticeps ; Eardley 1989: 4, syn. Macrocera neavei Vachal, 1910b: 325 , female holotype (RMCA) Democratic Republic of the Congo. Tetralonia neavei (Vachal) : Meade-Waldo 1914b: 400; Eardley 1989: 4, 7, syn. Eucara neavei (Vachal) : Cockerell 1933n: 364, syn. Tetralonia laticeps . Tetralonia sheffieldi Meade-Waldo, 1914b: 400–401 , female holotype (BMNH) Malawi; Cockerell 1916a: 210;

Cockerell 1930b: 342. Eucara sheffieldi (Meade-Waldo) : Cockerell 1933d: 456, syn.; Cockerell 1938b: 370. Tetralonia sheffieldi var. ferrugineipes Meade-Waldo, 1914b: 401–402 , female holotype (BMNH) Uganda; Eardley

1989: 4, 7, syn. Eucara macrognatha ferrugineipes (Meade-Waldo) : Cockerell 1933n: 362, 364. Tetralonia sheffieldi umbiloensis Cockerell, 1917b: 40 , female holotype (lost) South Africa; Cockerell 1933n: 363, syn. Eucara straminea Cockerell, 1933g: 136 , female holotype (BMNH) South Africa; Eardley 1989: 4, 7, syn.

Tetralonia mesotes (Eardley)

Kenya.

Eucara mesotes Eardley, 1989: 2–4 , 13–15, male holotype (MCZC) Kenya.

Tetralonia mesotes (Eardley) : Eardley and Urban 2006: 162.

Tetralonia nigropilosa Friese

Ivory Coast, South Africa. On flowers of: Ipomoea arborescens , Ipomoea pes-scaprae .

Tetralonia nigropilosa Friese, 1911a: 655 , 657–658, male lectotype [designated by Eardley, 1989] (SAMC) South Africa; Cockerell, 1916a: 210; Cockerell 1920b: 303; Eardley 1989: 15–18, 21; Rasmussen and Ascher 2008: 78.

Tetralonia (Thygatina) nigropilosa Friese : Michener 2000: 716 [by implication].

Tetralonia nigropilosa var. nigrosellata Cockerell, 1920b: 303 ; Eardley 1989: 15–18, 21, syn.

Tetralonia obscuriceps Friese

Burkina Faso, Mali, Namibia, Niger, Tanzania, Zimbabwe. On flowers of: Hibiscus esculentus , Ipomoea sp.

Tetralonia obscuriceps Friese, 1916a: 443 , male holotype (ZMHB) Tanzania; Cockerell 1936o: 557; Eardley 1989: 15, 17, 20–21; Rasmussen and Ascher 2008: 80.

Tetralonia (Thygatina) obscuriceps Friese : Michener 2000: 715 [by implication].

Tetralonia nudicauda Cockerell, 1936o: 557 , male holotype (BMNH) Namibia; Eardley 1989: 20, syn.

Tetralonia obscuripes Friese

Tanzania.

Tetralonia obscuripes Friese, 1905d: 22 (part), female holotype (ZMHB) Tanzania; Eardley 1989: 21 [male paratype = Tetralonia abessinica Friese ]; Rasmussen and Ascher 2008: 81.

Tetralonia (Thygatina) obscuripes Friese : Michener 2000: 715 [by implication].

Tetralonia paulyi (Eardley)

Burkina Faso.

Eucara paulyi Eardley, 1989: 2–4 , 8–9, male holotype (PCGB) Burkina Faso.

Tetralonia paulyi (Eardley) : Eardley and Urban 2006: 162.

Tetralonia penicillata (Friese)

Ethiopia, Kenya, Tanzania.

Anthophora (Eucara) penicillata Friese, 1905f: 241–242 , male holotype (ZMHB) Tanzania; Friese 1909a: 262, 265, 282; Friese 1909b: 140, 145, 165; Rasmussen and Ascher 2008: 85.

Eucara penicillata (Friese) : Cockerell, 1933n: 363; Cockerell 1938b: 369–370; Eardley 1989: 11–12, 14.

Tetralonia penicillata (Friese) : Eardley and Urban 2006: 162.

Tetralonia ruficollis (Friese) Lesotho, South Africa. Eucara ruficollis Friese, 1911a: 667 , male lectotype [designated by Eardley, 1989] ( SAMC) South Africa; Cockerell

1933k: 136; Cockerell 1933n: 362, 364; Friese 1941: 101; Eardley 1989: 2–4, 12–14; Rasmussen and Ascher

2008: 92.

Tetralonia ruficollis (Friese) : Eardley and Urban 2006: 162.

Tetralonia testacea Smith

Australian

Tetralonia testacea Smith, 1854: 301 , male lectotype [designated by Michener 1965: 306] (BMNH) “Africa” [misidentified Nomia bituberculata (Smith) , see Michener 1965].

Tetralonia trichardti Cockerell

South Africa.

Tetralonia trichardti Cockerell, 1933g: 134–135 , female lectotype [designated by Eardley, 1989] (BMNH) South Africa; Cockerell 1936o: 556–557; Eardley 1989: 21.

Tetralonia (Thygatina) trichardti Cockerell : Michener 2000: 716 [by implication].